: A North Eastern; B North Western; AB North Eastern & North Western; C South Eastern; D Tropical Florida; E Texas-Mexican Boundary; F Rocky Mountains; G Oregon & California; H New Mexico & Arizona Mexican Boundary

MANUAL OF THE TREES OF NORTH AMERICA
(EXCLUSIVE OF MEXICO)

BY
CHARLES SPRAGUE SARGENT
Director of the Arnold Arboretum of Harvard University
Author of The Silva of North America

WITH SEVEN HUNDRED AND EIGHTY-THREE ILLUSTRATIONS FROM DRAWINGS BY
CHARLES EDWARD FAXON
AND
MARY W. GILL

Second Edition

BOSTON AND NEW YORK
HOUGHTON MIFFLIN COMPANY
The Riverside Press Cambridge
1922

TO
M. R. S.
THE WISE AND KIND FRIEND OF THIRTY YEARS
THIS BOOK IS DEDICATED
WITH GRATITUDE AND AFFECTION

PREFACE TO THE SECOND EDITION

The studies of the trees of North America (exclusive of Mexico) which have been carried on by the agents and correspondents of the Arboretum in the sixteen years since the publication of the Manual of the Trees of North America have increased the knowledge of the subject and made necessary a new edition of this Manual. The explorations of these sixteen years have added eighty-nine species of trees and many recently distinguished varieties of formerly imperfectly understood species to the silva of the United States, and made available much additional information in regard to the geographical distribution of American trees. Further studies have made the reduction of seven species of the first edition to varieties of other species seem desirable; and two species, Amelanchier obovalis and Cercocarpus parvifolius, which were formerly considered trees, but are more properly shrubs, are omitted. The genus Anamomis is now united with Eugenia; and the Arizona Pinus strobiformis Sarg. (not Engelm.) is now referred to Pinus flexilis James.

Representatives of four Families and sixteen Genera which did not appear in the first edition are described in the new edition in which will be found an account of seven hundred and seventeen species of trees in one hundred and eighty-five genera, illustrated by seven hundred and eighty-three figures, or one hundred and forty-one figures in addition to those which appeared in the first edition.

An International Congress of Botanists which assembled in Vienna in 1905, and again in Brussels in 1910, adopted rules of nomenclature which the world, with a few American exceptions, has now generally adopted. The names used in this new Manual are based on the rules of this International Congress. These are the names used by the largest number of the students of plants, and it is unfortunate that the confusion in the names of American trees must continue as long as the Department of Agriculture, including the Forest Service of the United States, uses another and now generally unrecognized system.

The new illustrations in this edition are partly from drawings made by Charles Edward Faxon, who died before his work was finished; it was continued by the skillful pencil of Mary W. Gill, of Washington, to whom I am grateful for her intelligent coöperation.

It is impossible to name here all the men and women who have in the last sixteen years contributed to this account of American trees, and I will now only mention Mr. T. G. Harbison and Mr. E. J. Palmer, who as agents of the Arboretum have studied for years the trees of the Southeastern States and of the Missouri-Texas region, Professor R. S. Cocks, of Tulane University, who has explored carefully and critically the forests of Louisiana, and Miss Alice Eastwood, head of the Botanical Department of the California Academy of Sciences, who has made special journeys in Alaska and New Mexico in the interest of this Manual. Mr. Alfred Rehder, Curator of the Herbarium of the Arboretum, has added to the knowledge of our trees in several Southern journeys; and to him I am specially indebted for assistance and advice in the preparation of the keys to the different groups of plants found in this volume.

This new edition of the Manual contains the results of forty-four years of my continuous study of the trees of North America carried on in every part of the United States and in many foreign countries. If these studies in any way serve to increase the knowledge and the love of trees I shall feel that these years have not been misspent.

PREFACE

In this volume I have tried to bring into convenient form for the use of students the information concerning the trees of North America which has been gathered at the Arnold Arboretum during the last thirty years and has been largely elaborated in my Silva of North America.

The indigenous trees of no other region of equal extent are, perhaps, so well known as those that grow naturally in North America. There is, however, still much to be learned about them. In the southern states, one of the most remarkable extratropical regions in the world in the richness of its arborescent flora, several species are still imperfectly known, while it is not improbable that a few may have escaped entirely the notice of botanists; and in the northern states are several forms of Cratægus which, in the absence of sufficient information, it has been found impracticable to include in this volume. Little is known as yet of the silvicultural value and requirements of North American trees, or of the diseases that affect them; and one of the objects of this volume is to stimulate further investigation of their characters and needs.

The arrangement of families and genera adopted in this volume is that of Engler & Prantl’s Die Natürlichen Pflanzenfamilien, in which the procession is from a simpler to a more complex structure. The nomenclature is that of The Silva of North America. Descriptions of a few species of Cratægus are now first published, and investigations made since the publication of the last volume of The Silva of North America, in December, 1902, have necessitated the introduction of a few additional trees described by other authors, and occasional changes of names.

An analytical key to the families, based on the arrangement and character of the leaves, will lead the reader first to the family to which any tree belongs; a conspectus of the genera, embodying the important and easily discovered contrasting characters of each genus and following the description of each family represented by more than one genus, will lead him to the genus he is trying to determine; and a similar conspectus of the species, following the description of the genus, will finally bring him to the species for which he is looking. Further to facilitate the determination, one or more letters, attached to the name of the species in the conspectus following the description of the genus, indicate in which of the eight regions into which the country is divided according to the prevailing character of the arborescent vegetation that species grows (see map forming frontispiece of the volume). For example, the northeastern part of the country, including the high Appalachian Mountains in the southern states which have chiefly a northern flora, is represented by (A), and a person wishing to learn the name of a Pine-tree or of an Oak in that region need occupy himself only with those species which in the conspectus of the genus Quercus or Pinus are followed by the letter (A), while a person wishing to determine an Oak or a Pine-tree in Oregon or California may pass over all species which are not followed by (G), the letter which represents the Pacific coast region south of the state of Washington.

The sign of degrees (°) is used in this work to represent feet, and the sign of minutes (′) inches.

The illustrations which accompany each species and important variety are one half the size of nature, except in the case of a few of the large Pine cones, the flowers of some of the [viii] Magnolias, and the leaves and flower-clusters of the Palms. These are represented as less than half the size of nature in order to make the illustrations of uniform size. These illustrations are from drawings by Mr. Faxon, in which he has shown his usual skill and experience as a botanical draftsman in bringing out the most important characters of each species, and in them will be found the chief value of this Manual. For aid in its preparation I am indebted to him and to my other associates, Mr. Alfred Rehder and Mr. George R. Shaw, who have helped me in compiling the most difficult of the keys.

SYNOPSIS
OF THE FAMILIES OF PLANTS DESCRIBED IN THIS BOOK

Resinous trees; stems formed of bark, wood, or pith, and increasing in diameter by the annual addition of a layer of wood inside the bark; flowers unisexual; stamens numerous; ovules and seeds 2 or many, borne on the face of a scale, not inclosed in an ovary; embryo with 2 or more cotyledons; leaves straight-veined, without stipules.

Carpels or pistils consisting of a closed cavity containing the ovules and becoming the fruit.

Stems with woody fibres distributed irregularly through them, but without pith or annual layers of growth; parts of the flower in 3’s; ovary superior, 3-celled; embryo with a single cotyledon; leaves parallel-veined, persistent, without stipules.

Stems formed of bark, wood, or pith, and increasing by the addition of an annual layer of wood inside the bark; parts of the flower mostly in 4’s or 5’s; embryo with a pair of opposite cotyledons; leaves netted-veined.

Subdivision 1. Apetalæ. Flowers without a corolla and sometimes without a calyx.

Section 1. Flowers in unisexual aments (female flowers of Juglans and Quercus solitary or in spikes); ovary inferior (superior in Leitneriaceæ) when a calyx is present.

Section 2. Flowers unisexual (perfect in Ulmus); calyx regular, the stamens as many as its lobes and opposite them; ovary superior, 1-celled; seed 1.

Section 3. Flowers usually perfect; ovary superior or partly inferior, 1—4-celled; leaves simple, persistent in the North American species.

Subdivision 2. Petalatæ. Flowers with both calyx and corolla (without a corolla in Lauraceæ, in Liquidambar in Hamamelidaceæ, in Cercocarpus in Rosaceæ, in Euphorbiaceæ, in some species of Acer, in Reynosia, Condalia, and Krugiodendron in Rhamnaceæ, in Fremontia in Sterculiaceæ, in Chytraculia in Myrtaceæ, in Conocarpus in Combretaceæ and in some species of Fraxinus in Oleaceæ).

A. Ovary superior (partly inferior in Hamamelidaceæ; inferior in Malus, Sorbus, Heteromeles, Cratægus, and Amelanchier in Rosaceæ).

Section 2. Gamopetalæ. Corolla of united petals (divided in Elliottia in Ericaceæ, 0 in some species of Fraxinus in Oleaceæ).

A. Ovary superior (inferior in Vaccinium in Ericaceæ, partly inferior in Symplocaceæ and Styracaceæ).

ANALYTICAL KEY
TO THE GENERA OF PLANTS INCLUDED IN THIS BOOK, BASED CHIEFLY ON THE CHARACTER OF THE LEAVES

TREES OF NORTH AMERICA
(Exclusive of Mexico)

Class 1. GYMNOSPERMÆ.

Ovules and seeds borne on the face of a scale, not inclosed in an ovary; resinous trees, with stems increasing in diameter by the annual addition of a layer of wood inside the bark.

I. PINACEÆ.

Trees, with narrow or scale-like generally persistent clustered or alternate leaves and usually scaly buds. Flowers appearing in early spring, mostly surrounded at the base by an involucre of the more or less enlarged scales of the buds, unisexual, monœcious (diœcious in Juniperus), the male consisting of numerous 2-celled anthers, the female of scales bearing on their inner face 2 or several ovules, and becoming at maturity a woody cone or rarely a berry. Seeds with or without wings; seed-coat of 2 layers; embryo axile in copious albumen; cotyledons 2 or several. Of the twenty-nine genera scattered over the surface of the globe, but most abundant in northern temperate regions, thirteen occur in North America.

CONSPECTUS OF THE NORTH AMERICAN GENERA.

1. PINUS Duham. Pine.

Trees or rarely shrubs, with deeply furrowed and sometimes laminate or with thin and scaly bark, hard or often soft heartwood often conspicuously marked by dark bands of summer cells impregnated with resin, pale nearly white sapwood, and large branch-buds formed during summer and composed of minute buds in the axils of bud-scales, becoming the bracts of the spring shoot. Leaves needle-shaped, clustered, the clusters borne on deciduous spurs in the axils of scale-like primary leaves, inclosed in the bud by numerous scales lengthening and forming a more or less persistent sheath at the base of each cluster. Male flowers clustered at the base of leafy growing shoots of the year, each flower surrounded at the base by an involucre of 3—6 scale-like bracts, composed of numerous sessile anthers, imbricated in many ranks and surmounted by crest-like nearly orbicular connectives; the female subterminal or lateral, their scales in the axils of non-accrescent bracts. Fruit a woody cone maturing at the end of the second or rarely of the third season, composed of the hardened and woody scales of the flower more or less thickened on the exposed surface (the apophysis), with the ends of the growth of the previous year appearing as terminal or dorsal brown protuberances or scars (the umbo). Seeds usually obovoid, shorter or longer than their wings or rarely wingless; outer seed-coat crustaceous or thick, hard, and bony, the inner membranaceous; cotyledons 3—18, usually much shorter than the inferior radicle.

Pinus is widely distributed through the northern hemisphere from the Arctic Circle to the West Indies, the mountains of Central America, the Canary Islands, northern Africa, the Philippine Islands, and Sumatra. About sixty-six species are recognized. Of exotic species the so-called Scotch Pine, Pinus sylvestris L., of Europe and Asia, the Swiss Stone Pine, Pinus cembra L., and the Austrian Pine and other forms of Pinus nigra Arnold, from central and southern Europe, are often planted in the northeastern states, and Pinus Pinaster Ait., of the coast region of western France and the Mediterranean Basin is successfully cultivated in central and southern California. Pinus is the classical name of the Pine-tree.

The North American species can be conveniently grouped in two sections, Soft Pines and Pitch Pines.

SOFT PINES.

Wood soft, close-grained, light-colored, the sapwood thin and nearly white; sheaths of the leaf-clusters deciduous; leaves with one fibro-vascular bundle.

1. Pinus Strobus L. White Pine.

Leaves soft bluish green, whitened on the ventral side by 3—5 bands of stomata, 3′—5′ long, mostly turning yellow and falling in September in their second season, or persistent until the following June. Flowers: male yellow; female bright pink, with purple scale margins. Fruit fully grown in July of the second season, 4′—8′ long, opening and discharging its seeds in September; seeds narrowed at the ends, ¼′ long, red-brown mottled with black, about one fourth as long as their wings.

A tree, while young with slender horizontal or slightly ascending branches in regular whorls usually of 5 branches; at maturity often 100°, occasionally 220° high, with a tall straight stem 3°—4° or rarely 6° in diameter, when crowded in the forest with short branches forming a narrow head, or rising above its forest companions with long lateral branches sweeping upward in graceful curves, the upper branches ascending and forming a broad open irregular head, and slender branchlets coated at first with rusty tomentum, soon glabrous, and orange-brown in their first winter. Bark on young stems and branches thin, smooth, green tinged with red, lustrous during the summer, becoming 1′—2′ thick on old trunks and deeply divided by shallow fissures into broad connected ridges covered with small closely appressed purplish scales. Wood light, not strong, straight-grained, easily worked, light brown often slightly tinged with red; largely manufactured into lumber, shingles, and laths, used in construction, for cabinet-making, the interior finish of buildings, wooden ware, matches, and the masts of vessels.

Distribution. Newfoundland to Manitoba, southward through the northern states to Pennsylvania, northern and eastern (Belmont County) Ohio, northern Indiana, valley of the Rocky River near Oregon, Ogle County, Illinois, and central and southeastern Iowa, and along the Appalachian Mountains to eastern Kentucky and Tennessee and northern [4] Georgia; forming nearly pure forests on sandy drift soils, or more often in small groves scattered in forests of deciduous-leaved trees on fertile well-drained soil, also on the banks of streams, or on river flats, or rarely in swamps.

Largely planted as an ornament of parks and gardens in the eastern states, and in many European countries, where it grows with vigor and rapidity; occasionally used in forest planting in the United States.

2. Pinus monticola D. Don. White Pine.

Leaves blue-green, glaucous, whitened by 2—6 rows of ventral and often by dorsal stomata, mostly persistent 3 or 4 years. Flowers: male yellow; female pale purple. Fruit 5′—11′ long, shedding its seeds late in the summer or in early autumn; seeds narrowed at the ends, ⅓′ long, pale red-brown mottled with black, about one third as long as their wings.

A tree, often 100° or occasionally 150° high, with a trunk frequently 4°—5° or rarely 7°—8° in diameter, slender spreading slightly pendulous branches clothing young stems to the ground and in old age forming a narrow open often unsymmetrical pyramidal head, and stout tough branchlets clothed at first with rusty pubescence, dark orange-brown and puberulous in their first and dark red-purple and glabrous in their second season. Bark of young stems and branches thin, smooth, light gray, becoming on old trees ¾′—1½′ thick and divided into small nearly square plates by deep longitudinal and cross fissures, and covered by small closely appressed purple scales. Wood light, soft, not strong, close, straight-grained, light brown or red; sometimes manufactured into lumber, used in construction and the interior finish of buildings.

Distribution. Scattered through mountain forests from the basin of the Columbia River in British Columbia to Vancouver Island; on the mountains of northern Washington to the western slopes of the Rocky Mountains of northern Montana; on the coast ranges of Washington and Oregon; and on the Cascade and Sierra Nevada ranges southward to the Kern River valley, California; most abundant and of its greatest value in northern Idaho on the bottom-lands of streams tributary to Lake Pend Oreille; reaching the sea-level on the southern shores of the Straits of Fuca and elevations of 10,000° on the California Sierras.

Often planted as an ornamental tree in Europe, and occasionally in the eastern United States where it grows more vigorously than any other Pine-tree of western America.

3. Pinus Lambertiana Dougl. Sugar Pine.

Leaves stout, rigid, 3½′—4′ long, marked on the two faces by 2—6 rows of stomata; deciduous during their second and third years. Flowers: male light yellow; female pale green. Fruit fully grown in August and opening in October, 11′—18′ or rarely 21′ long; seeds ½′—⅝′ long, dark chestnut-brown or nearly black, and half the length of their firm dark brown obtuse wings broadest below the middle and ½′ wide.

A tree, in early life with remote regular whorls of slender branches often clothing the stem to the ground and forming an open narrow pyramid; at maturity 200°-220° high, with a trunk 6°—8° or occasionally 12° in diameter, a flat-topped crown frequently 60° or 70° across of comparatively slender branches sweeping outward and downward in graceful curves, and stout branchlets coated at first with pale or rufous pubescence, dark orange-brown during their first winter, becoming dark purple-brown. Bark on young stems and branches thin, smooth, dark green, becoming on old trunks 2′—3′ thick and deeply and irregularly divided into long thick plate-like ridges covered with large loose rich purple-brown or cinnamon-red scales. Wood light, soft, straight-grained, light red-brown; [6] largely manufactured into lumber and used for the interior finish of buildings, woodwork, and shingles. A sweet sugar-like substance exudes from wounds made in the heartwood.

Distribution. Mountain slopes and the sides of ravines and cañons; western Oregon from the valley of the north branch of the Santiam River southward on the Cascade and coast ranges; California along the northern and coast ranges to Sonoma County; along the western slopes of the Sierra Nevada, where it grows to its greatest size at elevations between 3000° and 7000°; reappearing on the Santa Lucia Mountains of the coast ranges; and on the high mountains in the southwestern part of the state from Santa Barbara County southward usually at elevations of 5000°—7000° above the sea; and on the San Pedro Mártir Mountains in Lower California.

Occasionally planted as an ornamental tree in western Europe and in the eastern states, the Sugar Pine has grown slowly in cultivation and shows little promise of attaining the large size and great beauty which distinguish it in its native forests.

4. Pinus flexilis James. Rocky Mountain White Pine.

Leaves stout, rigid, dark green, marked on all sides by 1—4 rows of stomata, 1½′—3′ long, deciduous in their fifth and sixth years. Flowers: male reddish; female clustered, bright red-purple. Fruit subcylindric, horizontal or slightly declining, green or rarely purple at maturity, 3′—10′ long, with narrow and more or less reflexed scales opening at maturity; seeds compressed, ⅓′—½′ long, dark red-brown mottled with black, with a thick shell produced into a narrow margin, their wings about ¹/₁₂′ wide, generally persistent on the scale after the seed falls.

A tree, usually 40°—50°, occasionally 80° high, with a short trunk 2°—5° in diameter, stout long-persistent branches ultimately forming a low wide round-topped head, and stout branchlets orange-green and covered at first with soft fine pubescence, usually soon glabrous and darker colored; at high elevations often a low spreading shrub. Bark of young stems and branches thin, smooth, light gray or silvery white, becoming on old trunks 1′—2′ thick, dark brown or nearly black, and divided by deep fissures into broad ridges broken into nearly square plates covered by small closely appressed scales. Wood light, soft, close-grained, pale clear yellow, turning red with exposure; occasionally manufactured into lumber.

Distribution. Eastern slope of the Rocky Mountains from Alberta to western Texas and westward on mountain ranges at elevations of 5000° to 12,000° to Montana, and southern California, reaching the western slopes of the Sierra Nevada at the head of King’s River near the summit of San Gorgonio Mountain and in Snow Cañon, San Bernardino Range; usually scattered singly or in small groves; forming open forests on the eastern foothills of the Rocky Mountains of Montana and on the ranges of central Nevada; attaining its largest size on those of northern New Mexico and Arizona.

5. Pinus albicaulis Engelm. White Pine.

Leaves stout, rigid, slightly incurved, dark green, marked by 1—3 rows of dorsal stomata, clustered at the ends of the branches, 1½′—2½′ long, persistent for from five to eight years. Flowers opening in July, scarlet. Fruit ripening in August, oval or subglobose, horizontal, [7] sessile, dark purple, 1½′—3′ long, with scales thickened, acute, often armed with stout pointed umbos, remaining closed at maturity; seeds wingless, acute, subcylindric or flattened on one side, ⅓′—½′ long, ⅓′ thick, with a thick dark chestnut-brown hard shell.

A tree, usually 20°—30° or rarely 60° high, generally with a short trunk 2°—4° in diameter, stout very flexible branches, finally often standing nearly erect and forming an open very irregular broad head, and stout dark red-brown or orange-colored branchlets puberulous for two years or sometimes glabrous; at high elevations often a low shrub, with wide-spreading nearly prostrate stems. Bark thin, except near the base of old trunks and broken by narrow fissures into thin narrow brown or creamy white plate-like scales. Wood light, soft, close-grained, brittle, light brown. The large sweet seeds are gathered and eaten by Indians.

Distribution. Alpine slopes and exposed ridges between 5000° and 12,000° elevation, forming the timber-line on many mountain ranges from latitude 53° north in the Rocky Mountains and British Columbia, southward to the Wind River and Salt River Ranges, Wyoming, the mountains of eastern Washington and Oregon, the Cascade Range, the mountains of northern California and the Sierra Nevada to Mt. Whitney.

6. Pinus Balfouriana Balf. Foxtail Pine.

Leaves stout, rigid, dark green and lustrous on the back, pale and marked on the ventral faces by numerous rows of stomata, 1′—1½′ long, persistent for ten or twelve years. Flowers: male dark orange-red; female dark purple. Fruit 3½′—5′ long, with scales armed with minute incurved prickles, dark purple, turning after opening dark red or mahogany color; seeds full and rounded at the apex, compressed at the base, pale, conspicuously mottled with dark purple, ⅓′ long, their wings narrowed and oblique at the apex, about 1′ long and ¼′ wide.

A tree, usually 30°—40° or rarely 90° high, with a trunk generally 1°—2° or rarely 5° in diameter, short stout branches forming an open irregular pyramidal picturesque head, and long rigid more or less spreading puberulous, soon glabrous, dark orange-brown ultimately dark gray-brown or nearly black branchlets, clothed only at the extremities with the long dense brush-like masses of foliage. Bark thin, smooth, and milky white on the stems and branches of young trees, becoming on old trees sometimes ¾′ thick, dark red-brown, deeply divided into broad flat ridges. [8] broken into nearly square plates separating on the surface into small closely appressed scales. Wood light, soft and brittle, pale reddish brown.

Distribution. California, on rocky slopes and ridges, forming scattered groves on Scott Mountain, Siskiyou County, at elevations of 5000°—6000°; on the mountains at the head of the Sacramento River; on Mt. Yolo Bally in the northern Coast Range, and on the southern Sierra Nevada up to elevations of 11,500°, growing here to its largest size and forming an extensive open forest on the Whitney Plateau east of the cañon of Kern River, and at the highest elevations often a low shrub, with wide-spreading prostrate stems.

7. Pinus aristata Engelm. Foxtail Pine. Hickory Pine.

Leaves stout or slender, dark green, lustrous on the back, marked by numerous rows of stomata on the ventral faces, 1′—1½′ long, often deciduous at the end of ten or twelve years or persistent four or five years longer. Flowers male dark orange-red; female dark purple. Fruit 3′—3½′ long, with scales armed with slender incurved brittle prickles nearly ¼′ long, dark purple-brown on the exposed parts, the remainder dull red, opening and scattering their seeds about the 1st of October; seeds nearly oval, compressed, light brown mottled with black, ¼′ long, their wings broadest at the middle, about ⅓′ long and ¼′ wide.

A bushy tree, occasionally 40°—50° high, with a short trunk 2°—3° in diameter, short stout branches in regular whorls while young, in old age growing very irregularly, the upper erect and much longer than the usually pendulous lower branches, and stout light orange-colored, glabrous, or at first puberulous, ultimately dark gray-brown or nearly black branchlets clothed at the ends with long compact brush-like tufts of foliage. Bark thin, smooth, milky white on the stems and branches of young trees, becoming on old trees ½′—¾′ thick, red-brown, and irregularly divided into flat connected ridges separating on the surface into small closely appressed scales. Wood light, soft, not strong, light red; occasionally used for the timbers of mines and for fuel.

Distribution. Rocky or gravelly slopes at the upper limit of tree growth and rarely below 8,000° above the sea from the outer range of the Rocky Mountains of Colorado to those of southern Utah, central and southern Nevada, southeastern California, and the San Francisco peaks of northern Arizona.

8. Pinus cembroides Zucc. Nut Pine. Piñon.

Leaves in 2 or 3-leaved clusters, slender, much incurved, dark green, sometimes marked by rows of stomata on the 3 faces, 1′—2′ long, deciduous irregularly during their third and fourth years. Flowers: male in short crowded clusters, yellow; female dark red. Fruit subglobose, 1′—2′ broad; seeds subcylindric or obscurely triangular, more or less compressed at the pointed apex, full and rounded at base, nearly black on the lower side and dark chestnut-brown on the upper, ½′—¾′ long, the margin of their outer coat adnate to the cone-scale.

A bushy tree, with a short trunk rarely more than a foot in diameter and a broad round-topped head, usually 15°—20° high, stout spreading branches, and slender dark orange-colored branchlets covered at first with matted pale deciduous hairs, dark brown and sometimes nearly black at the end of five or six years; in sheltered cañons on the mountains of Arizona and in Lower California occasionally 50° or 60° tall. Bark about ½′ thick, irregularly [9] divided by remote shallow fissures and separated on the surface into numerous large thin light red-brown scales. Wood light, soft, close-grained, pale clear yellow. The large oily seeds are an important article of food in northern Mexico, and are sold in large quantities in Mexican towns.

Distribution. Mountain ranges of central and southern Arizona, usually only above elevations of 6500°, often covering their upper slopes with open forests; in an isolated station on the Edwards Plateau on uplands and in cañons at the headwaters of the Frio and Nueces Rivers, Edwards and Kerr Counties, Texas; on the Sierra de Laguna, Lower California, and on many of the mountain ranges of northern Mexico; passing into the following varieties differing only in the number of the leaves in the leaf-clusters, and in their thickness.

Pinus cembroides var. Parryana Voss. Nut Pine. Piñon.

Leaves in 1—5 usually 4-leaved clusters, stout, incurved, pale glaucous green, marked on the three surfaces by numerous rows of stomata, 1¼′—1½′ long, irregularly deciduous, mostly falling in their third year.

A tree, 30°—40° high, with a short trunk occasionally 18′ in diameter, and thick spreading branches forming a compact regularpyramidal or in old age a low round-topped irregular head, and stout branchlets coated at first with soft pubescence, and light orange-brown. Bark ½′—¾′ thick, dark brown tinged with red, and divided by shallow fissures into broad flat connected ridges covered by thick closely appressed plate-like scales. Wood light, soft, close-grained, pale brown or yellow. The seeds form an important article of food for the Indians of Lower California.

Distribution. Arid mesas and low mountain slopes of Lower California southward to the foothills of the San Pedro Mártir Mountains, extending northward across the boundary of California to the desert slopes of the Santa Rosa Mountains, Riverside County, where it is common at elevations of 5000° above the sea-level.

Pinus cembroides var. edulis Voss. Nut Pine. Piñon.

Leaves in 2 or rarely in 3-leaved clusters, stout, semiterete or triangular, rigid, incurved, dark-green, marked by numerous rows of stomata, ¾′—1½′ long, deciduous during the third or not until the fourth or fifth year, dropping irregularly and sometimes persistent for eight or nine years.

A tree often 40°—50° high with a tall trunk occasionally 2° in diameter and short erect [10] branches forming a narrow head, or frequently with a short divided trunk and a low round-topped head of spreading branches, and thick branchlets orange color during their first and second years, finally becoming light gray or dark brown sometimes tinged with red. Bark ½′—¾′ thick and irregularly divided into connected ridges covered by small closely appressed light brown scales tinged with red or orange color. Wood light, soft, not strong, brittle, pale brown; largely employed for fuel and fencing, and as charcoal used in smelting; in western Texas occasionally sawed into lumber. The seeds form an important article of food among Indians and Mexicans, and are sold in the markets of Colorado and New Mexico.

Distribution. Eastern foothills of the outer ranges of the Rocky Mountains, from northern Colorado (Owl Cañon, Larimer County); to the extreme western part of Oklahoma (near Kenton, Cimarron County, G. W. Stevens) and to western Texas, westward to eastern Utah, southwestern Wyoming, and to northern and central Arizona; over the mountains of northern Mexico, and on the San Pedro Mártir Mountains, Lower California; often forming extensive open forests at the eastern base of the Rocky Mountains, on the Colorado plateau, and on many mountain ranges of northern and central Arizona up to elevations of 7000° above the sea.

Pinus cembroides var. monophylla Voss. Nut Pine. Piñon.

Leaves in 1 or 2-leaved clusters, rigid, incurved, pale glaucous green, marked by 18—20 rows of stomata, usually about 1½′ long, sometimes deciduous during their fourth and fifth seasons, but frequently persistent until their twelfth year.

A tree usually 15°—20°, occasionally 40°—50° high, with a short trunk rarely more than a foot in diameter and often divided near the ground into several spreading stems, short thick branches forming while the tree is young a broad rather compact pyramid, and in old age often pendulous and forming a low round-topped often picturesque head, and stout light orange-colored ultimately dark brown branchlets. Bark about ¾′ thick and divided by deep irregular fissures into narrow connected flat ridges broken on the surface into thin closely appressed light or dark brown scales tinged with red or orange color. Wood light, soft, weak, and brittle; largely used for fuel, and charcoal used in smelting. The seeds supply an important article of food to the Indians of Nevada and California.

Distribution. Dry gravelly slopes and mesas from the western base of the Wasatch Mountains of Utah, westward over the mountain ranges of Nevada to the eastern slopes of the southern Sierra Nevada, and to their western slope at the headwaters of the Tuolumne, Kings and Kern Rivers, and southward to northern Arizona and to the mountains [11] of southern California where it is common on the San Bernardino and San Jacinto Mountains between altitudes of 3500° and 7000°, and on the Sierra del Pinal, Lower California; often forming extensive open forests at elevations between 5000° and 7000°.

PITCH PINES.

Wood usually heavy, coarse-grained, generally dark-colored, with pale often thick sapwood; cones green at maturity (sometimes purple in 10 and 21) becoming various shades of brown; cone-scales more or less thickened, mostly armed; seeds shorter than their wings (except in 17 and 28); leaves with 2 fibro-vascular bundles.

9. Pinus leiophylla Schlecht. and Cham. Yellow Pine.

Leaves slender, pale glaucous green, marked by 6—8 rows of conspicuous stomata on each of the 3 sides, 2½′—4′ long, irregularly deciduous from their fourth season, their sheaths deciduous. Flowers: male yellow; female yellow-green. Fruit ovoid, horizontal or slightly declining, long-stalked, 1½′—2′ long, becoming light chestnut-brown and lustrous, maturing at the end of the third season, with scales only slightly thickened, their ultimately pale umbos armed with recurved deciduous prickles; seeds oval, rounded above and pointed below, about ⅛′ long, with a thin dark brown shell, their wings ⅓′ long and broadest near the middle.

A tree, rarely more than 40°—50° high, with a tall trunk sometimes 2° in diameter, stout slightly ascending branches forming a narrow open pyramidal or round-topped head of thin pale foliage, and slender bright orange-brown branchlets, soon becoming dull red-brown. Bark of old trunks ¾′—1½′ thick, dark reddish brown or sometimes nearly black, and deeply divided into broad flat ridges covered with thin closely appressed scales. Wood light, soft, not strong but durable, light orange color, with thick much lighter colored sapwood. Often forming coppice by the growth of shoots from the stump of cut trees.

Distribution. Mountain ranges of southern New Mexico and Arizona, usually at elevations between 6000° and 7000°; not common; more abundant on the Sierra Madre of northern Mexico and on several of the short ranges of Chihuahua and Sonora, and of a larger size in Mexico than in the United States.

10. Pinus ponderosa Laws. Yellow Pine. Bull Pine.

Leaves tufted at the ends of naked branches, in 2 or in 2 and 3-leaved clusters, stout, dark yellow-green, marked by numerous rows of stomata on the 3 faces, 5′—11′ long, mostly deciduous during their third season. Flowers: male yellow; female clustered or in pairs, dark red. Fruit ellipsoidal, horizontal or slightly declining, nearly sessile or short-stalked, 3′—6′ long, often clustered, bright green or purple when fully grown, becoming light reddish brown, with narrow scales much thickened at the apex and armed with slender prickles, mostly falling soon after opening and discharging their seeds, generally leaving the lower scales attached to the peduncle; seeds ovoid, acute, compressed at the apex, full and rounded below, ¼′ long, with a thin dark purple often mottled shell, their wings usually broadest below the middle, gradually narrowed at the oblique apex, 1′—1¼′ long, about 1′ wide.

A tree, sometimes 150°—230° high, with a massive stem 5°—8° in diameter, short thick many-forked often pendulous branches generally turned upward at the ends and forming [13] a regular spire-like head, or in arid regions a broader often round-topped head surmounting a short trunk, and stout orange-colored branchlets frequently becoming nearly black at the end of two or three years. Bark for 80—100 years broken into rounded ridges covered with small closely appressed scales, dark brown, nearly black or light cinnamon-red, on older trees becoming 2′—4′ thick and deeply and irregularly divided into plates sometimes 4°—5° long and 12′—18′ wide, and separating into thick bright cinnamon-red scales. Wood hard, strong, comparatively fine-grained, light red, with nearly white sapwood sometimes composed of more than 200 layers of annual growth; largely manufactured into lumber used for all sorts of construction, for railway-ties, fencing, and fuel.

Distribution. Mountain slopes, dry valleys, and high mesas from northwestern Nebraska and western Texas to the shores of the Pacific Ocean, and from southern British Columbia to Lower California and northern Mexico; extremely variable in different parts of the country in size, in the length and thickness of the leaves, size of the cones, and in the color of the bark. The form of the Rocky Mountains (var. scopulorum, Engelm.), ranging from Nebraska to Texas, and over the mountain ranges of Wyoming, eastern Montana and Colorado, and to northern New Mexico and Arizona, where it forms on the Colorado plateau with the species the most extensive Pine forests of the continent, has nearly black furrowed bark, rigid leaves in clusters of 2 or 3 and 3′—6′ long, and smaller cones, with thin scales armed with slender prickles hooked backward. More distinct is

Pinus ponderosa var. Jeffreyi Vasey.

This tree forms great forests about the sources of the Pitt River in northern California, [14] along the eastern slopes of the central and southern Sierra Nevada, growing often on the most exposed and driest ridges, and in southern California on the San Bernardino and San Jacinto ranges up to elevations of 7000° above the sea, on the Cuyamaca Mountains, and in Lower California on the Sierra del Pinal and the San Pedro Mártir Mountains.

A tree, 100° to nearly 200° high, with a tall massive trunk 4°—6° in diameter, covered with bright cinnamon-red bark deeply divided into large irregular plates, stiffer and more elastic leaves 4′—9′ long and persistent on the glaucous stouter branchlets for six to nine years, yellow-green staminate flowers, short-stalked usually purple cones 5′—15′ long, their scales armed with stouter or slender prickles usually hooked backward, and seeds often nearly ½′ long with larger wings.

Occasionally planted as an ornamental tree in eastern Europe, especially the variety Jeffreyi, which is occasionally successfully cultivated in the eastern states.

Pinus ponderosa var. arizonica Shaw. Yellow Pine.

Leaves tufted at the ends of the branches, in 3—5-leaved clusters, stout, rigid, dark green, stomatiferous on their 3 faces, 5′—7′ long, deciduous during their third season. Fruit ovoid, horizontal, 2′—2½′ long, becoming light red-brown, with thin scales much thickened at the apex and armed with slender recurved spines; seeds full and rounded below, slightly compressed towards the apex, ⅛′ long, with a thick shell, their wings broadest above the middle, about ⅓′ long and ¼′ wide.

A tree, 80°—100° high, with a tall straight massive trunk 3°—4° in diameter, thick spreading branches forming a regular open round-topped or narrow pyramidal head, and stout branchlets orange-brown and pruinose when they first appear, becoming dark gray-brown. Bark on young trunks dark brown or almost black and deeply furrowed, becoming on old trees 1½′—2′ thick and divided into large unequally shaped plates separating on the surface into thin closely appressed light cinnamon-red scales. Wood light, soft, not strong, rather brittle, light red or often yellow, with thick lighter yellow or white sapwood; in Arizona occasionally manufactured into coarse lumber.

Distribution. High cool slopes on the sides of cañons of the mountain ranges of southern Arizona at elevations between 6000° and 8000°, sometimes forming nearly pure forests; more abundant and of its largest size on the mountains of Sonora and Chihuahua.

11. Pinus palustris Mill. Long-leaved Pine. Southern Pine.

Leaves in crowded clusters, forming dense tufts at the ends of the branches, slender, flexible, pendulous, dark green, 8′—18′ long, deciduous at the end of their second year. Flowers in very early spring before the appearance of the new leaves, male in short dense clusters, dark rose-purple; female just below the apex of the lengthening shoot in pairs or in clusters of 3 or 4, dark purple. Fruit cylindric-ovoid, slightly curved, nearly sessile, horizontal or pendant, 6′—10′ long, with thin flat scales rounded at apex and armed with small [15] reflexed prickles, becoming dull brown; in falling leaving a few of the basal scales attached to the stem; seeds almost triangular, full and rounded on the sides, prominently ridged, about ½′ long, with a thin pale shell marked with dark blotches on the upper side, and wings widest near the middle, gradually narrowed to a very oblique apex, about 1¾′ long and ⁷/₁₆′ wide.

A tree, 100°—120° high, with a tall straight slightly tapering trunk usually 2°—2½° or occasionally 3° in diameter, stout slightly branched gnarled and twisted limbs covered with thin dark scaly bark and forming an open elongated and usually very irregular head one third to one half the length of the tree, thick orange-brown branchlets, and acute winter-buds covered by elongated silvery white lustrous scales divided into long spreading filaments forming a cobweb-like network over the bud. Bark of the trunk ¹/₁₆′—½′ thick, light orange-brown, separating on the surface into large closely appressed papery scales. Wood heavy, exceedingly hard, strong, tough, coarse-grained, durable, light red to orange color, with very thin nearly white sapwood; largely used as “southern pine” or “Georgia pine” for masts and spars, bridges, viaducts, railway-ties, fencing, flooring, the interior finish of buildings, the construction of railway-cars, and for fuel and charcoal. A large part of the naval stores of the world is produced from this tree, which is exceedingly rich in resinous secretions.

Distribution. Generally confined to a belt of late tertiary sands and gravels stretching along the coast of the Atlantic and Gulf states and rarely more than 125 miles wide, from southeastern Virginia to the shores of Indian River and the valley of the Caloosahatchee River, Florida, and along the Gulf coast to the uplands east of the Mississippi River, extending northward in Alabama to the southern foothills of the Appalachian Mountains and to central and western Mississippi (Hinds and Adams Counties); west of the Mississippi River to the valley of the Trinity River, Texas, and through eastern Texas and western Louisiana nearly to the northern borders of this state.

12. Pinus caribæa Morelet. Slash Pine. Swamp Pine.

Leaves stout, in crowded 2 and 3-leaved clusters, dark green and lustrous, marked by numerous bands of stomata on each face, 8′—12′ long, deciduous at the end of their second season. Flowers in January and February before the appearance of the new leaves, male in short crowded clusters, dark purple; female lateral on long peduncles, pink. Fruit ovoid or ovoid-conic, reflexed during its first year, pendant, 2′—6′ long, with thin flexible flat [16] scales armed with minute incurved or recurved prickles, becoming dark rich lustrous brown; seeds almost triangular, full and rounded on the sides, 1⅙′—1¼′ long, with a thin brittle dark gray shell mottled with black, and dark brown wings ¾′—1′ long, ¼′ wide, their thickened bases encircling the seeds and often covering a large part of their lower surface.

A tree, often 100° high, with a tall tapering trunk 2½°—3° in diameter, heavy horizontal branches forming a handsome round-topped head, and stout orange-colored ultimately dark branchlets. Bark ¾′—½′ thick, and separating freely on the surface into large thin scales. Wood heavy, exceedingly hard, very strong, durable, coarse-grained, rich dark orange color, with thick nearly white sapwood; manufactured into lumber and used for construction and railway-ties. Naval stores are largely produced from this tree.

Distribution. Coast region of South Carolina southward over the coast plain to the keys of southern Florida and along the Gulf coast to eastern Louisiana (Saint Tammany, Washington, southern Tangipahoa and eastern Livingston Parishes); common on the Bahamas, on the Isle of Pines, and on the lowlands of Honduras and eastern Guatemala; in the coast region of the southern states gradually replacing the Long-leaved Pine, Pinus palustris, Mill.

13. Pinus tæda L. Loblolly Pine. Old Field Pine.

Leaves slender, stiff, slightly twisted, pale green and somewhat glaucous, 6′—9′ long, marked by 10—12 rows of large stomata on each face, deciduous during their third year. Flowers opening from the middle of March to the first of May; male crowded in short spikes, yellow; female lateral below the apex of the growing shoot, solitary or clustered, short-stalked, yellow. Fruit oblong-conic to ovoid-cylindric, nearly sessile, 2′—6′ long, becoming light reddish brown, with thin scales rounded at the apex and armed with short stout straight or reflexed prickles, opening irregularly and discharging their seeds during the autumn and winter, and usually persistent on the branches for another year; seeds rhomboidal, full and rounded, ¼′ long, with a thin dark brown rough shell blotched with black, and produced into broad thin lateral margins, encircled to the base by the narrow border of their thin pale brown lustrous wing broadest above the middle, 1′ long, about ¼′ wide.

A tree, generally 80°—100° high, with a tall straight trunk usually about 2° but occasionally 5° in diameter, short thick much divided branches, the lower spreading, the upper ascending and forming a compact round-topped head, and comparatively slender glabrous branchlets brown tinged with yellow during their first season and gradually growing darker in their second year. Bark of the trunk ¾′—1½′ thick, bright red-brown, and irregularly [17] divided by shallow fissures into broad flat ridges covered with large thin closely appressed scales. Wood weak, brittle, coarse-grained, not durable, light brown, with orange-colored or often nearly white sapwood, often composing nearly half the trunk; largely manufactured into lumber, used for construction and the interior finish of buildings.

Distribution. Cape May, New Jersey, through southern Delaware and eastern Maryland and southward to the shores of Indian River and Tampa Bay, Florida, westward to middle North Carolina and through South Carolina and Georgia and the eastern Gulf states to the Mississippi River, extending into southern Tennessee and northeastern Mississippi; west of the Mississippi River from southern Arkansas and the southwestern part of Oklahoma through western Louisiana to the shores of the Gulf of Mexico, and through eastern Texas to the valley of the Colorado River; on the Atlantic coast often springing up on lands exhausted by agriculture; west of the Mississippi River one of the most important timber-trees, frequently growing in nearly pure forests on rolling uplands.

14. Pinus rigida Mill. Pitch Pine.

Leaves stout, rigid, dark yellow-green, marked on the 3 faces by many rows of stomata, 3′—5′ long, standing stiffly and at right angles with the branch, deciduous during their second year. Flowers: male in short crowded spikes, yellow or rarely purple; female often clustered and raised on short stout stems, light green more or less tinged with rose color. Fruit ovoid, acute at apex, nearly sessile, often clustered, 1′—3½′ long, becoming light brown, with thin flat scales armed with recurved rigid prickles, often remaining on the branches for ten or twelve years; seeds nearly triangular, full and rounded on the sides, ¼′ long, with a thin dark brown mottled roughened shell and wings broadest below the middle, gradually narrowed to the very oblique apex, ¾′ long, ⅓′ wide.

A tree, 50°—60° or rarely 100° high, with a short trunk occasionally 3° in diameter, thick contorted often pendulous branches covered with thick much roughened bark, forming a round-topped thick head, often irregular and picturesque, and [18] stout bright green branchlets becoming dull orange color during their first winter and dark gray-brown at the end of four or five years; often fruitful when only a few feet high. Bark of young stems thin and broken into plate-like dark red-brown scales, becoming on old trunks ¾′—1½′ thick, deeply and irregularly fissured, and divided into broad flat connected ridges separating on the surface into thick dark red-brown scales often tinged with purple. Wood light, soft, not strong, brittle, coarse-grained, very durable, light brown or red, with thick yellow or often white sapwood; largely used for fuel and in the manufacture of charcoal; occasionally sawed into lumber.

Distribution. Sandy plains and dry gravelly uplands, or less frequently in cold deep swamps; island of Mt. Desert, Maine, to the northern shores of Lake Ontario, and southward to southern Delaware and southern Ohio (Scioto County) and along the Appalachian Mountains to northern Georgia and to their western foothills in West Virginia, Kentucky, and Tennessee; very abundant in the coast region south of Massachusetts; sometimes forming pure forests in New Jersey and Pennsylvania.

Pinus rigida var. serotina Loud. Pond Pine. Marsh Pine.

Leaves in clusters of 3 or occasionally of 4, slender, flexuose, dark yellow-green, 6′—8′ long, marked by numerous rows of stomata on the 3 faces, deciduous during their third and fourth years. Flowers: male in crowded spikes, dark orange color; female clustered or in pairs on stout stems. Fruit subglobose to ovoid, full and rounded or pointed at apex, subsessile or short-stalked, horizontal or slightly declining, 2′—2½′ long, with thin nearly flat scales armed with slender incurved mostly deciduous prickles, becoming light yellow-brown at maturity, often remaining closed for one or two years and after opening long-persistent on the branches; seeds nearly triangular, often ridged below, full and rounded at the sides, ⅛′ long, with a thin nearly black roughened shell produced into a wide border, the wings broadest at the middle, gradually narrowed at the ends, ¾′ long, ¼′ wide.

A tree, usually 40°—50° or occasionally 70°—80° high, with a short trunk sometimes 3° but generally not more than 2° in diameter, stout often contorted branches more or less pendulous at the extremities, forming an open round-topped head, and slender branchlets dark green when they first appear, becoming dark orange color during their first winter and dark brown or often nearly black at the end of four or five years. Bark of the trunk ½′—¾′ thick, dark red-brown and irregularly divided by narrow shallow fissures into small plates separating on the surface into thin closely appressed scales. Wood very resinous, heavy, soft, brittle, coarse-grained, dark orange color, with thick pale yellow sapwood; occasionally manufactured into lumber.

Distribution. Low wet flats or sandy or peaty swamps; near Cape May, New Jersey, and southeastern Virginia southward near the coast to northern Florida and central Alabama.

15. Pinus radiata D. Don. Monterey Pine.

Leaves in 3, rarely in 2-leaved clusters, slender, bright rich green, 4′—6′ long, mostly deciduous during their third season. Flowers: male in dense spikes, yellow; female clustered, dark purple. Fruit ovoid, pointed at apex, very oblique at base, short-stalked, reflexed, 3′—7′ long, becoming deep chestnut-brown and lustrous, with scales much thickened and mammillate toward the base on the outer side of the cone, thinner on the inner side and at its apex, and armed with minute thickened incurved or straight prickles, long-persistent and often remaining closed on the branches for many years; seeds ellipsoidal, compressed, ¼′ long, with a thin brittle rough nearly black shell, their wings light brown, longitudinally striped, broadest above the middle, gradually narrowed and oblique at apex, 1′ long, ¼′ wide.

A tree, usually 40°—60° rarely 100°—115° high, with a tall trunk usually 1°—2° but occasionally 4½° in diameter, spreading branches forming a regular narrow open round-topped head, and slender branchlets light or dark orange color, at first often covered with a glaucous bloom, ultimately dark red-brown. Bark of the trunk 1½′—2′ thick, dark red-brown, and deeply divided into broad flat ridges broken on the surface into thick appressed plate-like scales. Wood light, soft, not strong, brittle, close-grained; occasionally used as fuel.

Distribution. In a narrow belt a few miles wide on the California coast from Pescadero to the shores of San Simeon Bay; in San Luis Obispo County near the village of Cambria; on the islands of Santa Rosa and Santa Cruz of the Santa Barbara group; and on Guadaloupe Island off the coast of Lower California; most abundant and of its largest size on Point Pinos south of the Bay of Monterey, California.

Largely planted for the decoration of parks in western and southern Europe, occasionally planted in the southeastern states and in Mexico, Australia, New Zealand, and other regions with temperate climates, and more generally in the coast region of the Pacific states from Vancouver Island southward than any other Pine-tree.

16. Pinus attenuata Lemm. Knob-cone Pine.

Leaves slender, firm and rigid, pale yellow or bluish green, marked by numerous rows of stomata on their 3 faces, 3′—7′, usually 4′—5′ long. Flowers: male orange-brown; female fascicled, often with several fascicles on the shoot of the year. Fruit elongated, conic, pointed, very oblique at base by the greater development of the scales on the outer side, whorled, short-stalked, strongly reflexed and incurved, 3′—6′ long, becoming light yellow-brown, with thin flat scales rounded at apex, those on the outer side being enlarged into prominent transversely flattened knobs armed with thick flattened incurved spines, those on the inner side of the cone slightly thickened and armed with minute recurved prickles, persistent on the stems and branches for thirty or forty years, sometimes becoming completely imbedded in the bark of old trunks, and usually not opening until the death of the tree; seeds ellipsoidal, compressed, acute at apex, ¼′ long, with a thin oblique shell, their wings broadest at the middle, gradually narrowed to the ends, 1¼′ long, ⅓′ wide.

A tree, usually about 20° high, with a trunk a foot in diameter, and often fruitful when [20] only 4° or 5° tall; occasionally growing to the height of 80°—100°, with a trunk 2½° thick, and frequently divided above the middle into two ascending stems, slender branches arrangedin regular whorls while the tree is young, and in old age forming a narrow round-topped straggling head of sparse thin foliage, and slender dark orange-brown branchlets growing darker during their second season. Bark of young stems and branches thin, smooth, pale brown, becoming at the base of old trunks ¼′—½′ thick and dark brown often tinged with purple, slightly and irregularly divided by shallow fissures and broken into large loose scales. Wood light, soft, not strong, brittle, coarse-grained, light brown, with thick sapwood sometimes slightly tinged with red.

Distribution. Dry mountain slopes from the valley of the Mackenzie River in Oregon over the mountains of southwestern Oregon, where it is most abundant and grows to its largest size, often forming pure forests over large areas, southward along the western slopes of the Cascade Mountains; in California on the northern cross ranges, the coast ranges from Trinity to Sonoma Counties, the western slopes of the Sierra Nevada to Mariposa County, and over the southern coast ranges from Santa Cruz to the dry arid southern slopes of the San Bernardino Mountains, where it forms a belt between City and East Twin Creeks at an altitude of 3500° above the sea.

17. Pinus Sabiniana Dougl. Digger Pine. Bull Pine.

Leaves stout, flexible, pendant, pale blue-green, marked on each face with numerous rows of pale stomata, 8′—12′ long, deciduous usually in their third and fourth years. Flowers: male yellow; female on stout peduncles, dark purple. Fruit oblong-ovoid, full and rounded at base, pointed, becoming light reddish brown, 6′—10′ long, long-stalked, pendulous, the scales narrowed into a stout incurved sharp hook, strongly reflexed toward the base of the cone and armed with spur-like incurved spines; seeds full and rounded below, somewhat compressed toward the apex, ¾′ long, ⅓′ wide, dark brown or nearly black, with a thick hard shell, encircled by their wings much [21] thickened on the inner rim, obliquely rounded at the broad apex and about ½ length of nuts.

A tree, usually 40°—50° but occasionally 80° high, with a trunk 3°—4° in diameter, divided generally 15°—20° above the ground into 3 or 4 thick secondary stems, clothed with short crooked branches pendant below and ascending toward the summit of the tree, and forming an open round-topped head remarkable for the sparseness of its foliage, and stout pale glaucous branchlets, becoming dark brown or nearly black during their second season. Bark of the trunk 1½′—2′ thick, dark brown slightly tinged with red or nearly black and deeply and irregularly divided into thick connected ridges covered with small closely appressed scales. Wood light, soft, not strong, close-grained, brittle, light brown or red with thick nearly white sapwood. Abietine, a nearly colorless aromatic liquid with the odor of oil of oranges, is obtained by distilling the resinous juices. The large sweet slightly resinous seeds formed an important article of food for the Indians of California.

Distribution. Scattered singly or in small groups over the dry foothills of western California, ranging from 500° up to 4000° above the sea-level and from the southern slopes of the northern cross ranges to the Tehachapi Mountains and the Sierra de la Liebre; most abundant and attaining its largest size on the eastern foothills of the Sierra Nevada near the centre of the state at elevations of about 2000°; here often the most conspicuous feature of the vegetation.

18. Pinus Coulteri D. Don. Pitch Pine.

Leaves tufted at the ends of the branches, stout, rigid, dark blue-green, marked by numerous bands of stomata on the 3 faces, 6′—12′ long, deciduous during their third and fourth seasons. Flowers: male yellow; female dark reddish brown. Fruit oblong-conic, short-stalked and pendant, 10′—14′ long, becoming light yellow-brown, with thick broad scales terminating in a broad, flat, incurved, hooked claw ½′—1½′ long, gradually opening in the autumn and often persistent on the branches for several years; seeds ellipsoidal, compressed, ½′ long, ¼′—⅓′ wide, dark chestnut-brown, with a thick shell, inclosed by their wings, broadest above the middle, oblique at apex, nearly 1′ longer than the seed, about ⅝′ wide.

A tree, 40°—90° high, with a trunk 1°—2½° in diameter, thick branches covered with dark scaly bark, long and mostly pendulous below, short and ascending above, and forming a loose unsymmetrical often picturesque head, and very stout branchlets dark orange-brown at first, becoming sometimes nearly black at the end of three or four years. Bark of the [22] trunk 1½′—2′ thick, dark brown or nearly black and deeply divided into broad rounded connected ridges covered with thin closely appressed scales. Wood light, soft, not strong, brittle, coarse-grained, light red, with thick nearly white sapwood; occasionally used for fuel. The seeds were formerly gathered in large quantities and eaten by the Indians of southern California.

Distribution. Scattered singly or in small groves through coniferous forests on the dry slopes and ridges of the coast ranges of California at elevations of 3000°—6000° above the sea, from Mount Diablo and the Santa Lucia Mountains to the San Bernardino and Cuyamaca Mountains; and on the Sierra del Pinal, Lower California; most abundant on the San Bernardino and San Jacinto ranges at elevations of about 5000°.

19. Pinus resinosa Ait. Red Pine. Norway Pine.

Leaves slender, soft and flexible, dark green and lustrous, 5′—6′ long, obscurely marked on the ventral faces by bands of minute stomata, deciduous during their fourth and fifth seasons. Flowers: male in dense spikes, dark purple; female terminal, short-stalked, scarlet. Fruit ovoid-conic, subsessile, 2′—2¼′ long, with thin slightly concave scales, unarmed, becoming light chestnut-brown and lustrous at maturity; shedding their seeds early in the autumn and mostly persistent on the branches until the following summer; seeds oval, compressed, ⅛′ long, with a thin dark chestnut-brown more or less mottled shell and wings broadest below the middle, oblique at apex, ¾′ long, ¼′—⅓′ broad.

A tree, usually 70°—80° or occasionally 120° high, with a tall straight trunk 2°—3° or rarely 5° in diameter, thick spreading more or less pendulous branches clothing the young stems to the ground and forming a broad irregular pyramid, and in old age an open round-topped picturesque head, and stout branchlets at first orange color, finally becoming light reddish brown. Bark of the trunk ¾′—1¼′ thick and slightly divided by shallow fissures into broad flat ridges covered by thin loose light red-brown scales. Wood light, hard, very close-grained, pale red, with thin yellow often nearly white sapwood; largely used in the construction of bridges and buildings, for piles, masts, and spars. The bark is occasionally used for tanning leather.

Distribution. Light sandy loam or dry rocky ridges, usually forming groves rarely more than a few hundred acres in extent and scattered through forests of other Pines and deciduous-leaved trees; occasionally on sandy flats forming pure forests; Nova Scotia to Lake St. John, westward through Quebec and central Ontario to the valley of the Winnipeg River, and southward to eastern Massachusetts, the mountains of northern Pennsylvania, and to central and southwestern (Port Huron) Michigan, Wisconsin, and Minnesota, most abundant, and growing to its largest size in the northern parts of these states; rare and local in eastern Massachusetts and southward.

Often planted for the decoration of parks, and the most desirable as an ornamental tree of the Pitch Pines which flourish in the northern states.

20. Pinus contorta Loud. Scrub Pine.

Leaves dark green, slender, 1′—1½′ long, marked by 6—10 rows of stomata on each face, mostly persistent 4—6 years. Flowers orange-red: male in short crowded spikes; female clustered or in pairs on stout stalks. Fruit ovoid to subcylindric, usually very oblique at base, horizontal or declining, often clustered, ¾′—2′ long, with thin slightly concave scales armed with long slender more or less recurved often deciduous prickles, and toward the base of the cone especially on the upper side developed into thick mammillate knobs, becoming light yellow-brown and lustrous, sometimes opening and losing their seeds as soon as ripe, or remaining closed on the branches and preserving the vitality of their seeds for many years; seeds oblique at apex, acute below, about ⅙′ long, with a thin brittle dark red-brown shell mottled with black and wings widest above the base, gradually tapering toward the oblique apex, ½′ long.

A tree, sometimes fertile when only a few inches high, usually 15°—20° or occasionally 30° tall, with a short trunk rarely more than 18′ in diameter, comparatively thick branches forming a round-topped compact and symmetrical or an open picturesque head, and stout branchlets light orange color when they first appear, finally becoming dark red-brown or occasionally almost black. Bark of the trunk ¾′—1′ thick, deeply and irregularly divided by vertical and cross fissures into small oblong plates covered with closely appressed dark red-brown scales tinged with purple or orange color. Wood light, hard, strong although brittle, coarse-grained, light brown tinged with red, with thick nearly white sapwood; occasionally used for fuel.

Distribution. Coast of Alaska, usually in sphagnum-covered bogs southward in the immediate neighborhood of the coast to the valley of the Albion River, Mendocino County, California; south of the northern boundary of the United States generally inhabiting sand dunes and barrens or occasionally near the shores of Puget Sound the margins of tide pools and deep wet swamps; spreading inland and ascending the coast ranges and western slopes of the Cascade Mountains, where it is not common and where it gradually changes its habit and appearance, the thick deeply furrowed bark of the coast form being found only near the ground, while the bark higher on the stems is thin, light-colored, and inclined to separate into scales, and the leaves are often longer and broader. This is

Pinus contorta var. latifolia S. Wats. Lodge-pole Pine.

Leaves yellow-green, usually about 2′ long, although varying from 1′—3′ in length and from ¹/₁₆′ to nearly ⅛′ in width. Fruit occasionally opening as soon as ripe but usually remaining closed and preserving the vitality of the seeds sometimes for twenty years.

A tree, usually 70°—80° but often 150° high, with a trunk generally 2°—3° but occasionally 5°—6° in diameter, slender much-forked branches frequently persistent nearly to the base of the stem, light orange-colored during their early years, somewhat pendulous below, ascending near the top of the tree, and forming a narrow pyramidal spire-topped head. [24] Bark of the trunk rarely more than ¼′ thick, close and firm, light orange-brown and covered by small thin loosely appressed scales. Wood light, soft, not strong, close, straight-grained and easily worked, not durable, light yellow or nearly white, with thin lighter colored sapwood; occasionally manufactured into lumber; also used for railway-ties, mine-timbers, and for fuel.

Distribution. Common on the Yukon hills in the valley of the Yukon River; on the interior plateau of northern British Columbia and eastward to the eastern foothills of the Rocky Mountains, covering with dense forests great areas in the basin of the Columbia River; forming forests on both slopes of the Rocky Mountains of Montana; on the Yellowstone plateau at elevations of 7000°-8000°; common on the mountains of Wyoming, and extending southward to southern Colorado; the most abundant coniferous tree of the northern Rocky Mountain region; common on the ranges of eastern Washington and Oregon, on the mountains of northern California, and southward along the Sierra Nevada, where it attains its greatest size and beauty in alpine forests at elevations between 8000° and 9500°; in southern California the principal tree at elevations between 7000° and 10,000° on the high peaks of the San Bernardino and San Jacinto Mountains; on the upper slopes of the San Pedro Mártir Mountains, Lower California.

21. Pinus Banksiana Lamb. Gray Pine. Jack Pine.

Leaves in remote clusters, stout, flat or slightly concave on the inner face, at first light yellow-green, soon becoming dark green, ¾′—1¼′ long, gradually and irregularly deciduous in their second or third year. Flowers: male in short crowded clusters, yellow; female clustered, dark purple, often with 2 clusters produced on the same shoot. Fruit oblong-conic, acute, oblique at base, sessile, usually erect and strongly incurved, 1½′—2′ long, dull purple or green when fully grown, becoming light yellow and lustrous, with thin stiff [25] scales often irregularly developed, and armed with minute incurved often deciduous prickles; seeds nearly triangular, full and rounded on the sides, ¹/₁₂′ long, with an almost black roughened shell and wings broadest at the middle, full and rounded at apex, ⅓′ long, ⅛′ wide.

A tree, frequently 70° high, with a straight trunk sometimes free of branches for 20°—30° and rarely exceeding 2° in diameter, long spreading branches forming an open symmetrical head, and slender tough flexible pale yellow-green branchlets turning dark purple during their first winter and darker the following year; often not more than 20°—30° tall, with a stem 10′—12′ in diameter; generally fruiting when only a few years old; sometimes shrubby with several low slender stems. Bark of the trunk thin, dark brown slightly tinged with red, very irregularly divided into narrow rounded connected ridges separating on the surface into small thick closely appressed scales. Wood light, soft, not strong, close-grained, clear pale brown or rarely orange color, with thick nearly white sapwood; used for fuel and occasionally for railway-ties and posts; occasionally manufactured into lumber.

Distribution. From Nova Scotia to the valley of the Athabasca River and down the Mackenzie to about latitude 65° north, ranging southward to the coast of Maine, northern New Hampshire and Vermont, the Island of Nantucket (Wauwinet, J. W. Harshburger), northern New York, the shores of Saginaw Bay, Michigan, the southern shores of Lake Michigan in Illinois, the valley of the Wisconsin River, Wisconsin, and central and southeastern Minnesota (with isolated groves in Root River valley, near Rushford, Fillmore County); abundant in central Michigan, covering tracts of barren lands; common and of large size in the region north of Lake Superior; most abundant and of its greatest size west of Lake Winnipeg and north of the Saskatchewan, here often spreading over great areas of sandy sterile soil.

22. Pinus glabra Walt. Spruce Pine. Cedar Pine.

Leaves soft, slender, dark green, 1½′—3′ long, marked by numerous rows of stomata, deciduous at the end of their second and in the spring of their third year. Flowers: male in short crowded clusters, yellow; female raised on slender slightly ascending peduncles. Fruit single or in clusters of 2 or 3, reflexed on short stout stalks, subglobose to oblong-ovoid, ½′—2′ long, becoming reddish brown and rather lustrous, with thin slightly concave scales armed with minute straight or incurved usually deciduous prickles; seeds nearly triangular, full and rounded on the sides, ⅛′ long, with a thin dark gray shell mottled with black and wings broadest below the middle, ⅝′ long, ¼′ wide.

A tree, usually 80°—100° or occasionally 120° high, with a trunk 2°—2½° or rarely 3½° in diameter, comparatively small horizontal branches, and slender flexible branchlets at first light red more or less tinged with purple, ultimately dark reddish brown. Bark of young trees and upper trunks smooth pale gray becoming on old stems ½′—¾′ thick, slightly and irregularly divided by shallow fissures into flat connected ridges. Wood light, soft, not strong, brittle, close-grained, light brown, with thick nearly white sapwood; occasionally used for fuel and rarely manufactured into lumber.

Distribution. Valley of the lower Santee River, South Carolina to middle and northwestern Florida; banks of the Alabama River, Dallas County, Alabama; eastern and southwestern Mississippi, and sandy banks of streams in northeastern Louisiana; usually growing singly or in small groves; attaining its largest size and often occupying areas of considerable extent in northwestern Florida.

23. Pinus echinata Mill. Yellow Pine. Short-leaved Pine.

Leaves in clusters of 2 and of 3, slender, flexible, dark blue-green, 3′—5′ long, beginning to fall at the end of their second season and dropping irregularly until their fifth year. Flowers: male in short crowded clusters, pale purple; female in clusters of 2 or 3 on stout ascending stems, pale rose color. Fruit ovoid to oblong-conic, subsessile and nearly horizontal or short-stalked and pendant, generally clustered, 1½′—2½′ long, becoming dull brown, with thin scales nearly flat below and rounded at the apex, armed with short straight or somewhat recurved frequently deciduous prickles; seeds nearly triangular, full and rounded on the sides, about ³/₁₆′ long, with a thin pale brown hard shell conspicuously mottled with black, their wings broadest near the middle, ½′ long, ⅛′ wide.

A tree, usually 80°—100° occasionally 120° high, with a tall slightly tapering trunk 3°—4° in diameter, a short pyramidal truncate head of comparatively slender branches, and stout brittle pale green or violet-colored branchlets covered at first with a glaucous bloom, becoming dark red-brown tinged with purple before the end of the first season, their bark beginning in the third year to separate into large scales. Bark of the trunk ¾′—1′ thick and broken into large irregularly shaped plates covered with small closely appressed light cinnamon-red scales. Wood very variable in quality, and in the thickness of the nearly white sapwood, heavy, hard, strong and usually coarse-grained, orange-colored or yellow-brown; largely manufactured into lumber.

Distribution. Long Island (near Northport), and Staten Island, New York, and southern Pennsylvania to northern Florida, and westward through the Gulf states to eastern Texas, through Arkansas to southwestern Oklahoma (near Page, Leflore County, G. W. Stevens) and to southern Missouri and southwestern Illinois and to eastern Tennessee and western West Virginia; most abundant and of its largest size west of the Mississippi River.

24. Pinus virginiana Mill. Jersey Pine. Scrub Pine.

Leaves in remote clusters, stout, gray-green, 1½′—3′ long, marked by many rows of minute stomata, gradually and irregularly deciduous during their third and fourth years. Flowers: male in crowded clusters, orange-brown; female on opposite spreading peduncles near the middle of the shoots of the year, generally a little below and alternate with 1 or 2 [27] lateral branchlets, pale green, 2′—3′ long, the scale-tips tinged with rose color. Fruit ovoid-conic, often reflexed, dark red-brown and lustrous, with thin nearly flat scales, and stout or slender persistent prickles, opening in the autumn and slowly shedding their seeds, turning dark reddish brown and remaining on the branches for three or four years; seeds nearly oval, full and rounded, ¼′ long, with a thin pale brown rough shell, their wings broadest at the middle, ⅓′ long, about ⅛′ wide.

A tree, usually 30°—40° high, with a short trunk rarely more than 18′ in diameter, long horizontal or pendulous branches in remote whorls forming a broad open often flat-topped pyramid, and slender tough flexible branchlets at first pale green or green tinged with purple and covered with a glaucous bloom, becoming purple and later light gray-brown; toward the western limits of its range a tree frequently 100° tall, with a trunk 2½°—3° in diameter. Bark of the trunk ¼′—½′ thick, broken by shallow fissures into flat plate-like scales separating on the surface into thin closely appressed dark brown scales tinged with red. Wood light, soft, not strong, brittle, coarse-grained, durable in contact with the soil, light orange color, with thick nearly white sapwood; often used for fuel and occasionally manufactured into lumber.

Distribution. Middle and southern New Jersey; Plymouth, Luzerne County, and central, southern and western Pennsylvania to Columbia County, Georgia, Dallas County, Alabama (near Selma, T. G. Harbison), and to the hills of northeastern Mississippi (Bear Creek near its junction with the Tennessee River, E. N. Lowe), through eastern and middle Tennessee to western Kentucky and to southeastern and southern (Scioto County) Ohio, and southern Indiana; usually small in the Atlantic states and only on light sandy soil, spreading rapidly over exhausted fields; of its largest size west of the Alleghany Mountains on the low hills of southern Indiana.

25. Pinus clausa Sarg. Sand Pine. Spruce Pine.

Leaves slender, flexible, dark green, 2′—3½′ long, marked by 10—20 rows of stomata, deciduous during their third and fourth years. Flowers: male in short crowded spikes, dark orange color; female lateral on stout peduncles. Fruit elongated ovoid-conic, often oblique at base, usually clustered and reflexed, 2′—3½′ long, nearly sessile or short-stalked, with convex scales armed with short stout straight or recurved prickles, becoming dark yellow-brown in autumn; some of the cones opening at once, others remaining closed for three or four years before liberating their seeds, ultimately turning to an ashy gray color; others still unopened becoming enveloped in the growing tissues of the stem and branches and finally entirely covered by them; seeds nearly triangular, compressed, ¼′ long, with a [28] black slightly roughened shell, their wings widest near or below the middle, ¾′ long, about ¼′ wide.

A tree, usually 15°—20° high, with a stem rarely a foot in diameter, generally clothed to the ground with wide-spreading branches forming a bushy flat-topped head, and slender tough flexible branchlets, pale yellow-green when they first appear, becoming light orange-brown and ultimately ashy gray; occasionally growing to the height of 70°—80° with a trunk 2° in diameter. Bark on the lower part of the trunk ⅓′—½′ thick, deeply divided by narrow fissures into irregularly shaped generally oblong plates separating on the surface into thin closely appressed bright red-brown scales; on the upper part of the trunk and on the branches thin, smooth, ashy gray. Wood light, soft, not strong, brittle, light orange color or yellow, with thick nearly white sapwood; occasionally used for the masts of small vessels.

Distribution. Coast of the Gulf of Mexico from southern Alabama to Peace Creek, western Florida; eastern Florida from the neighborhood of St. Augustine to New River, Dade County, covering sandy wind-swept plains near the coast; growing to its largest size and most abundant in the interior of the peninsula (Lake and Orange Counties).

26. Pinus muricata D. Don. Prickle-cone Pine.

Leaves in crowded clusters, thick, rigid, dark yellow-green, 4′—6′ long, beginning to fall in their second year. Flowers: male in elongated spikes, orange-colored; female short-stalked, whorled, 2 whorls often being produced on the shoot of the year. Fruit ovoid, oblique at base, sessile, in clusters of 3—5 or sometimes of 7, 2′—3½′ but usually about 3′ long, becoming light chestnut-brown and lustrous, with scales much thickened on the [29] outside of the cone, those toward its base produced into stout incurved knobs sometimes armed with stout flattened spur-like often incurved spines, and on the inside of the cone slightly flattened and armed with stout or slender straight prickles; often remaining closed for several years and usually persistent on the stem and branches during the entire life of the tree without becoming imbedded in the wood; seeds nearly triangular, ¼′ long, with a thin nearly black roughened shell, their wings broadest above the middle, oblique at apex, nearly 1′ long, ⅛′ wide.

A tree, usually 40°—50° but occasionally 90° high, with a trunk 2°—3° in diameter, thick spreading branches covered with dark scaly bark, in youth forming a regular pyramid, and at maturity a handsome compact round-topped head of dense tufted foliage, and stout branchlets dark orange-green at first, turning orange-brown more or less tinged with purple. Bark of the lower part of the trunk often 4′—6′ thick and deeply divided into long narrow rounded ridges roughened by closely appressed dark purplish brown scales. Wood light, very strong, hard, rather coarse-grained, light brown, with thick nearly white sapwood; occasionally manufactured into lumber.

Distribution. California coast region from Mendocino County southward, usually in widely separated localities to Point Reyes Peninsula, north of the Bay of San Francisco, and from Monterey to Coon Creek, San Luis Obispo County; in Lower California on Cedros Island and on the west coast between Ensenada and San Quentin; of its largest size and the common Pine-tree on the coast of Mendocino County.

27. Pinus pungens Lamb. Table Mountain Pine. Hickory Pine.

Leaves in crowded clusters, rigid, usually twisted, dark blue-green, 1¼′—2½′ long, deciduous during their second and third years. Flowers: male in elongated loose spikes, yellow; female clustered, long-stalked. Fruit ovoid-conic, oblique at base by the greater development of the scales on the outer than on the inner side, sessile, reflexed, in clusters usually of 3 or 4, or rarely of 7 or 8, 2′—3½′ long, becoming light brown and lustrous, with thin tough scales armed with stout hooked curved spines produced from much thickened mammillate knobs, opening as soon as ripe and gradually shedding their seeds, or often remaining closed for two or three years longer, and frequently persistent on the branches for eighteen or twenty years; seeds almost triangular, full and rounded on the sides, nearly ¼′ long, with a thin conspicuously roughened light brown shell, their wings widest below the middle, gradually narrowed to the ends, 1′ long, ¼′ wide.

A tree, when crowded in the forest occasionally 60° high, with a trunk 2°—3° in diameter, and a few short branches near the summit forming a narrow round-topped head; in open ground usually 20°—30° tall, and often fertile when only a few feet high, with a short thick trunk frequently clothed to the ground, and long horizontal branches, the lower pendulous toward the extremities, the upper sweeping in graceful upward curves and forming a flat-topped often irregular head, and stout branchlets, light orange color when they first appear, [30] soon growing darker and ultimately dark brown. Bark on the lower part of the trunk ¾′—1′ thick and broken into irregularly shaped plates separating on the surface into thin loose dark brown scales tinged with red, higher on the stem, and on the branches dark brown and broken into thin loose scales. Wood light, soft, not strong, brittle, very coarse-grained, pale brown, with thick nearly white sapwood; somewhat used for fuel, and in Pennsylvania manufactured into charcoal.

Distribution. Dry gravelly slopes and ridges of the Appalachian Mountains from southern Pennsylvania to North Carolina, eastern Tennessee and northern Georgia, sometimes ascending to elevations of 3000°, with isolated outlying stations in eastern Pennsylvania, western New Jersey, Maryland, the District of Columbia and Virginia; often forming toward the southern limits of its range pure forests of considerable extent.

28. Pinus Torreyana Carr. Torrey Pine.

Leaves forming great tufts at the ends of the branches, stout, dark green, conspicuously marked on the 3 faces by numerous rows of stomata, 8′—13′ long. Flowers from January to March; male yellow, in short dense heads; female subterminal on long stout peduncles. Fruit broad-ovoid, spreading or reflexed on long stalks, 4′—6′ in length, becoming deep chestnut-brown, with thick scales armed with minute spines; mostly deciduous in their fourth year and in falling leaving a few of the barren scales on the stalk attached to the branch; seeds oval, more or less angled, ¾′—1′ long, dull brown and mottled on the lower side, light yellow-brown on the upper side, with a thick hard shell, nearly surrounded by their dark brown wings often nearly ½′ long.

A tree, usually 30°—40° high, with a short trunk about 1° in diameter, or occasionally 50°—60° tall, with a long straight slightly tapering stem 2½° in diameter, stout spreading and often ascending branches, and very stout branchlets bright green in their first season, becoming light purple and covered with a metallic bloom the following year, ultimately nearly black. Bark ¾′—1′ thick, deeply and irregularly divided into broad flat ridges covered by large thin closely appressed light red-brown scales. Wood light, soft, not strong, coarse-grained, light yellow, with thick yellow or nearly white sapwood; occasionally used for fuel. The large edible seeds are gathered in large quantities and are eaten raw or roasted.

Distribution. Only in a narrow belt a few miles long on the coast near the mouth of the Soledad River just north of San Diego and on the island of Santa Rosa, California; the least widely distributed Pine-tree of the United States.

Now planted in the parks of San Diego, California, and in New Zealand, growing rapidly in cultivation, and promising to attain a much larger size than on its native cliffs.

2. LARIX Adans. Larch.

Tall pyramidal trees, with thick sometimes furrowed scaly bark, heavy heartwood, thin pale sapwood, slender remote horizontal often pendulous branches, elongated leading branchlets, short thick spur-like lateral branchlets, and small subglobose buds, their inner scales accrescent and marking the lateral branchlets with prominent ring-like scars. Leaves awl-shaped, triangular and rounded above, or rarely 4-angled, spirally disposed and remote on leading shoots, on lateral branchlets in crowded fascicles, each leaf in the axil of a deciduous bud-scale, deciduous. Flowers solitary, terminal, the staminate globose, oval or oblong, sessile or stalked, on leafless branches, yellow, composed of numerous spirally arranged anthers with connectives produced above them into short points, the pistillate appearing with the leaves, short-oblong to oblong, composed of few or many green nearly orbicular stalked scales in the axes of much longer mucronate usually scarlet bracts. Fruit a woody ovoid-oblong conic or subglobose short-stalked cone composed of slightly thickened suborbicular or oblong-obovate concave scales, shorter or longer than their bracts, gradually decreasing from the centre to the ends of the cone, the small scales usually sterile. Seeds nearly triangular, rounded on the sides, shorter than their wings; the outer seed-coat crustaceous, light brown, the inner membranaceous, pale chestnut-brown and lustrous; cotyledons usually 6, much shorter than the inferior radicle.

Larix is widely distributed over the northern and mountainous region of the northern hemisphere from the Arctic Circle to the mountains of West Virginia and Oregon in the New World, and to central Europe, the Himalayas, Siberia, Korea, western China, and Japan in the Old World. Ten species are recognized. Of the exotic species the European Larix decidua, Mill., has been much planted for timber and ornament in the northeastern states, where the Japanese Larix Kœmpferi, Sarg., also flourishes.

CONSPECTUS OF THE NORTH AMERICAN SPECIES.

1. Larix laricina K. Koch. Tamarack. Larch.

Leaves linear, triangular, rounded above, prominently keeled on the lower surface, ¾′—1¼′ long, bright green, conspicuously stomatiferous when they first appear; turning yellow and falling in September or October. Flowers: male subglobose and sessile; female oblong, with light-colored bracts produced into elongated green tips, and nearly orbicular rose-red scales. Fruit on stout incurved stems, subglobose, rather obtuse, ½′—¾′ long, composed of about 20 scales slightly erose on their nearly entire margins, rather longer than broad and twice as long as their bracts, bright chestnut-brown at maturity; usually falling during their second year; seeds ⅛′ long, about one third as long as their light chestnut-brown wings broadest near the middle and obliquely rounded at apex.

A tree, 50°—60° high, with a trunk 18′—20′ in diameter, small horizontal branches forming during the early life of the tree a narrow regular pyramidal head always characteristic of this tree when crowded in the forest, or with abundant space sweeping out in graceful [32] curves, often becoming contorted and pendulous and forming a broad open frequently picturesque head, and slender leading branchlets often covered at first with a glaucous bloom, becoming light orange-brown during their first winter and conspicuous from the small globose dark red lustrous buds. Bark ½′—¾′ thick, separating into thin closely appressed rather bright reddish brown scales. Wood heavy, hard, very strong, rather coarse-grained, very durable, light brown; largely used for the upper knees of small vessels, fence-posts, telegraph-poles, and railway-ties.

Distribution. At the north often on well-drained uplands, southward in cold deep swamps which it often clothes with forests of closely crowded trees, from Labrador to the Arctic Circle, ranging west of the Rocky Mountains to latitude 65° 35′ north, and southward through Canada and the northern states to northern and eastern Pennsylvania, Garrett County, Maryland (Oakland to Thayerville), and Preston County, West Virginia (Cranesville Swamp), northern Indiana and Illinois, and northeastern Minnesota; along the eastern foothills of the Rocky Mountains to about latitude 53° and between the Yukon River and Cook Inlet, Alaska (Larix alaskensis Wight.); very abundant in the interior of Labrador, where it is the largest tree; common along the margins of the barren lands stretching beyond the sub-Arctic forest to the shores of the Arctic Sea; attaining its largest size north of Lake Winnipeg on low benches which it occasionally covers with open forests; on the eastern slopes of the northern Rocky Mountains usually at elevation from 600°—1700° above the sea; rare and local toward the southern limits of its range.

Occasionally planted as an ornamental tree in the northeastern states, growing rapidly and attaining in cultivation a large size and picturesque habit.

2. Larix occidentalis Nutt. Tamarack.

Leaves triangular, rounded on the back, conspicuously keeled below, rigid, sharp-pointed, 1′—1¾′ long, about ¹/₃₂′ wide, light pale green, turning pale yellow early in the autumn. Flowers: male short-oblong; female oblong, nearly sessile, with orbicular scales and bracts produced into elongated tips. Fruit oblong, short-stalked, 1′—1½′ long, with numerous thin stiff scales nearly entire and sometimes a little reflexed on their margins, much shorter than their bracts, more or less thickly coated on the lower surface below the middle with hoary tomentum, and standing after the escape of the seeds at right angles to the axis of the cone, or often becoming reflexed; seeds nearly ¼′ long, with a pale brown shell, one half to two thirds as long as the thin fragile pale wings broadest near the middle and obliquely rounded at apex.

A tree, sometimes 180° high, with a tall tapering naked trunk 6°—8° in diameter, or on dry soil and exposed mountain slopes usually not more than 100° tall, with a short narrow pyramidal head of small branches clothed with scanty foliage, or occasionally with a larger crown of elongated drooping branches, stout branchlets covered when they first appear with soft pale pubescence, usually soon glabrous, bright orange-brown in their first year, ultimately becoming dark gray-brown, and dark chestnut-brown winter-buds about ⅛′ in diameter. Bark of young stems thin, dark-colored and scaly, becoming near the base of old trunks 5′ or 6′ thick and broken into irregularly shaped oblong plates often 2° long and covered with thin closely appressed light cinnamon-red scales. Wood very heavy, exceedingly hard and strong, close-grained, very durable in contact with the soil, bright light red, with thin nearly white sapwood; largely used for railway-ties and fence-posts, and manufactured into lumber used in cabinet-making and the interior finish of buildings.

Distribution. Moist bottom-lands and on high benches and dry mountain sides generally at elevations between 2000° and 7000° above sea-level, usually singly or in small groves, through the basin of the upper Columbia River from southern British Columbia to the western slopes of the continental divide of northern Montana, and to the eastern slopes of the Cascade Mountains of Washington and northern Oregon; most abundant and of its largest size on the bottom-lands of streams flowing into Flat Head Lake in northern Montana, and in northern Idaho.

Occasionally planted in the eastern states and in Europe, but in cultivation showing little promise of attaining a large size or becoming a valuable ornamental or timber-tree.

3. Larix Lyallii Parl. Tamarack.

Leaves 4-angled, rigid, short-pointed, pale blue-green, 1′—1½′ long. Flowers: male short-oblong; female ovoid-oblong, with dark red or occasionally pale yellow-green scales and dark purple bracts abruptly contracted into elongated slender tips. Fruit ovoid, rather acute, 1½′—2′ long, subsessile or raised on a slender stalk coated with hoary tomentum, with dark reddish purple or rarely green erose scales, fringed and covered on their lower surface with matted hairs at maturity spreading nearly at right angles and finally much reflexed, much shorter than their dark purple very conspicuous long-tipped bracts; seeds full and rounded on the sides, ⅛′ long and about half as long as their light red lustrous wings broadest near the base with nearly parallel sides.

A tree, usually 25°—50° high, with a trunk generally 18′—20′ but rarely 3°—4° in diameter, and remote elongated exceedingly tough persistent branches sometimes pendulous, developing very irregularly and often abruptly ascending at the extremities, stout branchlets [34] coated with hoary tomentum usually persistent until after their second winter, ultimately becoming nearly black, and prominent winter-buds with conspicuous long white matted hairs fringing the margins of their scales and often almost entirely covering the bud. Bark of young trees and of the branches thin, rather lustrous, smooth, and pale gray tinged with yellow, becoming loose and scaly on larger stems and on the large branches of old trees, and on fully grown trunks ½′—¾′ thick and slightly divided by shallow fissures into irregularly shaped plates covered by thin dark-red brown loosely attached scales. Wood heavy, hard, coarse-grained, light reddish brown.

Distribution. Near the timber-line on mountain slopes at elevations of 4000°—8000°, from southern Alberta on the eastern slope of the Rocky Mountains and from the interior of southern British Columbia, southward along the eastern slopes of the Cascade Mountains of northern Washington to Mt. Stewart at the head of the north fork of the Yakima River, and along the continental divide to the middle fork of Sun River, Montana, forming here a forest of considerable size at elevations of 7000°—8000°, and on the Bitter Root Mountains to the headwaters of the south fork of the Clearwater River, Idaho.

3. PICEA Dietr. Spruce.

Pyramidal trees, with tall tapering trunks often stoutly buttressed at the base, thin scaly bark, soft pale wood containing numerous resin-canals, slender whorled twice or thrice ramified branches, their ultimate divisions stout, glabrous or pubescent, and leaf-buds usually in 3’s, the 2 lateral in the axils of upper leaves. Leaves linear, spirally disposed, extending out from the branch on all sides or occasionally appearing 2-ranked by the twisting of those on its lower side, mostly pointing to the end of the branch, entire, articulate on prominent persistent rhomboid ultimately woody bases, keeled above and below, 4-sided and stomatiferous on the 4 sides, or flattened and stomatiferous on the upper and occasionally on the lower side, persistent from seven to ten years, deciduous in drying. Flowers terminal or in the axils of upper leaves, the male usually long-stalked, composed of numerous spirally arranged anthers with connectives produced into broad nearly circular toothed crests, the female oblong, oval or cylindric, with rounded or pointed scales, each in the axis of an accrescent bract shorter than the scale at maturity. Fruit an ovoid or oblong, cylindric pendant cone, crowded on the upper branches or in some species scattered over the upper half of the tree. Seeds ovoid or oblong, usually acute at base, much shorter than their wings; outer seed-coat crustaceous, light or dark brown, the inner membranaceous, pale chestnut-brown; cotyledons 4—15.

Picea is widely distributed through the colder and temperate regions of the northern hemisphere, some species forming great forests on plains and high mountain slopes. Thirty-seven species are now recognized, ranging from the Arctic Circle to the slopes of the southern Appalachian Mountains and to those of northern New Mexico and Arizona in the New World, and to central and southeastern Europe, the Caucasus, the Himalayas, western China, Formosa and Japan. Of exotic species the so-called Norway Spruce, Picea Abies Karst., one of the most valuable timber-trees of Europe, has been largely planted for ornament and shelter in the eastern states, where the Caucasian Picea orientalis Carr., and some of the Japanese species also flourish.

CONSPECTUS OF THE NORTH AMERICAN SPECIES.

1. Picea mariana B. S. P. Black Spruce.

Leaves slightly incurved above the middle, abruptly contracted at apex into short callous tips, pale blue-green and glaucous at maturity, ¼′—¾′ long, hoary on the upper surface from the broad bands of stomata, and lustrous and slightly stomatiferous on the lower surface. Flowers: male subglobose, with dark red anthers; female oblong-cylindric, with obovate purple scales rounded above, and oblong purple glaucous bracts rounded and denticulate at apex. Fruit ovoid, pointed, gradually narrowed at the base into short strongly incurved stalks, ½′—1½′ long, with rigid puberulous scales rounded or rarely somewhat pointed at apex and more or less erose on the notched pale margins, turning as they ripen dull gray-brown and becoming as the scales gradually open and slowly discharge their seeds almost globose; sometimes remaining on the branches for twenty or thirty years, the oldest close to the base of the branches near the trunk; seeds oblong, narrowed to the acute base, about ⅛′ long, very dark brown, with delicate pale brown wings broadest above the middle, very oblique at the apex, about ½′ long, ⅛′ wide.

A tree, usually 20°—30° and occasionally 100° high, with a trunk 6′—12′ and rarely 3° in diameter, and comparatively short branches generally pendulous with upward curves, forming an open irregular crown, light green branchlets coated with pale pubescence, soon beginning to grow darker, and during their first winter light cinnamon-brown and covered with short rusty pubescence, their thin brown bark gradually becoming glabrous and beginning to break into small thin scales during their second year; at the extreme north [36] sometimes cone-bearing when only 2°—3° high. Winter-buds ovoid, acute, light reddish brown, puberulous, about ⅛′ long. Bark ¼′—½′ thick and broken on the surface into thin rather closely appressed gray-brown scales. Wood light, soft, not strong, pale yellow-white, with thin sapwood; probably rarely used outside of Manitoba and Saskatchewan, except in the manufacture of paper pulp. Spruce-gum, the resinous exudations of the Spruce-trees of northeastern America, is gathered in considerable quantities principally in northern New England and Canada, and is used as a masticatory. Spruce-beer is made by boiling the branches of the Black and Red Spruces.

Distribution. At the north on well-drained bottom-lands and the slopes of barren stony hills, and southward in sphagnum-covered bogs, swamps, and on their borders, from Labrador to the valley of the Mackenzie River in about latitude 65° north, and, crossing the Rocky Mountains, through the interior of Alaska to the valley of White River; southward through Newfoundland, the maritime provinces, eastern Canada and the northeastern United States to central Pennsylvania, and along the Alleghany Mountains to northern Virginia; and from the eastern foothills of the Rocky Mountains in Alberta, through northern Saskatchewan and northern Manitoba, and south to northeastern and northern Minnesota, and central Wisconsin and Michigan; very abundant at the far north and the largest coniferous tree of Saskatchewan and northern Manitoba, covering here large areas and growing to its largest size; common in Newfoundland and all the provinces of eastern Canada except southern Ontario; in the United States less abundant, of small size, and usually only in cold sphagnum swamps (var. brevifolia Rehd.)

Occasionally planted as an ornamental tree, the Black Spruce is short-lived in cultivation and one of the least desirable of all Spruce-trees for the decoration of parks and gardens.

2. Picea rubra Link. Red Spruce.

Leaves more or less incurved above the middle, acute or rounded and furnished at the apex with short callous points, dark green often slightly tinged with yellow, very lustrous, marked on the upper surface by 4 rows and on the lower less conspicuously by 2 rows of stomata on each side of the prominent midrib, ½′—⅝′ long, nearly ¹/₁₆′ wide. Flowers: male oval, almost sessile, bright red; female oblong-cylindric, with thin rounded scales reflexed and slightly erose on their margins, and obovate bracts rounded and laciniate above. Fruit on very short straight or incurved stalks, ovoid-oblong, gradually narrowed from near the middle to the acute apex, 1¼′—2′ long, with rigid puberulous scales entire or slightly toothed at the apex; bright green or green somewhat tinged with purple when [37] fully grown, becoming light reddish brown and lustrous at maturity, beginning to fall as soon as the scales open in the autumn or early winter, and generally disappearing from the branches the following summer; seeds dark brown, about ⅛′ long, with short broad wings full and rounded above the middle.

A tree, usually 70°—80° and occasionally 100° high, with a trunk 2°—3° in diameter, branches long-persistent on the stem and clothing it to the ground, forming a narrow rather conical head, or soon disappearing below from trees crowded in the forest, stout pubescent light green branchlets, becoming bright reddish brown or orange-brown during their first winter, glabrous the following year, and covered in their third or fourth year with scaly bark. Winter-buds ovoid, acute, ¼′—⅓′ long, with light reddish brown scales. Bark ¼′—½′ thick, and broken into thin closely appressed irregularly shaped red-brown scales. Wood light, soft, close-grained, not strong, pale slightly tinged with red, with paler sapwood usually about 2′ thick; largely manufactured into lumber in the northeastern states, Pennsylvania, and Virginia, and used for the flooring and construction of houses, for the sounding-boards of musical instruments, and in the manufacture of paper-pulp.

Distribution. Well-drained uplands and mountain slopes, often forming a large part of extensive forests, from Prince Edward Island and the valley of the St. Lawrence southward to the coast of Massachusetts, along the interior hilly part of New England, New York, and northern Pennsylvania and on the slopes of the Alleghany Mountains at elevations above 2500 feet from West Virginia to North Carolina and Tennessee.

Occasionally planted in the eastern states and in Europe as an ornamental tree, but growing in cultivation more slowly than any other Spruce-tree.

3. Picea glauca Voss. White Spruce.

Leaves crowded on the upper side of the branches by the twisting of those on the lower side, incurved, acute or acuminate with rigid callous tips, pale blue and hoary when they first appear, becoming dark blue-green or pale blue, marked on each of the 4 sides by 3 or 4 rows of stomata, ⅓′—¾′ long. Flowers: male pale red, soon appearing yellow from the thick covering of pollen; female oblong-cylindric, with round nearly entire pale red or yellow-green scales, broader than long, and nearly orbicular denticulate bracts. Fruit nearly sessile or borne on short thin straight stems, oblong-cylindric, slender, slightly narrowed to the ends, rather obtuse at apex, usually about 2′ long, pale green sometimes tinged with red when fully grown, becoming at maturity pale brown and lustrous, with nearly orbicular scales, rounded, truncate, and slightly emarginate, or rarely narrowed at apex, and very thin, flexible and elastic at maturity, usually deciduous in the autumn or during the following winter; seeds about ⅛′ long, pale brown, with narrow wings gradually widened from the base to above the middle and very oblique at the apex.

A tree, with disagreeable smelling foliage, rarely more than 60°—70° tall, with a trunk [38] not more than 2° in diameter, long comparatively thick branches densely clothed with stout rigid laterals sweeping out in graceful upward curves, and forming a broad-based rather open pyramid often obtuse at the apex, stout glabrous branchlets orange-brown during their first autumn and winter, gradually growing darker grayish brown. Winter-buds broadly ovoid, obtuse, covered by light chestnut-brown scales with thin often reflexed ciliate margins. Bark ¼′—½′ thick, separating irregularly into thin plate-like light gray scales more or less tinged with brown. Wood light, soft, not strong, straight-grained light yellow, with hardly distinguishable sapwood; manufactured into lumber in the eastern provinces of Canada and in Alaska, and used in construction, for the interior finish of buildings, and for paper-pulp.

Distribution. Banks and borders of streams and lakes, ocean cliffs, and in the north the rocky slopes of low hills, from Labrador along the northern frontier of the forest nearly to the shores of the Arctic Sea, reaching Behring Strait in 66° 44′ north latitude, and southward down the Atlantic coast to southern Maine, northern New Hampshire, Vermont, and New York, shores of Saginaw Bay, Michigan, northern Wisconsin and Minnesota, and through the interior of Alaska.

The variety (var. albertiana Sarg.) of the Gaspé Peninsula and the valleys of the Black Hills of South Dakota and of the Rocky Mountains of northern Wyoming, Montana, Alberta and northward, is a tree with a narrow pyramidal head, sometimes 150° high, with a trunk 3° to 4° in diameter, and shorter and rather broader cones than those of the typical White Spruce of the east, although not shorter or as short as the cones of that tree in the extreme north.

Often planted in Canada, northern New England, and northern Europe as an ornamental tree; in southern New England and southward suffering from heat and dryness.

4. Picea Engelmannii Engelm. White Spruce. Engelmann Spruce.

Leaves soft and flexible, with acute callous tips, slender, nearly straight or slightly incurved on vigorous sterile branches, stouter, shorter, and more incurved on fertile branches, 1′—1⅛′ long, marked on each face by 3—5 rows of stomata, covered at first with a glaucous bloom, soon becoming dark blue-green or pale steel-blue. Flowers: male dark purple; female bright scarlet, with pointed or rounded and more or less divided scales, and oblong bracts rounded or acute or acuminate and denticulate at apex or obovate-oblong and abruptly acuminate. Fruit oblong-cylindric to ellipsoidal, gradually narrowed to the ends, usually about 2′ long, sessile or very short-stalked, produced in great numbers on the upper branches, horizontal and ultimately pendulous, light green somewhat tinged with scarlet when fully grown, becoming light chestnut-brown and lustrous, with thin flexible slightly concave scales, generally erose-dentate or rarely almost entire on the margins, usually broadest at the middle, wedge-shaped below, and gradually contracted above into a truncate or acute apex, or occasionally obovate and rounded above; mostly deciduous in the autumn or early in their first winter soon after the escape of the seeds; seeds obtuse [39] at the base, nearly black, about ⅛′ long and much shorter than their broad very oblique wings.

A tree, with disagreeable smelling foliage sometimes 120° high, with a trunk 3° in diameter, spreading branches produced in regular whorls and forming a narrow compact pyramidal head, gracefully hanging short lateral branches, and comparatively slender branchlets pubescent for three or four years, light or dark orange-brown or gray tinged with brown during their first winter, their bark beginning to separate into small flaky scales in their fourth or fifth year; at its highest altitudes low and stunted with elongated branches pressed close to the ground. Winter-buds conic or slightly obtuse, with pale chestnut-brown scales scarious and often free and slightly reflexed on the margins. Bark ¼′—½′ thick, light cinnamon-red, and broken into large thin loose scales. Wood light, soft, not strong, close-grained, pale yellow tinged with red, with thick hardly distinguishable sapwood; largely manufactured into lumber used in the construction of buildings; also employed for fuel and charcoal. The bark is sometimes employed in tanning leather.

Distribution. High mountain slopes, often forming great forests from the mountains of Alberta, British Columbia and Alaska, southward over the interior mountain systems of the continent to southern New Mexico (the Sacramento Mountains) and northern Arizona, from elevations of 5000° at the north up to 11,500° and occasionally to 12,000° at the south, and westward through Montana and Idaho to the eastern slopes of the Cascade Mountains of Washington and Oregon; attaining its greatest size and beauty north of the northern boundary of the United States.

Occasionally planted as an ornamental tree in the New England states and northern Europe, where it grows vigorously and promises to attain a large size; usually injured in western Europe by spring frosts.

5. Picea pungens Englm. Blue Spruce. Colorado Spruce.

Leaves strongly incurved, especially those on the upper side of the branches, stout, rigid, acuminate and tipped with long callous sharp points, 1′—1⅛′ long on sterile branches, often not more than half as long on the fertile branches of old trees, marked on each side by 4—7 rows of stomata, dull bluish green on some individuals and light or dark steel-blue or silvery white on others, the blue colors gradually changing to dull blue-green at the end of three or four years. Flowers: male yellow tinged with red; female with broad oblong or slightly obovate pale green scales truncate or slightly emarginate at the denticulate apex, and acute bracts. Fruit produced on the upper third of the tree, sessile or short-stalked, oblong-cylindric, [40] slightly narrowed at the ends, usually about 3′ long, green more or less tinged with red when fully grown at midsummer, becoming pale chestnut-brown and lustrous, with flat tough rhombic scales flexuose on the margins, and acute, rounded or truncate at the elongated erose apex; seeds ⅛′ long or about half the length of their wings, gradually widening to above the middle and full and rounded at apex.

A tree, usually 80°—100° or occasionally 150° high, with a trunk rarely 3° in diameter and occasionally divided into 3 or 4 stout secondary stems, rigid horizontal branches disposed on young trees in remote whorls and decreasing regularly in length from below upward, the short stout stiff branchlets pointing forward and making flat-topped masses of foliage; branches on old trees short and remote, with stout lateral branches forming a thin ragged pyramidal crown; branchlets stout, rigid, glabrous, pale glaucous green, becoming bright orange-brown during the first winter and ultimately light grayish brown. Winter-buds stout, obtuse or rarely acute, ¼′—½′ long, with thin pale chestnut-brown scales usually reflexed on the margins. Bark of young trees gray or gray tinged with cinnamon-red and broken into small oblong plate-like scales, becoming on the lower part of old trunks ¾′—1½′ thick and deeply divided into broad rounded ridges covered with small closely appressed pale gray or occasionally bright cinnamon-red scales. Wood light, soft, close-grained, weak, pale brown or often nearly white, with hardly distinguishable sapwood.

Distribution. Banks of streams or on the first benches above them singly or in small groves at elevations between 6500° and 11,000° above the sea; Colorado and eastern Utah northward to the northern end of the Medicine Bow Mountains and on the Laramie Range in southern and on the Shoshone and Teton Mountains in northwestern Wyoming, and southward into northern New Mexico (Sierra Bianca, alt. 8000°—11,000°, Sacramento Mountains, Pecos River National Forest).

Often planted as an ornamental tree in the eastern and northern states and in western and northern Europe, especially individuals with blue foliage; very beautiful in early life but in cultivation soon becoming unsightly from the loss of the lower branches.

6. Picea Breweriana S. Wats. Weeping Spruce.

Leaves abruptly narrowed and obtuse at apex, straight or slightly incurved, rounded and obscurely ridged and dark green and lustrous on the lower surface, flattened and conspicuously marked on the upper surface by 4 or 5 rows of stomata on each side of the prominent midrib, ¾′—1⅛′ long, ¹/₁₆′—¹/₁₀′ wide. Flowers: male dark purple; female oblong-cylindric, with obovate scales rounded above and reflexed on the entire margins, and oblong bracts laciniately divided at their rounded or acute apex. Fruit ellipsoidal, gradually narrowed from the middle to the ends, acute at apex, rather oblique at base, suspended on straight slender stalks, deep rich purple or green more or less tinged with purple when fully grown, becoming light orange-brown, 2′—4′ long, with thin broadly ovate flat scales longer than broad, rounded at apex, opening late in the autumn after the escape of the [41] seeds, often becoming strongly reflexed and very flexible; usually remaining on the branches until their second winter; seeds acute at base, full and rounded on the sides, ⅛′ long, dark brown, and about one quarter the length of their wings broadest toward the full and rounded apex.

A tree, usually 80°—100° high, with a trunk 2°—3° in diameter above the swelling of its enlarged and gradually tapering base, and furnished to the ground with crowded branches, those at the top of the tree short and slightly ascending, with comparatively short pendulous lateral branches, those lower on the tree horizontal or pendulous and clothed with slender flexible whip-like laterals often 7°—8° long and not more than ¼′ thick and furnished with numerous long thin lateral branchlets, their ultimate divisions slender, coated with fine pubescence persistent until their third season, bright red-brown during their first winter, gradually growing dark gray-brown. Winter-buds conic, light chestnut-brown, ¼′ long and ⅛′ thick. Bark ½′—¾′ thick, broken into long thin closely appressed scales dull red-brown on the surface. Wood heavy, soft, close-grained, light brown or nearly white, with thick hardly distinguishable sapwood.

Distribution. Dry mountain ridges and peaks near the timber-line on both slopes of the Siskiyou Mountains on the boundary between California and Oregon, forming small groves at elevations of about 7000° above the sea; on a high peak west of Marble Mountain in Siskiyou County, California; on the coast ranges of southwestern Oregon at elevations of 4000°—5000°.

7. Picea sitchensis Carr. Tideland Spruce. Sitka Spruce.

Leaves standing out from all sides of the branches and often nearly at right angles to them, frequently bringing their white upper surface to view by a twist at their base, straight or slightly incurved, acute or acuminate with long callous tips, slightly rounded, green, lustrous, and occasionally marked on the lower surface with 2 or 3 rows of small conspicuous stomata on each side of the prominent midrib, flattened, obscurely ridged and almost covered with broad silvery white bands of numerous rows of stomata on the upper surface, ½′—1⅛′ long and ¹/₁₆′—¹/₁₂′ wide, mostly persistent 9—11 years. Flowers: male at the ends of the pendant lateral branchlets, dark red; female on rigid terminal shoots of the branches of the upper half of the tree, with nearly orbicular denticulate scales, often slightly truncate above and completely hidden by their elongated acuminate bracts. Fruit oblong-cylindric, short-stalked, yellow-green often tinged with dark red when fully grown, becoming lustrous and pale yellow or reddish brown, 2½′—4′ long, with thin stiff elliptic scales rounded toward the apex, denticulate above the middle, and nearly twice as long as their lanceolate denticulate [42] bracts; deciduous mostly during their first autumn and winter; seeds full and rounded, acute at the base, pale reddish brown, about ⅛′ long, with narrow oblong slightly oblique wings ⅓′—½′ in length.

A tree, usually about 100° high, with a conspicuously tapering trunk often 3°—4° in diameter above its strongly buttressed and much-enlarged base, occasionally 200° tall, with a trunk 15°—16° in diameter, horizontal branches forming an open loose pyramid and on older trees clothed with slender pendant lateral branches frequently 2°—3° long, and stout rigid glabrous branchlets pale green at first, becoming dark or light orange-brown during their first autumn and winter and finally dark gray-brown; at the extreme northwestern limits of its range occasionally reduced to a low shrub. Winter-buds ovoid, acute or conical, ¼′—½′ long, with pale chestnut-brown acute scales, often tipped with short points and more or less reflexed above the middle. Bark ¼′—½′ thick and broken on the surface into large thin loosely attached dark red-brown or on young trees sometimes bright cinnamon-red scales. Wood light, soft, not strong, straight-grained, light brown tinged with red, with thick nearly white sapwood; largely manufactured into lumber used in the interior finish of buildings, for fencing, boat-building, aeroplanes, cooperage, wooden-ware, and packing-cases.

Distribution. Moist sandy, often swampy soil, or less frequently at the far north on wet rocky slopes, from the eastern end of Kadiak Island, southward through the coast region of Alaska, British Columbia, Washington, and Oregon to Mendocino County, California; in Washington, occasionally ranging inland to the upper valley of the Nesqually River.

Often planted in western and central Europe and occasionally in the middle Atlantic states as an ornamental tree.

4. TSUGA Carr. Hemlock.

Tall pyramidal trees, with deeply furrowed astringent bark bright cinnamon-red except on the surface, soft pale wood, nodding leading shoots, slender scattered horizontal often pendulous branches, the secondary branches three or four times irregularly pinnately ramified, with slender round glabrous or pubescent ultimate divisions, the whole forming graceful pendant masses of foliage, and minute winter-buds. Leaves flat or angular, obtuse and often emarginate or acute at apex, spirally disposed, usually appearing almost 2-ranked by the twisting of their petioles, those on the upper side of the branch then much shorter than the others, abruptly narrowed into short petioles jointed on ultimately woody persistent bases, with stomata on the lower surface; on one species not 2-ranked, and of nearly equal length, with stomata on both surfaces. Flowers solitary, the male in the axils of leaves of the previous year, globose, composed of numerous subglobose anthers, with connectives produced into short gland-like tips, the female terminal, erect, with nearly circular scales slightly longer or shorter than their membranaceous bracts. Fruit [43] an ovoid-oblong, oval, or oblong-cylindric obtuse usually pendulous nearly sessile green or rarely purple cone becoming light or dark reddish brown, with concave suborbicular or ovate-oblong scales thin and entire on the margins, much longer than their minute bracts, persistent on the axis of the cone after the escape of the seeds. Seeds furnished with resin-vesicles, ovoid-oblong, compressed, nearly surrounded by their much longer obovate-oblong wings; outer seed-coat crustaceous, light brown, the inner membranaceous, pale chestnut-brown, and lustrous; cotyledons 3—6, much shorter than the inferior radicle.

Tsuga is confined to temperate North America, Japan, central and southwestern China, Formosa, and the Himalayas; nine species have been distinguished.

CONSPECTUS OF THE NORTH AMERICAN SPECIES.

1. Tsuga canadensis Carr. Hemlock.

Leaves, rounded and rarely emarginate at apex, dark yellow-green, lustrous and obscurely grooved especially toward the base on the upper surface, marked on the lower surface by 5 or 6 rows of stomata on each side of the low broad midrib, ⅓′—⅔′ long, about ¹/₁₆′ wide, deciduous in their third season from dark orange-colored persistent bases. Flowers: male light yellow; female pale green, with broad bracts coarsely laciniate on the margins and shorter than their scales. Fruit on slender puberulous stalks often ¼′ long, ovoid, acute, ½′—¾′ long, with broad-obovate scales almost as wide as long, and broad truncate bracts slightly laciniate on the margins, opening and gradually losing their seeds during [44] the winter and mostly persistent on the branches until the following spring; seeds ¹/₁₆′ long, usually with 2 or 3 large oil-vesicles, nearly half as long as their wings broad at the base and gradually tapering to the rounded apex.

A tree, usually 60°—70°, and occasionally 100° high, with a trunk 2°—4° in diameter, gradually and conspicuously tapering toward the apex, long slender horizontal or pendulous branches, persistent until overshadowed by other trees, and forming a broad-based rather obtuse pyramid, and slender light yellow-brown pubescent branchlets, growing darker during their first winter and glabrous and dark red-brown tinged with purple in their third season. Winter-buds obtuse, light chestnut-brown, slightly puberulous, about ¹/₁₆′ long. Bark ½′—¾′ thick, deeply divided into narrow rounded ridges covered with thick closely appressed scales varying from cinnamon-red to gray more or less tinged with purple. Wood light, soft, not strong, brittle, coarse-grained, difficult to work, liable to wind-shake and splinter, not durable when exposed to the air, light brown tinged with red, with thin somewhat darker sapwood; largely manufactured into coarse lumber employed for the outside finish of buildings. The astringent inner bark affords the largest part of the material used in the northeastern states and Canada in tanning leather. From the young branches oil of hemlock is distilled.

Distribution. Scattered through upland forests and often covering the northern slopes of rocky ridges and the steep rocky banks of narrow river-gorges from Nova Scotia to eastern Minnesota (Carleton County), and southward through the northern states to Newcastle County, Delaware, cliffs of Tuckahoe Creek, Queen Anne’s County, Maryland, southern Michigan, southern Indiana (bank of Back Creek near Leesville, Laurence County), southwestern Wisconsin, and along the Appalachian Mountains to northern Georgia, and in northern Alabama; most abundant and frequently an important element of the forest in New England, northern New York, and western Pennsylvania; attaining its largest size near streams on the slopes of the high mountains of North Carolina and Tennessee.

Largely cultivated with numerous seminal varieties as an ornamental tree in the northern states, and in western and central Europe.

2. Tsuga caroliniana Engelm. Hemlock.

Leaves retuse or often emarginate at apex, dark green, lustrous and conspicuously grooved on the upper surface, marked on the lower surface by a band of 7 or 8 rows of stomata on each side of the midrib, ⅓′—¾′ long, about ¹/₁₂′ wide, deciduous from the orange-red bases during their fifth year. Flowers: male tinged with purple; female purple, with broadly ovate bracts, scarious and erose on the margins and about as long as their scales. Fruit on short stout stalks, oblong, 1′—1½′ long, with narrow-oval scales gradually narrowed and rounded at apex, rather abruptly contracted at base into distinct stipes, thin, concave, puberulous on the outer surface, twice as long as their broad pale bracts, spreading nearly at right angles to the axis of the cone [45] at maturity, their bracts rather longer than wide, wedge-shaped, pale, nearly truncate or slightly pointed at the broad apex; seeds ⅙′ long, with numerous small oil-vesicles on the lower side, and one quarter as long as the pale lustrous wings broad or narrow at the base and narrowed to the rounded apex.

A tree, usually 40°—50°, or occasionally 70° high, with a trunk rarely exceeding 2° in diameter, short stout often pendulous branches forming a handsome compact pyramidal head, and slender light orange-brown pubescent branchlets, usually becoming glabrous and dull brown more or less tinged with orange during their third year. Winter-buds obtuse, dark chestnut-brown, pubescent, nearly ⅛′ long. Bark of the trunk ¾′—1¼′ thick, red-brown, and deeply divided into broad flat connected ridges covered with thin closely appressed plate-like scales. Wood light, soft, not strong, brittle, coarse-grained, pale brown tinged with red, with thin nearly white sapwood.

Distribution. Rocky banks of streams usually at elevations between 2500° and 3000° on the Blue Ridge from southwestern Virginia to northern Georgia, generally singly or in small scattered groves of a few individuals.

Occasionally planted as an ornamental tree in the northern states, and in western Europe.

3. Tsuga heterophylla Sarg. Hemlock.

Leaves rounded at apex, conspicuously grooved, dark green and very lustrous on the upper surface, marked below by broad white bands of 7—9 rows of stomata, abruptly contracted at the base into slender petioles, ¼′—¾′ long and ¹/₁₆′—¹/₁₂′ wide, mostly persistent 4—7 years. Flowers: male yellow; female purple and puberulous, with broad bracts gradually narrowed to an obtuse point and shorter than their broadly ovate slightly scarious scales. Fruit oblong-ovoid, acute, sessile, ¾′—1′ long, with slightly puberulous oval scales, often abruptly narrowed near the middle, and dark purple puberulous bracts rounded and abruptly contracted at apex; seeds ⅛′ long, furnished with occasional oil-vesicles, one third to one half as long as their narrow wings.

A tree, frequently 200° high, with a tall trunk 6°—10° in diameter, and short slender usually pendulous branches forming a narrow pyramidal head, and slender pale yellow-brown branchlets ultimately becoming dark reddish brown, coated at first with long pale hairs, and pubescent or puberulous for five or six years. Winter-buds ovoid, bright chestnut-brown, about ¹/₁₆′ long. Bark on young trunks thin, dark orange-brown, and [46] separated by shallow fissures into narrow flat plates broken into delicate scales, becoming on fully grown trees 1′—1½′ thick and deeply divided into broad flat connected ridges covered with closely appressed brown scales more or less tinged with cinnamon-red. Wood light, hard and tough, pale brown tinged with yellow, with thin nearly white sapwood; stronger and more durable than the wood of the other American hemlocks; now largely manufactured into lumber used principally in the construction of buildings. The bark is used in large quantities in tanning leather; from the inner bark the Indians of Alaska obtain one of their principal articles of vegetable food.

Distribution. Southeastern Alaska, southward near the coast to southern Mendocino County, California, extending eastward over the mountains of southern British Columbia, northern Washington, Idaho and Montana, to the western slopes of the continental divide, and through Oregon to the western slopes of the Cascade Mountains, sometimes ascending in the interior to elevations of 6000° above the sea; most abundant and of its largest size on the coast of Washington and Oregon; often forming a large part of the forests of the northwest coast.

4. Tsuga Mertensiana Sarg. Mountain Hemlock. Black Hemlock.

Leaves standing out from all sides of the branch, remote on leading shoots and crowded on short lateral branchlets, rounded and occasionally obscurely grooved or on young plants sometimes conspicuously grooved on the upper surface, rounded and slightly ribbed on the lower surface, bluntly pointed, often more or less curved, stomatiferous above and below, with about 8 rows of stomata on each surface, light bluish green or on some individuals pale blue, ½′—1′ long, about ¹/₁₆′ wide, abruptly narrowed into nearly straight or slightly twisted petioles articulate on bases as long or rather longer than the petioles; irregularly deciduous during their third and fourth years. Flowers: male borne on slender pubescent drooping stems, violet-purple; female erect, with delicate lustrous dark purple or yellow-green bracts gradually narrowed above into slender often slightly reflexed tips and much longer than their scales. Fruit sessile, oblong-cylindric, narrowed toward the blunt apex and somewhat toward the base, erect until more than half grown, pendulous or rarely erect at maturity, ⅝′—3′ long, with thin delicate oblong-obovate scales gradually contracted from above the middle to the wedge-shaped base, rounded at the slightly thickened more or less erose margins, puberulous on the outer surface, usually bright bluish purple or occasionally pale yellow-green, four or five times as long as their short-pointed dark purple or brown bracts; seeds light brown, ⅛′ long, often marked on the [47] surface next their scales with 1 or 2 large resin-vesicles, with wings nearly ½′ long, broadest above the middle, gradually narrowed below, slightly or not at all oblique at the rounded apex.

A tree, usually 70°—100° but occasionally 150° high, with a slightly tapering trunk 4°—5° in diameter, gracefully pendant slender branches furnished with drooping frond-like lateral branches, their ultimate divisions erect and forming an open pyramid surmounted by the long drooping leading shoot, and thin flexible or sometimes stout rigid branchlets light reddish brown and covered for two or three years with short pale dense pubescence, becoming grayish brown and very scaly. Winter-buds acute, about ⅛′ long, the scales of the outer ranks furnished on the back with conspicuous midribs produced into slender deciduous awl-like tips. Bark 1′—1½′ thick, deeply divided into connected rounded ridges broken into thin closely appressed dark cinnamon scales shaded with blue or purple. Wood light, soft, not strong, close-grained, pale brown or red, with thin nearly white sapwood; occasionally manufactured into lumber.

Distribution. Exposed ridges and slopes at high altitudes along the upper border of the forest from southeastern Alaska, southward over the mountain ranges of British Columbia to the Olympic Mountains of Washington, and eastward to the western slopes of the Selkirk Mountains in the interior of southern British Columbia, and along the Bitter Root Mountains to the headwaters of the Clearwater River, Idaho; along the Cascade Mountains of Washington and Oregon, on the mountain ranges of northern California, and along the high Sierra Nevada to the cañon of the south fork of King’s River, California; in Alaska occasionally descending to the sea-level, and toward the southern limits of its range often ascending to elevations of 10,000°.

Often planted as an ornamental tree in western and central Europe, and rarely in the eastern United States.

5. PSEUDOTSUGA Carr.

Pyramidal trees, with thick deeply furrowed bark, hard strong wood, with spirally marked wood-cells, slender usually horizontal irregularly whorled branches clothed with slender spreading lateral branches forming broad flat-topped masses of foliage, ovoid acute leaf-buds, the lateral buds in the axils of upper leaves, their inner scales accrescent and marking the branchlets with ring-like scars. Leaves petiolate, linear, flat, rounded and obtuse or acuminate at apex, straight or incurved, grooved on the upper side, marked on the lower side by numerous rows of stomata on each side of the prominent midrib, spreading nearly at right angles with the branch. Flowers solitary, the male axillary, scattered along the branches, oblong-cylindric, with numerous globose anthers, their connectives terminating in short spurs, the female terminal or in the axils of upper leaves, composed of spirally arranged ovate rounded scales much shorter than their acutely 2-lobed bracts, with midribs produced into elongated slender tips. Fruit an ovoid-oblong acute pendulous cone maturing in one season, with rounded concave rigid scales persistent on the axis of the cone after the escape of the seeds, and becoming dark red-brown, much shorter than the 2-lobed bracts with midribs ending in rigid woody linear awns, those at the base of the cone without scales and becoming linear-lanceolate by the gradual suppression of their lobes. Seeds nearly triangular, full, rounded and dark-colored on the upper side and pale on the lower side, shorter than their oblong wings infolding the upper side of the seeds in a dark covering; outer seed-coat thick and crustaceous, the inner thin and membranaceous; cotyledons 6—12, much shorter than the inferior radicle.

Pseudotsuga is confined to western North America, southern Japan, southwestern China and Formosa. Four species are recognized.

Pseudotsuga, a barbarous combination of a Greek with a Japanese word, indicates the relation of these trees with the Hemlocks.

CONSPECTUS OF THE NORTH AMERICAN SPECIES.

1. Pseudotsuga taxifolia Britt. Douglas Spruce. Red Fir.

Leaves straight or rarely slightly incurved, rounded and obtuse at apex, or acute on leading shoots, ¾′—1¼′ long, ¹/₁₆′—¹/₁₂′ wide, dark yellow-green or rarely light or dark bluish green, occasionally persistent until their sixteenth year. Flowers: male orange-red; female with slender elongated bracts deeply tinged with red. Fruit pendant on long stout stems, 4′—6½′ long, with thin slightly concave scales rounded and occasionally somewhat elongated at apex, usually rather longer than broad, when fully grown at midsummer slightly puberulous, dark blue-green below, purplish toward the apex, bright red on the closely appressed margins, and pale green bracts becoming slightly reflexed above the middle, ⅕′—¼′ wide, often extending ½′ beyond the scales; seeds light reddish brown and lustrous above, pale and marked below with large irregular white spots, ¼′ long, nearly ⅛′ wide, almost as long as their dark brown wings broadest just below the middle, oblique above and rounded at the apex.

A tree, often 200° high, with a trunk 3°—4° in diameter, frequently taller, with a trunk 10°—12° in diameter, but in the dry interior of the continent rarely more than 80°—100° high, with a trunk hardly exceeding 2°—3° in diameter, slender crowded branches densely clothed with long pendulous lateral branches, forming while the tree is young an open pyramid, soon deciduous from trees crowded in the forest, often leaving the trunk naked for two thirds of its length and surmounted by a comparatively small narrow head sometimes becoming flap-topped by the lengthening of the upper branches, and slender branchlets pubescent for three or four years, pale orange color and lustrous during their first season, becoming bright reddish brown and ultimately dark gray-brown. Winter-buds ovoid, acute, the terminal bud often ¼′ long and nearly twice as large as the lateral buds. Bark on young trees smooth, thin, rather lustrous, dark gray-brown, usually becoming on old trunks 10′—12′ thick, and divided into oblong plates broken into great broad rounded and irregularly connected ridges separating on the surface into small thick closely appressed [49] dark red-brown scales. Wood light, red or yellow, with nearly white sapwood; very variable in density, quality, and in the thickness of the sapwood; largely manufactured into lumber in British Columbia, western Washington and Oregon, and used for all kinds of construction, fuel, railway-ties, and piles; known commercially as “Oregon pine.” The bark is sometimes used in tanning leather.

Distribution. From about latitude 55° north in the Rocky Mountains and from the head of the Skeena River in the coast range, southward through all the Rocky Mountain system to the mountains of western Texas, southern New Mexico and Arizona, and of northern Mexico, and from the Big Horn and Laramie Ranges in Wyoming and from eastern base of the Rocky Mountains of Colorado to the Pacific coast, but absent from the arid mountains in the great basin between the Wahsatch and the Sierra Nevada ranges and from the mountains of southern California; most abundant and of its largest size near the sea-level in the coast region of southern British Columbia and of Washington and Oregon, and on the western foothills of the Cascade Mountains; ascending on the California Sierras to elevations of 5500°, and on the mountains of Colorado to between 6000° and 11,000°, above the sea.

Often planted for timber and ornament in temperate Europe, and for ornament in the eastern and northern states, where only the form from the interior of the continent flourishes. (P. glauca Mayr.)

2. Pseudotsuga macrocarpa Mayr. Hemlock.

Leaves acute or acuminate, terminating in slender rigid callous tips, apparently 2-ranked by the conspicuous twist of their petioles, incurved above the middle, ¾′—1¼′ long, about ¹/₁₆′ wide, dark bluish gray. Flowers: male pale yellow, inclosed for half their length in conspicuous involucres of the lustrous bud-scales; female with pale green bracts tinged with red. Fruit produced on the upper branches and occasionally on those down to the middle of the tree, short-stalked, with scales near the middle of the cone 1½′—2′ across, stiff, thick, concave, rather broader than long, rounded above, abruptly wedge-shaped at the base, puberulous on the outer surface, often nearly as long as their comparatively short and narrow bracts with broad midribs produced into short flattened flexible tips; seeds full and rounded on both sides, rugose, dark chestnut-brown or nearly black and lustrous above, pale reddish brown below, ½′ long, ⅜′ wide, with a thick brittle outer coat, and wings broadest near the middle, about ½′ long, nearly ¼′ wide, and rounded at the apex.

A tree, usually 40°—50° and rarely 90° high, with a trunk 3°—4° in diameter, remote elongated branches pendulous below, furnished with short stout pendant or often erect laterals forming an open broad-based symmetrical pyramidal head, slender branchlets dark reddish [50] brown and pubescent during their first year, becoming glabrous and dark or light orange-brown and ultimately gray-brown. Winter buds ovoid, acute, usually not more than ⅛′ long, often nearly as broad as long. Bark 3′—6′ thick, dark reddish brown, deeply divided into broad rounded ridges covered with thick closely appressed scales. Wood heavy, hard, strong, close-grained, not durable; occasionally manufactured into lumber; largely used for fuel.

Distribution. Steep rocky mountain slopes in southern California at elevations of 3000°—5000° above the sea, often forming open groves of considerable extent, from the Santa Inez Mountains in Santa Barbara County to the Cuyamaca Mountains.

6. ABIES Link. Fir.

Tall pyramidal trees, with bark containing numerous resin-vesicles, smooth, pale, and thin on young trees, often thick and deeply furrowed in old age, pale and usually brittle wood, slender horizontal wide-spreading branches in regular remote 4 or 5-branched whorls, clothed with twice or thrice forked lateral branches forming flat-topped masses of foliage gradually narrowed from the base to the apex of the branch, the ultimate divisions stout, glabrous or pubescent, and small subglobose or ovoid winter branch-buds usually thickly covered with resin, or in one species large and acute, with thin loosely imbricated scales. Leaves linear, sessile, on young plants and on lower sterile branches flattened and mostly grooved on the upper side, or in one species 4-sided, rounded and usually emarginate at apex, appearing 2-ranked by a twist near their base or occasionally spreading from all sides of the branch, only rarely stomatiferous above, on upper fertile branches and leading shoots usually crowded, more or less erect, often incurved or falcate, thick, convex on the upper side, or quadrangular in some species and then obtuse, or acute at apex and frequently stomatiferous on all sides; persistent usually for eight or ten years, in falling leaving small circular scars. Flowers axillary, from buds formed the previous season on branchlets of the year, surrounded at the base by conspicuous involucres of enlarged bud-scales, the male very abundant on the lower side of branches above the middle of the tree, oval or oblong-cylindric with yellow or scarlet anthers surmounted by short knob-like projections, the female usually on the upper side only of the topmost branches, or in some species scattered also over the upper half of the tree, erect, globose, ovoid or oblong, their scales imbricated in many series, obovate, rounded above, cuneate below, much shorter than their acute or dilated mucronate bracts. Fruit an erect ovoid or oblong-cylindric cone, its scales closely imbricated, thin, incurved at the broad apex and generally narrowed below into long stipes, decreasing in size and sterile toward the ends of the cone, falling at maturity with their bracts and seeds from the stout tapering axis of the cone long-persistent on the branch. Seeds furnished with large conspicuous resin-vesicles, ovoid or oblong, acute at base, covered on the upper side and infolded below on the lower side by the base of their thin wing abruptly enlarged at the oblique apex; seed-coat thin, of 2 layers, the outer thick, coriaceous, the inner membranaceous; cotyledons 4—10, much shorter than the inferior radicle.

Abies is widely distributed in the New World from Labrador and the valley of the Athabasca River to the mountains of North Carolina, and from Alaska through the Pacific and Rocky Mountain regions to the highlands of Guatemala, and in the Old World from Siberia and the mountains of central Europe to southern Japan, central China, Formosa, the Himalayas, Asia Minor, and the highlands of northern Africa. Thirty-three species are now recognized. Several exotic species are cultivated in the northern and eastern states; of these the best known and most successful as ornamental trees are Abies Nordmanniana, Spach, of the Caucasus, Abies cilicica Carr., of Asia Minor, Abies cephalonica Loud., a native of Cephalonia, Abies Veitchii Lindl., and Abies homolepis S. & Z., of Japan, and Abies pinsapo, Boiss., of the Spanish Sierra Nevada.

CONSPECTUS OF THE NORTH AMERICAN SPECIES.

1. Abies Fraseri Poir. Balsam Fir. She Balsam.

Leaves obtusely short-pointed or occasionally slightly emarginate at apex, dark green and lustrous on the upper surface, marked on the lower surface by wide bands of 8—12 [52] rows of stomata, ½′ to nearly 1′ long, about ¹/₁₆′ wide. Flowers: male yellow tinged with red; female with scales rounded above, much broader than long and shorter than their oblong pale yellow-green bracts rounded at the broad apex terminating in a slender elongated tip. Fruit oblong-ovoid or nearly oval, rounded at the somewhat narrowed apex, dark purple, puberulous, about 2½′ long, with scales twice as wide as long, at maturity nearly half covered by their pale yellow-green reflexed bracts; seeds ⅛′ long, with dark lustrous wings much expanded and very oblique at apex.

A tree, usually 30°—40° and rarely 70° high, with a trunk occasionally 2½° in diameter, and rather rigid branches forming an open symmetrical pyramid and often disappearing early from the lower part of the trunk, and stout branchlets pubescent for three or four years, pale yellow-brown during their first season, becoming dark reddish brown often tinged with purple, and obtuse orange-brown winter-buds. Bark ¼′—½′ thick, covered with thin closely appressed bright cinnamon-red scales, generally becoming gray on old trees. Wood light, soft, not strong, coarse-grained, pale brown, with nearly white sapwood; occasionally manufactured into lumber.

Distribution: Appalachian Mountains; Cheat Mountain, near Cheat Bridge, Randolph County, West Virginia, and from southwestern Virginia to western North Carolina and eastern Tennessee, often forming forests of considerable extent at elevations between 4000° and 6000° above the sea-level.

Occasionally planted in the parks and gardens of the northern states and of Europe, but short-lived in cultivation and of little value as an ornamental tree.

2. Abies balsamea Mill. Balsam Fir.

Leaves dark green and lustrous on the upper surface, silvery white on the lower surface, with bands of 4—8 rows of stomata, ½′ long on cone-bearing branches to 1¼′ long on the sterile branches of young trees, straight, acute or acuminate, with short or elongated rigid callous tips, spreading at nearly right angles to the branch on young trees and sterile branches, on the upper branches of older trees often broadest above the middle, rounded or obtusely short-pointed at apex, occasionally emarginate on branches at the top of the tree. Flowers: male yellow, more or less deeply tinged with reddish purple; female with nearly orbicular purple scales much shorter than their oblong-obovate serrulate pale yellow-green bracts emarginate with a broad apex abruptly contracted into a long slender recurved tip. Fruit oblong-cylindric, gradually narrowed to the rounded apex, puberulous, dark rich purple, 2′—4′ long, with scales usually longer than broad, generally almost twice as long; rarely not as long as their bracts, (var. phanerolepis Fern.); seeds about ¼′ long and rather shorter than their light brown wings.

A tree, 50°—60° high, with a trunk usually 12′—18′; or rarely 30′ in diameter, spreading branches forming a handsome symmetrical slender-pyramid, the lower branches soon dying from trees crowded in the forest, and slender branchlets pale yellow-green and coated with fine pubescence at first, becoming light gray tinged with red, and often when four or five years old with purple. Winter-buds nearly globose, ⅛′—¼′ in diameter, with lustrous dark orange-green scales. Bark on old trees often ½′ thick, rich brown, much broken on the surface into small plates covered with scales. Wood light, soft, not strong, coarse-grained, perishable, pale brown streaked with yellow, with thick lighter colored sapwood; occasionally made into lumber principally used for packing-cases. From the bark of this tree oil of fir used in the arts and in medicine is obtained.

Distribution. From the interior of the Labrador peninsula westward to the shores of Lesser Slave Lake, southward through Newfoundland, the maritime provinces of Canada, Quebec and Ontario, northern New England, northern New York, northern Michigan to the shores of Saginaw Bay, and northern Minnesota and northeastern Iowa, and along the Appalachian Mountains from western Massachusetts and the Catskills of New York to the high mountains of southwestern Virginia; common and often forming a considerable part of the forest on low swampy ground; on well-drained hillsides sometimes singly in forests of spruce or forming small almost impenetrable thickets; in northern Wisconsin and vicinity occurs a form with longer and more crowded leaves and larger cones (var. macrocarpa Kent); near the timber-line on the mountains of New England and New York reduced to a low almost prostrate shrub.

Sometimes planted in the northern states in the neighborhood of farmhouses, but usually short-lived and of little value as an ornamental tree in cultivation; formerly but now rarely cultivated in European plantations; a dwarf form (var. hudsonica Englm.) growing only a few inches high and spreading into broad nests is often cultivated.

3. Abies lasiocarpa Nutt. Balsam Fir.

Leaves marked on the upper surface but generally only above the middle with 4 or 5 rows of stomata on each side of the conspicuous midrib and on the lower surface by 2 broad bands each of 7 or 8 rows, crowded, nearly erect by the twist at their base, on lower branches 1′—1¾′ long, about ¹/₁₂′ wide, and rounded and occasionally emarginate at apex, on upper branches somewhat thickened, usually acute, generally not more than ½′ long, on leading shoots flattened, closely appressed, with long slender rigid points. Flowers: male dark indigo-blue, turning violet when nearly ready to open; female with dark violet-purple obovate scales much shorter than their strongly reflexed bracts contracted into slender tips. Fruit oblong-cylindric, rounded, truncate or depressed at the narrowed apex, dark purple, puberulous, 2½′—4′ long, with scales gradually narrowed from the broad rounded or nearly truncate apex to the base, usually longer than broad, about three times as long as their oblong-obovate red-brown bracts laciniately cut on the margins, rounded, emarginate and abruptly contracted at the apex into long slender tips; seeds ¼′ long, with dark lustrous wings covering nearly the entire surface of the scales.

A tree, usually 80°—100°, occasionally 175°, or southward rarely more than 50° high, with a trunk 2°—5° in diameter, short crowded tough branches, usually slightly pendulous near the base of the tree, generally clothing the trunks of the oldest trees nearly to their base and forming dense spire-like slender heads, and comparatively stout branchlets coated for three or four years with fine rufous pubescence, or rarely glabrous before the end of their first season, pale orange-brown, ultimately gray or silvery white. Winter-buds subglobose, ⅛′—¼′ thick, covered with light orange-brown scales. Bark becoming on old trees ¾′—1½′ thick, divided by shallow fissures and roughened by thick closely appressed cinnamon-red scales; on the San Francisco Mountains, Arizona, thicker and spongy (var. arizonica Lem.). Wood light, soft, not strong, pale brown or nearly white, with light-colored sapwood; little used except for fuel.

Distribution. High mountain slopes and summits from about latitude 61° in Alaska, southward along the coast ranges to the Olympic Mountains of Washington, over all the [54] high mountain ranges of British Columbia and Alberta, and southward along the Cascade Mountains of Washington and Oregon to the neighborhood of Crater Lake, over the mountain ranges of eastern Washington and Oregon, and of Idaho, Wyoming, Colorado, and Utah to the San Francisco peaks of northern Arizona, and on the Sandia and Mogollon Mountains of New Mexico.

Occasionally planted as an ornamental tree in the northern United States and in northern Europe, but of little value in cultivation.

4. Abies grandis Lindl. White Fir.

Leaves thin and flexible, deeply grooved very dark green and lustrous on upper surface, silvery white on lower surface, with two broad bands of 7—10 rows of stomata, on sterile branches remote, rounded and conspicuously emarginate at apex, 1½′—2¼′ long, usually about ⅛′ wide, spreading in two ranks nearly at right angles to the branch, on cone-bearing branches more crowded, usually 1′—1½′ long, less spreading or nearly erect, blunt-pointed or often notched at apex, on vigorous young trees ½′—¾′ long, acute or acuminate, [55] usually persistent 4—10 years. Flowers: male pale yellow sometimes tinged with purple; female light yellow-green, with semiorbicular scales and short-oblong bracts emarginate and denticulate at the broad obcordate apex furnished with a short strongly reflexed tip. Fruit cylindric, slightly narrowed to the rounded and sometimes retuse apex, puberulous, bright green, 2′—4′ long, with scales usually about two thirds as long as wide, gradually or abruptly narrowed from their broad apex and three or four times as long as their short pale green bracts; seeds ⅜′ in length, light brown, with pale lustrous wings ½′—⅝′ long and nearly as broad as their abruptly widened rounded apex.

A tree, in the neighborhood of the coast 250°—300° high, with a slightly tapering trunk often 4° in diameter, long somewhat pendulous branches sweeping out in graceful curves, and comparatively slender pale yellow-green puberulous branchlets becoming light reddish brown or orange-brown and glabrous in their second season; on the mountains of the interior rarely more than 100° tall, with a trunk usually about 2° in diameter, often smaller and much stunted at high elevations. Winter-buds subglobose, ⅛′—¼′ thick. Bark becoming sometimes 2′ thick at the base of old trees and gray-brown or reddish brown and divided by shallow fissures into low flat ridges broken into oblong plates roughened by thick closely appressed scales. Wood light, soft, coarse-grained, not strong nor durable, light brown, with thin lighter colored sapwood; occasionally manufactured into lumber in western Washington and Oregon and used for the interior finish of buildings, packing-cases, and wooden-ware.

Distribution. Northern part of Vancouver Island southward in the neighborhood of the coast to northern Sonoma County, California, and along the mountains of northern Washington and Idaho to the western slopes of the continental divide in northern Montana, and to the mountains of eastern Oregon; near the coast scattered on moist ground through forests of other conifers; common in Washington and northern Oregon from the sea up to elevations of 4000°; in the interior on moist slopes in the neighborhood of streams from 2500° up to 7000° above the sea; in California rarely ranging more than ten miles inland or ascending to altitudes of more than 1500° above the sea.

Occasionally planted in the parks and gardens of temperate Europe, where it grows rapidly and promises to attain a large size; rarely planted in the United States.

5. Abies concolor Lindl. & Gord. White Fir.

Leaves crowded, spreading in 2 ranks and more or less erect from the strong twist at their base, pale blue or glaucous, becoming dull green at the end of two or three years, with 2 broad bands of stomata on the lower, and more or less stomatiferous on the upper surface, on lower branches flat, straight, rounded, acute or acuminate at apex, 2′—3′ long, about ¹/₁₆′ wide, on fertile branches and on old trees frequently thick, keeled above, usually falcate, acute or rarely notched at apex, ¾′—1½′ long, often ⅛′ wide. Flowers: male dark red or rose color; female with broad rounded scales, and oblong strongly reflexed obcordate bracts laciniate above the middle and abruptly contracted at apex into short points. Fruit oblong, slightly narrowed from near the middle to the ends, rounded or obtuse at apex, 3′—5′ long, puberulous, grayish green, dark purple or bright canary-yellow, with scales much broader than long, gradually and regularly narrowed from the rounded apex, rather more than twice as long as their emarginate or nearly truncate bracts broad at the apex and terminating in short slender tips; seeds ⅓′—½′ long, acute at base, dark dull brown, with lustrous rose-colored wings widest near the middle and nearly truncate at apex.

A tree, on the California sierras 200°—250° high, with a trunk often 6° in diameter or in the interior of the continent rarely more than 125° tall, with a trunk seldom exceeding 3° in diameter, a narrow spire-like crown of short stout branches clothed with long lateral branches pointing forward and forming great frond-like masses of foliage, and glabrous lustrous comparatively stout branchlets dark orange color during their first season, becoming light grayish green or pale reddish brown, and ultimately gray or grayish brown. Winter-buds subglobose, ⅛′—¼′ thick. Bark becoming on old trunks sometimes 5′—6′ thick near the ground and deeply divided into broad rounded ridges broken on the surface into irregularly [56] shaped plate-like scales. Wood very light, soft, coarse-grained and not strong nor durable, pale brown or sometimes nearly whiter occasionally manufactured into lumber, in northern California used for packing-cases and butter-tubs.

Distribution. Rocky Mountains of southern Colorado, westward to the mountain ranges of California, extending northward into northern Oregon, and southward over the mountains of New Mexico and Arizona into northern Mexico and Lower California (Mt. San Pedro Mártir Mountains); the only Fir-tree in the arid regions of the Great Basin, of southern New Mexico and Arizona, and of the mountain forests of southern California.

Often planted as an ornamental tree in Europe (the California form usually as A. Lowiana Murr.) and in the eastern states where it grows more vigorously than other Fir-trees.

6. Abies amabilis Forbes. White Fir.

Leaves deeply grooved, very dark green and lustrous on the upper surface, silvery white on the lower, with broad bands of 6 or 8 rows of stomata between the prominent midribs and incurved margins, on sterile branches obtuse and rounded, or notched or occasionally acute at apex, ¾′—1¼′ long, ¹/₁₆′—¹/₁₂′ wide, often broadest above the middle, erect by a twist at their base, very crowded, those on the upper side of the branch much shorter than those on the lower and usually parallel with and closely appressed against it, on fertile branches acute or acuminate with callous tips, occasionally stomatiferous on the upper surface near the apex, ½′—¾′ long; on vigorous leading shoots acute, with long rigid points, closely appressed or recurved near the middle, about ¾′ long and nearly ⅛′ wide. Flowers: male red; female with broad rounded scales and rhombic dark purple lustrous bracts erose above the middle and gradually contracted into broad points. Fruit oblong, slightly narrowed to the rounded and often retuse apex, deep rich purple, puberulous, 3½′—6′ long, with scales 1′—1⅛′ wide, nearly as long as broad, gradually narrowed from the rounded apex and rather more than twice as long as their reddish rhombic or oblong-obovate bracts terminating in long slender tips; seeds light yellow-brown, ½′ long, with oblique pale brown lustrous wings about ¾′ long.

A tree, often 250° tall, or at high altitudes and in the north usually not more than 70°—80° tall, with a trunk 4°—6° in diameter, in thick forests often naked for 150°, but in open situations densely clothed to the ground with comparatively short branches sweeping down in graceful curves, and stout branchlets clothed for four or five years with soft fine pubescence, light orange-brown in their first season, becoming dark purple and ultimately reddish brown. Winter-buds nearly globose, ⅛′—¼′ thick, with closely imbricated lustrous purple scales. Bark on trees up to 150 years old thin, smooth, pale or silvery white, [57] becoming near the ground on old trees 1½′—2½′ thick, and irregularly divided into comparatively small plates covered with small closely appressed reddish brown or reddish gray scales. Wood light, hard, not strong, close-grained, pale brown, with nearly white sapwood; in Washington occasionally manufactured into lumber used in the interior finish of buildings.

Distribution. High mountain slopes and benches from southeastern Alaska (Boca de Quadra Inlet and Sandfly Bay), to Vancouver Island and southward along the coast ranges to Saddle Mountain near Astoria, Oregon, and on the Cascade Mountains to the slopes of Old Bailey Mountain, Oregon, ranging from the sea level at the north to elevations of from 3000°—6000° southward; attaining its largest size on the Olympic Mountains of Washington, where it is the most common Fir-tree.

Occasionally cultivated as an ornamental tree in the eastern states and in western Europe, but without developing the beauty which distinguishes this species in its native forests.

7. Abies nobilis Lindl. Red Fir.

Leaves marked on the upper surface with a deep sharply defined groove, rounded and obscurely ribbed on the lower surface, stomatiferous above and below, dark or light blue-green, often very glaucous during their first season, crowded in several rows, those on the lower side of the branch two-ranked by the twisting of their bases, the others crowded, strongly incurved, with the points erect or pointing away from the end of the branch, on young plants and on the lower sterile branches of old trees flat, rounded, usually slightly notched at apex, 1′—1½′ long, about ¹/₁₆′ wide, on fertile branches much thickened and almost equally 4-sided, acuminate, with long rigid callous tips, ½′—¾′ long, on leading shoots flat, gradually narrowed from the base, acuminate, with long rigid points, about 1′ long. Flowers: male reddish purple; female often scattered over the upper part of the tree, with broad rounded scales much shorter than their nearly orbicular bracts erose on the margins and contracted above into slender elongated strongly reflexed tips. Fruit oblong-cylindric, slightly narrowed but full and rounded at apex, 4′—5′ long, purple or olive-brown, pubescent, with scales about one third wider than long, gradually narrowed from the rounded apex to the base, or full at the sides, rounded and denticulate above the middle and sharply contracted and wedge-shaped below, nearly or entirely covered by their strongly reflexed pale green spatulate bracts full and rounded above, fimbriate on the margins, with broad midribs produced into short broad flattened points; seeds ½′ long, pale reddish brown, about as long as their wings, gradually narrowed from below to the nearly truncate slightly rounded apex.

A tree, in old age with a comparatively broad somewhat rounded head, usually 150°—200° and occasionally 250° high, with a trunk 6°—8° in diameter, short rigid branches, short stout remote lateral branches standing out at right angles, and slender reddish brown branchlets [58] puberulous for four or five years and generally pointing forward. Winter-buds ovoid-oblong, red-brown, about ⅛′ long. Bark becoming on old trunks 1′—2′ thick, bright red-brown, and deeply divided into broad flat ridges irregularly broken by cross fissures and covered with thick closely appressed scales. Wood light, hard, strong, rather close-grained, pale brown streaked with red, with darker colored sapwood; occasionally manufactured into lumber and used under the name of larch for the interior finish of buildings and for packing-cases.

Distribution. Slopes of Mt. Baker in northern Washington and southward to the valley of the Mackenzie River, Oregon, and the Siskiyou Mountains, California, at elevations of from 2000°—5000° above the sea; most abundant and often forming extensive forests on the Cascade Mountains of Washington; less abundant and of smaller size on the eastern and northern slopes of these mountains. In Oregon sometimes called Larch.

Often planted in western and central Europe as an ornamental tree, and in the eastern states hardy in sheltered positions as far north as Massachusetts.

8. Abies magnifica A. Murr. Red Fir.

Leaves almost equally 4-sided, ribbed above and below, with 6—8 rows of stomata on each of the 4 sides, pale and very glaucous during their first season, later becoming blue-green, persistent usually for about ten years; on young plants and lower branches oblanceolate, somewhat flattened, rounded, bluntly pointed, ¾′—1½′ long, ¹/₁₆′ wide, those on the lower side of the branch spreading in 2 nearly horizontal ranks by the twist at their base, on upper, especially on fertile branches, much thickened, with more prominent [59] midribs, acute, with short callous tips, ⅓′ long on the upper side of the branch to 1¼′ long on the lower side, crowded, erect, strongly incurved, completely hiding the upper side of the branch, on leading shoots ¾′ long, erect and acuminate, with long rigid points pressed against the stem. Flowers: male dark reddish purple; female with rounded scales much shorter than their oblong pale green bracts terminating in elongated slender tips more or less tinged with red. Fruit oblong-cylindric, slightly narrowed to the rounded, truncate, or retuse apex, dark purplish brown, puberulous, from 6′—9′ long, with scales often 1½′ wide and about two thirds as wide as long, gradually narrowed to the cordate base, somewhat longer or often two thirds as long as their spatulate acute or acuminate bracts slightly serrulate above the middle and often sharply contracted and then enlarged toward the base; seeds dark reddish brown, ¾′ long, about as wide as their lustrous rose-colored obovate cuneate wings nearly truncate and often ¾′ wide at apex.

A tree, in old age occasionally somewhat round-topped, frequently 200° high, with a trunk 8°—10° in diameter and often naked for half the height of the tree, comparatively short small branches, the upper somewhat ascending, the lower pendulous, and stout light yellow-green branchlets pointing forward, slightly puberulous during their first season, becoming light red-brown and lustrous and ultimately gray or silvery white. Winter-buds ovoid, acute, ¼′⅓′ long, their bright chestnut-brown scales with prominent midribs produced into short tips. Bark becoming 4′—6′ thick near the ground, deeply divided into broad rounded ridges broken by cross fissures and covered by dark red-brown scales. Wood light, soft, not strong, comparatively durable, light red-brown, with thick somewhat darker sapwood; largely used for fuel, and in California occasionally manufactured into coarse lumber employed in the construction of cheap buildings and for packing-cases.

Distribution. Cascade Mountains of southern Oregon, southward over the mountain ranges of northern California (summits of the Trinity and Salmon Mountains and on the inner north coast ranges), and along the western slope of the Sierra Nevada to the divide between White and Kern Rivers; common in southern Oregon at elevations between 5000° and 7000° above the sea, forming sometimes nearly pure forests; very abundant on the Sierra Nevada, and the principal tree in the forest belt at elevations between 6000° and 9000°; ascending towards the southern extremity of its range to over 10,000°. Small stunted trees from the neighborhood of Meadow Lake, Sierra County, California, with yellowish cones have been described as var. xancocarpa Lemm.

Often planted as an ornamental tree in western and central Europe, and sometimes hardy in the United States as far north as eastern Massachusetts.

Abies magnifica var. shastensis Lemm. Red Fir.

On the mountains of southern Oregon and at high elevations on those of northern California, and on the southern Sierra Nevada, occurs this form distinguished only by the [60] longer rounded or obtusely pointed (not acute) bright yellow bracts which sometimes cover nearly half their scales.

9. Abies venusta K. Koch. Silver Fir.

Leaves thin, flat, rigid, linear or linear-lanceolate, gradually or abruptly narrowed toward the base, often falcate, especially on fertile branches, acuminate, with long slender callous tips, dark yellow-green, lustrous and slightly rounded on the upper surface marked below the middle with an obscure groove, silvery white or on old leaves pale on the lower surface, with bands of 8—10 rows of stomata between the broad midrib and the thickened strongly revolute margins, 2-ranked from the conspicuous twist near their base and spreading at nearly right angles to the branch, or pointing forward on upper fertile branches, 1½′—2¼′ long, on leading shoots standing out at almost right angles, rounded on the upper surface, more or less incurved above the middle, 1½′—1¾′ long, about ⅛′ wide. Flowers: male produced in great numbers near the base of the branchlets on branches from the middle of the tree upward, pale yellow; female near the ends of the branchlets of the upper branches only, with oblong scales rounded above and nearly as long as their cuneate obcordate yellow-green bracts ending in slender elongated awns. Fruit on stout peduncles sometimes ½′ long, oval or subcylindric, full and rounded at apex, glabrous, pale purple-brown, 3′—4′ long, with thin scales strongly incurved above, obtusely short-pointed at apex, obscurely denticulate on the thin margins, about one third longer than their oblong-obovate obcordate pale yellow-brown bracts terminating in flat rigid tips 1′—1¾′ long, above the middle of the cone pointing toward its apex and often closely appressed to its sides, below the middle spreading toward its base and frequently much recurved, firmly attached to the cone-scales and deciduous with them from the thick conical sharp-pointed axis of the cone; seeds dark red-brown, about ⅜′ long, and nearly as long as their oblong-obovate pale reddish brown lustrous wings rounded at the apex.

A tree, 100°—150° high, with a trunk sometimes 3° in diameter, comparatively short slender usually pendulous branches furnished with long sinuous rather remote lateral branches sparsely clothed with foliage, forming a broad-based pyramid abruptly narrowed 15°—20° from the top of the tree into a thin spire-like head, and stout glabrous light reddish brown branchlets covered at first with a glaucous bloom. Winter-buds ovoid, acute, ¾′—1′ long, ¼′—⅓′ thick, with very thin, loosely imbricated, pale chestnut-brown, acute, boat-shaped scales. Bark becoming near the base of the tree ½′—¾′ thick, light reddish brown, slightly [61] and irregularly fissured and broken into thick closely appressed scales. Wood heavy, not hard, coarse-grained, light brown tinged with yellow, with paler sapwood.

Distribution. In the moist bottoms of cañons and on dry rocky summits, usually at elevations of about 3000° above the sea on both slopes of the outer western ridge of the Santa Lucia Mountains, Monterey County, California.

Occasionally and successfully grown as an ornamental tree in the milder parts of Great Britain and in northern Italy; not hardy in the eastern United States.

7. SEQUOIA Endl.

Resinous aromatic trees, with tall massive lobed trunks, thick bark of 2 layers, the outer composed of fibrous scales, the inner thin, close and firm, soft, durable, straight-grained red heartwood, thin nearly white sapwood, short stout horizontal branches, terete lateral branchlets deciduous in the autumn, and scaly or naked buds. Leaves ovate-lanceolate or linear and spreading in 2 ranks especially on young trees and branches, or linear, acute, compressed, keeled on the back and closely appressed or spreading at apex, the two forms appearing sometimes on the same branch or on different branches of the same tree. Flowers minute, solitary, monœcious, appearing in early spring from buds formed the previous autumn, the male terminal in the axils of upper leaves, oblong or ovoid, surrounded by an involucre of numerous imbricated ovate, acute, and apiculate bracts, with numerous spirally disposed filaments dilated into ovoid acute subpeltate denticulate connectives bearing on their inner face 2—5 pendulous globose 2-valved anther-cells; the female terminal, ovoid or oblong, composed of numerous spirally imbricated ovate scales abruptly keeled on the back, the keels produced into short or elongated points closely adnate to the short ovule-bearing scales rounded above and bearing below their upper margin in 2 rows 5—7 ovules at first erect, becoming reversed. Fruit an ovoid or short-oblong pendulous cone maturing during the first or second season, persistent after the escape of the seeds, its scales formed by the enlargement of the united flower and ovuliferous scales, becoming woody, bearing large deciduous resin-glands, gradually enlarged upward and widening at the apex into a narrow thickened oblong disk transversely depressed through the middle and sometimes tipped with a small point. Seeds 5—7 under each scale, oblong-ovoid, compressed; seed-coat membranaceous, produced into broad thin lateral wings; cotyledons 4—6, longer than the inferior radicle.

Sequoia, widely scattered with several species over the northern hemisphere during the cretaceous and tertiary epochs, is now confined to the coast of Oregon and California and the mountains of California, where two species exist.

The name of the genus is formed from Sequoiah, the inventor of the Cherokee alphabet.

CONSPECTUS OF THE NORTH AMERICAN SPECIES.

1. Sequoia sempervirens Endl. Redwood.

Leaves of secondary branches and of lower branches of young trees lanceolate, more or less falcate, acute or acuminate and usually tipped with slender rigid points, slightly thickened on the revolute margins, decurrent at the base, spreading in 2 ranks by a half-turn at their base, ¼′—½′ long, about ⅛′ wide, obscurely keeled and marked above by 2 narrow bands of stomata, glaucous and stomatiferous below on each side of their conspicuous midrib, on leading shoots disposed in many ranks, more or less spreading or appressed, ovate or ovate-oblong, incurved at the rounded apiculate apex, thickened, rounded, and stomatiferous on the lower surface, concave, prominently keeled and covered with stomata [62] on the upper surface, usually about ¼′ long; dying and turning reddish brown at least two years before falling. Flowers opening in December or January; male oblong, obtuse; female with about 20 broadly ovate acute scales tipped with elongated and incurved or short points. Fruit ripening in October, oblong, ¾′—1′ long, ½′ broad, its scales gradually enlarged from slender stipes abruptly dilated above into disks penetrated by deep narrow grooves, and usually without tips; seeds about ¹/₁₆′ long, light brown, with wings as broad as their body.

A tree, from 200°—340° high, with a slightly tapering and irregularly lobed trunk usually free of branches for 75°—100°, usually 10°—15°, rarely 28° in diameter at the much buttressed base, slender branches, clothed with branchlets spreading in 2 ranks and forming while the tree is young an open narrow pyramid, on old trees becoming stout and horizontal, and forming a narrow rather compact and very irregular head remarkably small in proportion to the height and size of the trunk, and slender leading branchlets covered at the end of three or four years after the leaves fall with cinnamon-brown scaly bark; when cut producing from the stump numerous vigorous long-lived shoots. Buds with numerous loosely imbricated ovate acute scales persistent on the base of the branchlet. Bark 6′—12′ thick, divided into rounded ridges and separated on the surface into long narrow dark brown fibrous scales often broken transversely and in falling disclosing the bright cinnamon-red inner bark. Wood light, soft, not strong, close-grained, easily split and worked, very durable in contact with the soil, clear light red; largely manufactured into lumber and used for shingles, fence-posts, railway-ties, wine-butts, and in buildings.

Distribution. Valley of the Chetco River, Oregon, 8 miles north of the California state line, southward near the coast to Monterey County, California; rarely found more than twenty or thirty miles from the coast, or beyond the influence of the ocean fogs, or over 3000° above the sea-level; often forming in northern California pure forests occupying the sides of ravines and the banks of streams; southward growing usually in small groves scattered among other trees; most abundant and of its largest size north of Cape Mendocino.

Often cultivated as an ornamental tree in the temperate countries of Europe, and occasionally in the southeastern United States.

2. Sequoia gigantea Decne. Big Tree.

Leaves ovate and acuminate, or lanceolate, rounded and thickened on the lower surface, concave on the upper surface, marked by bands of stomata on both sides of the obscure midrib, rigid, sharp-pointed, decurrent below, spreading or closely appressed above the middle, ⅛′—¼′ or on leading shoots ½′ long. Flowers opening in late winter and early spring; male in great profusion over the whole tree, oblong-ovoid, with ovate acute or acuminate connectives; female with 25—40 pale yellow scales slightly keeled on the back and gradually [63] narrowed into long slender points. Fruit maturing in the second year, ovoid-oblong, 2′—3½′ long, 1½′—2¼′ wide, dark reddish brown, the scales gradually thickened upward from the base to the slightly dilated apex, ¾′—1¼′ long, and ¼′—½′ wide, deeply pitted in the middle, often furnished with an elongated reflexed tip and on the upper side near the base with two or three large deciduous resin-glands; seeds linear-lanceolate, compressed, ⅛′—¼′ long, light brown, surrounded by laterally united wings broader than the body of the seed, apiculate at the apex, often very unequal.

A tree, at maturity usually about 275° high, with a trunk 20° in diameter near the ground, occasionally becoming 320° tall, with a trunk 35° in diameter, much enlarged and buttressed at base, fluted with broad low rounded ridges, in old age naked often for 150° with short thick horizontal branches, slender leading branchlets becoming after the disappearance of the leaves reddish brown more or less tinged with purple and covered with thin close or slightly scaly bark and naked buds. Bark 1°—2° thick, divided into rounded lobes 4°—5° wide, corresponding to the lobes of the trunk, separating into loose light cinnamon-red fibrous scales, the outer scales slightly tinged with purple. Wood very light, soft, not strong, brittle and coarse-grained, turning dark on exposure; manufactured into lumber and used for fencing, in construction, and for shingles.

Distribution. Western slopes of the Sierra Nevada of California, in an interrupted belt at elevations of 5000°—8400° above the level of the sea, from the middle fork of the American River to the head of Deer Creek just south of latitude 36°; north of King’s River in isolated groves, southward forming forests of considerable extent, and best developed on the north fork of the Tule River.

Universally cultivated as an ornamental tree in all the countries of western and southern Europe; and occasionally in the middle eastern United States.

8. TAXODIUM Rich. Bald Cypress.

Resinous trees, with furrowed scaly bark, light brown durable heartwood, thin white sapwood, erect ultimately spreading branches, deciduous usually 2-ranked lateral branchlets, scaly globose buds, and stout horizontal roots often producing erect woody projections (knees). Leaves spirally disposed, pale and marked with stomata below on both sides of the obscure midrib, dark green above, linear-lanceolate, spreading in 2 ranks, or scale-like and appressed on lateral branchlets, the two forms appearing on the same or on different branches of the same tree or on separate trees, deciduous. Flowers unisexual, from buds formed the previous year; male in the axils of scale-like bracts in long terminal drooping panicles, with 6—8 stamens opposite in 2 ranks, their filaments abruptly enlarged [64] into broadly ovate peltate yellow connectives bearing on their inner face in 2 rows 4—9 2-valved pendulous anther-cells; female scattered near the ends of the branches of the previous year, subglobose, composed of numerous ovate spirally arranged long-pointed scales, adnate below to the thickened fleshy ovuliferous scales bearing at their base 2 erect bottle-shaped ovules. Fruit a globose or obovoid short-stalked woody cone maturing the first year and persistent after the escape of the seeds, formed from the enlargement and union of the flower and ovule-bearing scales abruptly dilated from slender stipes into irregularly 4-sided disks often mucronate at maturity, bearing on the inner face, especially on the stipes, large dark glands filled with blood-red fragrant liquid resin. Seeds in pairs under each scale, attached laterally to the stipes, erect, unequally 3-angled; seed-coat light brown and lustrous, thick, coriaceous or corky, produced into 3 thick unequal lateral wings and below into a slender elongated point; cotyledons 4—9, shorter than the superior radicle.

Taxodium, widely distributed through North America and Europe in Miocene and Pliocene times, is now confined to the southern United States and Mexico. Two species are distinguished.

The generic name, from τάξος and εῖδος, indicates a resemblance of the leaves to those of the Yew-tree.

1. Taxodium distichum Rich. Bald Cypress. Deciduous Cypress.

Leaves on distichously spreading branchlets, apiculate, ½′—¾′ long, about ¹/₁₂′ wide, light bright yellow-green or occasionally silvery white below; or on the form with pendulous compressed branchlets long-pointed, keeled and stomatiferous below, concave above more or less spreading at the free apex, about ½′ long; in the autumn turning with the branchlets dull orange-brown before falling. Flowers: panicles of staminate flowers 4′—5′ long, 1½′—2′ wide, with slender red-brown stems, obovoid flower-buds nearly ⅛′ long, pale silvery-gray during winter and purple when the flowers expand in the spring. Fruit usually produced in pairs at the end of the branch or irregularly scattered along it for several inches, nearly globose or obovoid, rugose, about 1′ in diameter, the scales generally destitute of tips; seeds with wings nearly ¼′ long, ⅛′ wide.

A tree, with a tall lobed gradually tapering trunk, rarely 12° and generally 4°—5° in diameter above the abruptly enlarged strongly buttressed usually hollow base, occasionally 150° tall, in youth pyramidal, with slender branches often becoming elongated and slightly pendulous, in old age spreading out into a broad low rounded crown often 100° across, and slender branchlets light green when they first appear, light red-brown and rather lustrous [65] during their first winter, becoming darker the following year, the lateral branchlets deciduous, 3′—4′ long, spreading at right angles to the branch, or in the form with acicular leaves pendulous or erect and often 6′ long. Bark 1′—2′ thick, light cinnamon-red and divided by shallow fissures into broad flat ridges separating on the surface into long thin closely appressed fibrous scales. Wood light, soft, not strong, easily worked, light or dark brown, sometimes nearly black; largely used for construction, railway-ties, posts, fences, and in cooperage.

Distribution. River swamps usually submerged during several months of the year, low wet banks of streams, and the wet depressions of Pine-barrens from southern New Jersey and southern Delaware southward generally near the coast to the Everglade Keys, southern Florida, and through the Gulf-coast region to the valley of Devil River, Texas, through Louisiana to southern Oklahoma, through southern and western Arkansas to southeastern Missouri, and through western and northern Mississippi to Tishomingo County, and in western Tennessee and Kentucky to southern Illinois and southwestern Indiana; most common and of its largest size in the south Atlantic and Gulf states, often covering with nearly pure forests great river swamps. From the coast of North Carolina to southern Florida, southern Alabama and eastern and western Louisiana the form with acicular leaves (Taxodium distichum var. imbricarium, Sarg.) is not rare as a small tree in Pine-barren ponds and swamps.

Often cultivated as an ornamental tree in the northern United States, and in the countries of temperate Europe, especially the var. imbricarium (as Glyptostrobus sinensis Hort. not Endl.).

9. LIBOCEDRUS Endl.

Tall resinous aromatic trees, with scaly bark, spreading branches, flattened branchlets disposed in one horizontal plane and forming an open 2-ranked spray and often ultimately deciduous, straight-grained durable fragrant wood, and naked buds. Leaves scale-like, in 4 ranks, on leading shoots nearly equally decussate, closely compressed or spreading, dying and becoming woody before falling, on lateral flattened branchlets much compressed, conspicuously keeled, and nearly covering those of the other ranks; on seedling plants linear-lanceolate and spreading. Flowers monœcious, solitary, terminal, the two sexes on different branchlets; male oblong, with 12—16 decussate filaments dilated into broad connectives usually bearing 4 subglobose anther-cells; female oblong, subtended at base by several pairs of leaf-like scales slightly enlarged and persistent under the fruit, composed of 6 acuminate short-pointed scales, those of the upper and middle ranks much larger than those of the lower rank, ovate or oblong, fertile and bearing at the base of a minute accrescent ovuliferous scale 2 erect ovules. Fruit an oblong cone maturing in one season, with subcoriaceous scales marked at the apex by the free thickened mucronulate border of the enlarged flower-scales, those of the lowest pair ovate, thin, reflexed, much shorter than the oblong thicker scales of the second pair widely spreading at maturity; those of the third pair confluent into an erect partition. Seeds in pairs, erect on the base of the scale; seed-coat membranaceous, of 2 layers, produced into thin unequal lateral wings, one narrow, the other broad, oblique, nearly as long as the scale; cotyledons 2, about as long as the superior radicle.

Libocedrus is confined to western North America, western South America, where it is distributed from Chile to Patagonia, New Zealand, New Caledonia, New Guinea, Formosa, and southwestern China. Eight species are distinguished.

Libocedrus, from λιβάς and Cedrus, relates to the resinous character of these trees.

1. Libocedrus decurrens Torr. Incense Cedar.

Leaves oblong-obovate, decurrent and closely adnate on the branchlets except at the callous apex, ⅛′ long on the ultimate lateral branchlets to nearly ½′ long on leading shoots, those of the lateral ranks gradually narrowed and acuminate at apex, keeled and glandular [66] on the back, and nearly covering the flattened obscurely glandular-pitted and abruptly pointed leaves of the inner ranks. Flowers appearing in January on the ends of short lateral branchlets of the previous year; male tingeing the tree with gold during the winter and early spring, ovate, nearly ¼′ long, with nearly orbicular or broadly ovate connectives, rounded, acute or acuminate at the apex and slightly erose on the margins; female subtended by 2—6 pairs of leaf-like scales, with ovate acute light yellow-green slightly spreading scales. Fruit ripening and discharging its seeds in the autumn, oblong, ¾′—1′ long, pendulous, light red-brown; seeds oblong-lanceolate, ⅓′—½′ long, semiterete and marked below by a conspicuous pale basal hilum; inner layer of the seed-coat penetrated by elongated resin-chambers, filled with red liquid balsamic resin.

A tree, usually 80°—100° or rarely 150° high, with a tall straight slightly and irregularly lobed trunk tapering from a broad base, 3° or 4° or occasionally 6° or 7° in diameter, slender branches erect at the top of the tree, below sweeping downward in bold curves, forming a narrow open feathery crown becoming in old age irregular in outline by the greater development of a few ultimately upright branches forming secondary stems, and stout branchlets somewhat flattened and light yellow-green at first, turning light red-brown during the summer and ultimately brown more or less tinged with purple, the lateral branchlets much flattened, 4′—6′ long, and usually deciduous at the end of the second or third season. Bark ½′—1′ thick, bright cinnamon-red, and broken into irregular ridges covered with closely appressed plate-like scales. Wood light, soft, close-grained very durable in contact with the soil, light reddish brown, with thin nearly white sapwood; often injured by dry rot but largely used for fencing, laths and shingles, the interior finish of buildings, for furniture, and in the construction of flumes.

Distribution. Singly or in small groves from the southeastern slope of Mt. Hood, Oregon, and southward along the Cascade Mountains; on the high mountains of northern California, on the western slopes of the Sierra Nevada, and in Alpine County on their eastern slope, on the Washoe Mountains, western Nevada, in the California coast ranges from the Santa Lucia Mountains, Monterey County to the high mountains in the southern part of the state; on the Sierra del Pinal and the San Pedro Mártir Mountains, Lower California; most abundant and of its largest size on the Sierra Nevada, of central California at elevations of 5000°—7000° above the sea.

Often cultivated as an ornamental tree in western and central Europe, where it grows rapidly and promises to attain a large size; hardy and occasionally planted in the New England and middle Atlantic states.

10. THUJA L. Arbor-vitæ.

Resinous aromatic trees, with thin scaly bark, soft durable straight-grained heartwood, thin nearly white sapwood, slender spreading or erect branches, pyramidal heads, flattened lateral pendulous branchlets disposed in one horizontal plane, forming a flat frond-like spray and often finally deciduous, and naked buds. Leaves decussate, scale-like, acute, stomatiferous on the back, on leading shoots appressed or spreading, rounded or slightly keeled on the back, narrowed into long slender points, on lateral branchlets much compressed in the lateral ranks, prominently keeled and nearly covering those of the other ranks; on seedling plants linear-lanceolate, acuminate, spreading or reflexed. Flowers minute, monœcious, from buds formed the previous autumn, terminal, solitary, the two sexes usually on different branchlets; male ovoid, with 4—6 decussate filaments, enlarged into suborbicular peltate connectives bearing on their inner face 2—4 subglobose anther-cells; female oblong, with 8—12 oblong acute scales opposite in pairs, the ovuliferous scales at their base bearing usually 2 erect bottle-shaped ovules. Fruit an ovoid-oblong erect pale cinnamon-brown cone maturing in one season, its scales thin (thick in one species), leathery, oblong, acute, marked near the apex by the thickened free border of the enlarged flower-scales, those of the 2 or 3 middle ranks largest and fertile. Seeds usually 2, erect on the base of the scale, ovoid, acute, compressed, light chestnut-brown; seed-coat membranaceous, produced except in one species into broad lateral wings distinct at the apex; cotyledons 2, longer than the superior radicle.

Thuja is confined to northeastern and northwestern America, to Japan, Korea and northern China. Five species are recognized. Of the exotic species the Chinese Thuja orientalis, L., with many varieties produced by cultivation, is frequently planted in the United States, especially in the south and west, for the decoration of gardens, and is distinguished from the other species by the thick umbonate scales of the cone, only the 4 lower scales being fertile, and by the thick rounded dark red-purple seeds without wings.

CONSPECTUS OF THE NORTH AMERICAN SPECIES.

1. Thuja occidentalis L. White Cedar. Arbor-vitæ.

Leaves on leading shoots often nearly ¼′ long, long-pointed and usually conspicuously glandular, on lateral branchlets much flattened, rounded and apiculate at apex, without glands or obscurely glandular-pitted, about ⅛′ long. Flowers opening in April and May, liver color. Fruit ripening and discharging its seeds in the early autumn, ⅓′—½′ long; seeds ⅛′ long, the thin wings as wide as the body.

A tree, 50°—60° high, with a short often lobed and buttressed trunk, occasionally 6° although usually not more than 2°—3° in diameter, often divided into 2 or 3 stout secondary stems, short horizontal branches soon turning upward and forming a narrow compact pyramidal head, light yellow-green branchlets paler on the lower surface than on the upper, changing with the death of the leaves during their second season to light cinnamon-red, growing darker the following year, gradually becoming terete and abruptly enlarged at the base and finally covered with smooth lustrous dark orange-brown bark, and marked by conspicuous scars left by the falling of the short pendulous lateral branchlets. Bark ¼′—⅓′ thick, light red-brown often tinged with orange color and broken by shallow fissures into narrow flat connected ridges separating into elongated more or less persistent scales. Wood light, soft, brittle, very coarse-grained, durable, fragrant, pale yellow-brown; largely used in Canada and the northern states for fence-posts, rails, railway-ties, and shingles. Fluid extracts and tinctures made from the young branchlets are sometimes used in medicine.

Distribution. Frequently forming nearly impenetrable forests on swampy ground or [68] often occupying the rocky banks of streams, from Nova Scotia and New Brunswick, northwestward to the mouth of the Saskatchewan, and southward through eastern Canada to southern New Hampshire, central Massachusetts, New York, central Ohio, northern Indiana and Illinois, and Minnesota; occasionally on the high mountains of Virginia, West Virginia, and northeastern Tennessee, and on the mountains of western Burke County, North Carolina, at an altitude of 3000 feet; very common at the north, less abundant and of smaller size southward.

Often cultivated, with many, often dwarf, forms produced in nurseries, as an ornamental tree and for hedges; and in Europe from the middle of the sixteenth century.

2. Thuja plicata D. Don. Red Cedar. Canoe Cedar.

Leaves on leading shoots ovate, long-pointed, often conspicuously glandular on the back, frequently ¼′ long, on lateral branchlets ovate, apiculate, without glands or obscurely glandular-pitted, usually not more than ⅛′ long, mostly persistent 2—5 years. Flowers about ¹/₁₂′ long, dark brown. Fruit ripening early in the autumn, clustered near the ends of the branches, much reflexed, ½′ long, with thin leathery scales, conspicuously marked near the apex by the free border of the flower-scale furnished with short stout erect or recurved dark mucros; seeds often 3 under each fertile scale, rather shorter than their usually slightly unequal wings about ¼′ long.

A tree, frequently 200° high, with a broad gradually tapering buttressed base sometimes 15° in diameter at the ground and in old age often separating toward the summit into 2 or 3 erect divisions, short horizontal branches, usually pendulous at the ends, forming a dense narrow pyramidal head, and slender much compressed branchlets often slightly zigzag, light bright [69] yellow-green during their first year, then cinnamon-brown, and after the falling of the leaves, lustrous and dark reddish brown often tinged with purple, the lateral branchlets 5′—6′ long, light green and lustrous on the upper surface, somewhat paler on the lower surface, turning yellow and falling generally at the end of their second season. Bark bright cinnamon-red, ½′—¾′ thick, irregularly divided by narrow shallow fissures into broad ridges rounded on the back and broken on the surface into long narrow rather loose plate-like scales. Wood light, soft, not strong, brittle, coarse-grained, easily split, dull brown tinged with red; largely used in Washington and Oregon for the interior finish of buildings, doors, sashes, fences, shingles, and in cabinet-making and cooperage. From this tree the Indians of the northwest coast split the planks used in the construction of their lodges, carved the totems which decorate their villages, and hollowed out their great war canoes, and from the fibres of the inner bark made ropes, blankets, and thatch for their cabins.

Distribution. Singly and in small groves on low moist bottom-lands or near the banks of mountain streams, from the sea-level to elevations of 6000° in the interior, from Baranoff Island, Alaska, southward along the coast ranges of British Columbia, western Washington, and Oregon, where it is the most abundant and grows to its largest size, and through the California-coast region to Mendocino County, ranging eastward along many of the interior ranges of British Columbia, northern Washington, Idaho, and Montana to the western slope of the continental divide.

Often cultivated as an ornamental tree in the parks and gardens of western and central Europe where it has grown rapidly and vigorously, and occasionally in the middle and north Atlantic states.

11. CUPRESSUS L. Cypress.

Resinous trees, with bark often separating into long shred-like scales, fragrant durable usually light brown heartwood, pale yellow sapwood, stout erect branches often becoming horizontal in old age, slender 4-angled branchlets, and naked buds. Leaves scale-like, ovate, acute, acuminate, or bluntly pointed at apex, with slender spreading or appressed tips, thickened, rounded, and often glandular on the back, opposite in pairs, becoming brown and woody before falling; on vigorous leading shoots and young plants needle-shaped or linear-lanceolate and spreading. Flowers minute, monœcious, terminal, yellow, the two sexes on separate branchlets; the male oblong, of numerous decussate stamens, with short filaments enlarged into broadly ovate connectives bearing 2—6 globose pendulous anther-cells; female oblong or subglobose, composed of 6—10 thick decussate scales bearing in several rows at the base of the ovuliferous scale numerous erect bottle-shaped ovules. Fruit an erect nearly globose cone maturing in the second year, composed of the much thickened ovule-bearing scales of the flower, abruptly dilated, clavate and flattened at the apex, bearing the remnants of the flower-scales developed into a short central more or less thickened mucro or boss; long-persistent on the branch after the escape of the seeds. Seeds numerous, in several rows, erect, thick, and acutely angled or compressed, with thin lateral wings; seed-coat of 2 layers, the outer thin and membranaceous, the inner thicker and crustaceous; cotyledons 3 or 4, longer than the superior radicle.

Cupressus with ten or twelve species is confined to Pacific North America and Mexico in the New World and to southeastern Europe, southwestern Asia, the Himalayas, and China in the Old World. Of the exotic species Cupressus sempervirens L., of southeastern Europe and southwestern Asia, and especially its pyramidal variety, are often planted for ornament in the south Atlantic and Pacific states.

CONSPECTUS OF THE NORTH AMERICAN SPECIES

1. Cupressus macrocarpa Gord. Monterey Cypress.

Leaves dark green, bluntly pointed, eglandular, and ¼′—½′ long; deciduous at the end of three or four years. Flowers opening late in February or early in March, yellow. Fruit clustered on short stout stems subglobose, slightly puberulous, 1′—1½′ in diameter, composed of 4 or 6 pairs of scales, with broadly ovoid thickened or occasionally on the upper scales subconical bosses, the scales of the upper and lower pairs being smaller than the others and sterile; seeds about 20 under each fertile scale, angled, light chestnut-brown, about ¹/₁₆′ long.

A tree, often 60°—70° high, with a short trunk 2°—3° or exceptionally 5°—6° in diameter, slender erect branches forming a narrow or broad bushy pyramidal head, becoming stout and spreading in old age into a broad flat-topped crown, and stout branchlets covered when the leaves fall at the end of three or four years with thin light or dark reddish brown bark separating into small papery scales. Bark ¾′—1′ thick and irregularly divided into broad flat connected ridges separating freely into narrow elongated thick persistent scales, dark red-brown on young stems and upper branches, becoming at last almost white on old and exposed trunks. Wood heavy, hard and strong, very durable, close-grained.

Distribution. Coast of California south of the Bay of Monterey, occupying an area about two miles long and two hundred yards wide from Cypress Point to the shores of Carmel Bay, with a small grove on Point Lobos, the southern boundary of the bay.

Universally cultivated in the Pacific states from Vancouver Island to Lower California, and often used in hedges and for wind-breaks; occasionally planted in the southeastern states; much planted in western and southern Europe, temperate South America, and in Australia and New Zealand.

2. Cupressus Goveniana Gord.

Leaves acutely pointed, dark green. Flowers: male obscurely 4-angled, with broadly ovate peltate connectives: female with 6—10 ovate pointed scales. Fruit usually sessile, [71] subglobose ¼′—⅞′ in diameter, its scales terminating in small bosses; seeds compressed, black, or dark brown, papillose, about ⅛′ long.

A tree rarely 75° high, with a tall trunk up to 2°10′ in diameter, often not more than 25° high, more often a shrub with numerous stems 1°—15° tall, ascending branches, and comparatively stout bright reddish brown branchlets, becoming purple and ultimately dark reddish brown; often beginning to produce fertile cones when only 1° or 2° tall. Bark bright reddish brown, about ¼′ thick, and divided by shallow fissures into flat ridges separating on the surface into long thread-like scales. Wood soft, very coarse-grained, pale reddish brown.

Distribution. California: pine barrens on the western slope of Point Pinos Ridge two miles west of Monterey, and on alkaline soil in a narrow belt beginning about three quarters of a mile from the shore of Mendocino County and extending inland for three or four miles from Ten Mile Run on the north to the Navarro River on the south; arborescent and also of its smallest size only in this northern station.

3. Cupressus Sargentii Jeps. Sargent’s Cypress.

Leaves obscurely glandular or without glands, dark green, pungently aromatic, ¹/₁₆′—⅛′ long, turning bright red-brown in drying and falling at the end of three or four years; on young plants ⅛′—¼′ long. Flowers: male with thin slightly erose connectives: female of 6 or 8 acute slightly spreading scales. Fruit often in crowded clusters, short-stalked, subglobose, ½′—1′ in diameter, reddish brown or purple, lustrous, puberulous, its 6 or 8 scales with broadly ovoid generally rounded and flattened and rarely short-obconic bosses; seeds brown, lustrous, often glaucous, with an acute margin, ⅙′ long, about 20 under each fertile scale.

A tree, shrub, or small bushy tree rarely more than 15° or 16° high, with a short trunk 2° in diameter, slender erect or spreading branches forming a handsome open head, and thin branchlets covered with close smooth bark, at first orange-colored, becoming bright reddish brown, and ultimately purple or dark brown. Bark ¼′—½′ thick, dark grayish brown, irregularly divided into narrow ridges covered with thin persistent oblong scales. Wood light, soft, not strong, light brown, with thick nearly white sapwood.

Distribution. California: dry mountain slopes usually between altitudes of 1300° and 2300° in few widely isolated stations, Red Mountain, Mendocino County, to Mt. Tamalpais, Marin County; Cedar Mountain, Alameda County; Santa Cruz Mountains, Santa Cruz County; Santa Lucia Mountains, Monterey County; often covering great areas on the hills of Marin County with dense thickets only a few feet high.

Occasionally cultivated as C. Goveniana in western and southern Europe as an ornamental tree.

4. Cupressus Macnabiana A. Murr. Cypress.

Leaves acute or rounded at apex, rounded and conspicuously glandular on the back, deep green, often slightly glaucous, usually not more than ¹/₁₆′ long. Flowers in March and April, male nearly cylindric, obtuse, with broadly ovate rounded connectives: female subglobose, with broadly ovate scales short-pointed and rounded at apex. Fruit oblong, subsessile or raised on a slender stalk, ½′—1′ long, dark reddish brown more or less covered with a glaucous bloom, slightly puberulous, especially along the margins of the 6 or rarely 8 scales, their prominent bosses thin and recurved on the lower scales, and much thickened, conical, and more or less incurved on the upper scales; seeds dark chestnut-brown, usually rather less than ¹/₁₆′ long, with narrow wings.

A tree in Oregon occasionally 80° high with a tall trunk sometimes 3½° in diameter, southward rarely more than 30° high, with a short trunk 12′—15′ in diameter, slender branches covered with close smooth compact bark, bright purple after the falling of the leaves, soon becoming dark brown; more often a shrub with numerous stems 6°—12° tall forming a broad open irregular head. Bark thin, dark reddish brown, broken into brown flat ridges, and separating on the surface into elongated thin slightly attached long-persistent scales. Wood light, soft, very close-grained.

Distribution. Rare and local, usually in small groves; dry ridges of Mount Steve and adjacent mountains up to altitudes of 5300°, Josephine County, southwestern Oregon; California; on lava beds, southeastern Siskiyou and southwestern Mono Counties (C. Bakeri); dry hills and low slopes, Mt. Ætna, in central Napa County; through Lake County to Red Mountain on the east side of Ukiah Valley, Mendocino County; in Trinity County between Shasta and Whiskeytown; and on the Sierra Nevada (Red Hill, Piute Mountains near Bodfish) Kern County, at an altitude of 5000° (C. nevadensis).

5. Cupressus guadaloupensis S. Wats. Tecate Cypress.

Leaves acute, rounded and minutely glandular-pitted or eglandular on the back, light blue-green, about ¹/₁₆′ long. Fruit on stout stems ¼′—⅓′ in length, subglobose to short-oblong, ¾′—1¼′ in diameter, puberulous especially along the margins of the six or eight scales, with prominent flattened or conic acute often incurved bosses; seeds about 70 under each scale, short-oblong, nearly square, light chestnut-brown up to ¼′ in length, with a narrow wing.

A tree in California sometimes 20°—25° in height, with a short slender or on exposed mountain slopes a trunk occasionally 2° or 3° in diameter, few short spreading or ascending branches forming an open head, and light red-brown lustrous branchlets becoming purplish. Bark smooth, lustrous, without resin or fibres, mahogany brown, the thin scales in falling leaving pale marks.

Distribution. San Diego County, California, rare and local; valley of the San Luis Rey River between Valley Centre and Pala; at altitudes between 1100° and 4000° in the gulches and on the summit of Mt. Tecate on the border between the United States and Lower California; on a mountain below Descanso and Pine Valley; in Cedar Cañon between El-nido and Dulzura; in Lower California on San Pedro Mártir Mountain and Guadaloupe Island. The insular form is a larger tree often with larger gland-pits on the leaves, and now often cultivated in California, western Europe, and in other countries with temperate climates.

6. Cupressus arizonica Greene. Cypress.

Leaves obtusely pointed, rounded, eglandular or rarely glandular-pitted on the back, pale green, ¹/₁₆′ long, dying and turning red-brown in their second season, generally falling four years later. Flowers: male oblong, obtuse, their 6 or 8 stamens with broadly ovate acute yellow connectives slightly erose on the margins: female not seen. Fruit on stout pedicels ¼′—½′ in length, subglobose, rather longer than broad, wrinkled, dark red-brown and covered with a glaucous bloom, the six or eight scales with stout flattened incurved prominent bosses; seeds oblong to nearly triangular, dark red-brown, ¹/₁₆′—⅛′ long with a thin narrow wing.

A conical tree 40°—70° high with a trunk 2°—4° in diameter, and stout spreading branches covered with bark separating into thin plates, leaving a smooth red surface, and branchlets [74] dark gray after the leaves fall. Bark on young trunks separating into large irregular curling thin scales, on old trees becoming dark red-brown and fibrous.

Distribution. Mountains above Clifton, Greenlee County, eastern Arizona; on the San Francisco Mountains, Socorro County, and San Luis Mountains, Grant County, western New Mexico; and in Chihuahua. Passing into

Cupressus arizonica var. bonita Lemm.

Differing from the type in the prominent oblong or circular glandular depressions on the backs of the leaves.

A tree 30°—70° high, with a trunk 18′—24′ or rarely 5° in diameter, erect branches forming a rather compact conical head. Bark of the trunk and large branches thin, smooth, dark reddish brown, separating into small curled scale-like plates, becoming on old trees dark gray and fibrous. Wood heavy, hard, pale straw color with lighter-colored sapwood, [75] durable in contact with the ground, somewhat used for fence-posts, corral-piles, mine-timbers and in log cabins.

Distribution. Gravelly slopes and moist gulches often in groups of considerable size at altitudes between 4000° and 7000°, Arizona; near Camp Verde, Tonto Basin; Natural Bridge, Payson, etc.; on the Chiracahua Mountains (J. W. Toumey, July, 1894); on the Santa Rita and Santa Catalina Mountains, and in Oak Creek Cañon twenty miles south of Flagstaff (P. Lowell, June, 1911).

12. CHAMÆCYPARIS.

Tall resinous pyramidal trees, with thin scaly or deeply furrowed bark, nodding leading shoots, spreading branches, flattened, often deciduous or ultimately terete branchlets 2-ranked in one horizontal plane, pale fragrant durable heartwood, thin nearly white sapwood, and naked buds. Leaves scale-like, ovate, acuminate, with slender spreading or appressed tips, opposite in pairs, becoming brown and woody before falling, on vigorous sterile branches and young plants needle-shaped or linear-lanceolate and spreading. Flowers minute, monœcious, terminal, the two sexes on separate branchlets; the male oblong, of numerous decussate stamens, with short filaments enlarged into ovate connectives decreasing in size from below upward and bearing usually 2 pendulous globose anther-cells; the female subglobose, composed of usually 6 decussate peltate scales bearing at the base of the ovuliferous scales 2—5 erect bottle-shaped ovules. Fruit an erect globose cone maturing at the end of the first season, surrounded at the base by the sterile lower scales of the flowers, and formed by the enlargement of the ovule-bearing scales, abruptly dilated, club-shaped and flattened at the apex, bearing the remnants of the flower-scales as short prominent points or knobs; persistent on the branches after the escape of the seeds. Seeds 1—5, erect on the slender stalk-like base of the scale, subcylindric and slightly compressed; seed-coat of 2 layers, the outer thin and membranaceous, the inner thicker and crustaceous, produced into broad lateral wings; cotyledons 2, longer than the superior radicle.

Chamæcyparis is confined to the Atlantic and Pacific coast regions of North America, and to Japan and Formosa. Six species are distinguished. Of exotic species the Japanese Retinosporas, Chamæcyparis obtusa Endl., and Chamæcyparis pisifera Endl., with their numerous abnormal forms are familiar garden plants in all temperate regions.

Chamæcyparis is from χαμαί, on the ground, and κυπάρισσος, cypress.

CONSPECTUS OF THE NORTH AMERICAN SPECIES.

1. Chamæcyparis thyoides B. S. P. White Cedar.

Leaves closely appressed, or spreading at the apex especially on vigorous leading shoots, keeled and glandular or conspicuously glandular-punctate on the back, dark dull blue-green or pale below, at the north becoming russet-brown during the winter, ¹/₁₆′—⅛′ long, dying during the second season and then persistent for many years. Flowers: male composed of 5 or 6 pairs of stamens, with ovate connectives rounded at apex, dark brown below the middle, nearly black toward the apex: female subglobose, with ovate acute [76] spreading pale liver-colored scales and black ovules. Fruit ¼′ in diameter, sessile on a short leafy branch, light green, covered with a glaucous bloom when fully grown, later bluish purple and very glaucous, finally becoming dark red-brown, its scales terminating in ovate acute, often reflexed bosses; seeds 1 or 2 under each fertile scale, ovoid, acute, full and rounded at the base, slightly compressed, gray-brown, about ⅛′ long, with wings as broad as the body of the seed and dark red-brown.

A tree, 70°—80° high, with a tall trunk usually about 2 and occasionally 3°—4° in diameter, or northward much smaller, slender horizontal branches forming a narrow spire-like head, and 2-ranked compressed branchlets disposed in an open fan-shaped more or less deciduous spray, the persistent branchlets gradually becoming terete, light green tinged with red, light reddish brown during their first winter, and then dark brown, their thin close bark separating slightly at the end of three or four years into small papery scales. Bark ¾′—1′ thick, light reddish brown, and divided irregularly into narrow flat connected ridges often spirally twisted round the stem, separating on the surface into elongated loose or closely appressed plate-like scales. Wood light, soft, not strong, close-grained, slightly fragrant, light brown tinged with red; largely used in boat-building and cooperage, for wooden ware, shingles, the interior finish of houses, fence-posts, and railway-ties.

Distribution. Cold swamps usually immersed during several months of the year, often forming dense pure forests; near Concord, New Hampshire, southern Maine, southward only near the coast to northern Florida, and westward to southwestern Mississippi; most abundant south of Massachusetts Bay; comparatively rare east of Boston and west of Mobile Bay.

Occasionally planted as an ornamental tree in the eastern states and in the countries of temperate Europe.

2. Chamæcyparis nootkatensis Sudw. Yellow Cypress. Sitka Cypress.

Leaves rounded, eglandular or glandular-pitted on the back, dark blue-green, closely appressed, about ⅛′ long, on vigorous leading branchlets somewhat spreading and often ¼′ long, with more elongated and sharper points, beginning to die at the end of their second year and usually falling during the third season. Flowers: male on lateral branchlets of the previous year, composed of 4 or 5 pairs of stamens, with ovate rounded slightly erose light yellow connectives: female clustered near the ends of upper branchlets, dark liver color, the fertile scales each bearing 2—4 ovules. Fruit ripening in September and October. [77] nearly ½′ in diameter, dark red-brown, with usually 4 or 6 scales tipped with prominent erect pointed bosses and frequently covered with conspicuous resin-glands; seeds 2—4 under each scale, ovoid, acute, slightly flattened, about ¼′ long, dark red-brown, with thin light red-brown wings often nearly twice as wide as the body of the seed.

A tree, frequently 120° high, with a tall trunk 5°—6° in diameter, horizontal branches forming a narrow pyramidal head, stout distichous somewhat flattened or terete light yellow branchlets often tinged with red at first, dark or often bright red-brown during their third season, ultimately paler and covered with close thin smooth bark. Bark ½′—¾′ thick, light gray tinged with brown, irregularly fissured, and separated on the surface into large thin loose scales. Wood hard, rather brittle, very close-grained, exceedingly durable, bright clear yellow, with very thin nearly white sapwood; fragrant with an agreeable resinous odor; used in boat and shipbuilding, the interior finish of houses, and the manufacture of furniture.

Distribution. Islands of Prince William Sound, Alaska, and southward over the coast mountains of Alaska and British Columbia, and along the Cascade Mountains of Washington and Oregon to the northeastern slopes of Mt. Jefferson, extending eastward to the headwaters of the Yakima River on the eastern slope of the range; on Whiskey Peak of the Siskiyou Mountains in the southeastern corner of Josephine County, Oregon and about two miles from the California line; most abundant and of its largest size near the coast of Alaska and northern British Columbia, ranging from the sea-level up to altitudes of 3000°; at high elevations on the Cascade Mountains sometimes a low shrub.

Occasionally cultivated, with its several abnormal forms, as an ornamental tree in the middle Atlantic states and in California, and commonly in the countries of western and central Europe.

3. Chamæcyparis Lawsoniana Parl. Port Orford Cedar. Lawson Cypress.

Leaves bright green or pale below, conspicuously glandular on the back, usually not more than ¹/₁₆′ long on lateral branchlets, on leading shoots often spreading at the apex, ⅛′ to nearly ¼′ long, usually dying, turning bright red-brown and falling during their third year. Flowers: male with bright red connectives bearing usually 2 pollen-sacs: female with dark ovate acute spreading scales, each bearing 2—4 ovules. Fruit clustered on the upper lateral branchlets and produced in great profusion, ripening in September and October, about ⅓′ in diameter, green and glaucous when full grown, red-brown and often covered with a bloom at maturity, its scales with thin broadly ovate acute reflexed bosses; seeds 2—4 under each fertile scale, ovoid, acute, slightly compressed, ⅛′ long, light chestnut-brown, with broad thin wings.

A tree, often 200° high, with a tall trunk frequently 12° in diameter above its abruptly enlarged base, a spire-like head of small horizontal or pendulous branches clothed with [78] remote flat spray frequently 6′—8′ long. Bark often 10′ thick at the base of old trees and 3′—4′ thick on smaller stems, dark reddish brown, with 2 distinct layers, the inner ⅛′—¼′ thick, darker, more compact, and firmer than the outer, divided into great broad-based rounded ridges separated on the surface into small thick closely appressed scales. Wood light, hard, strong, very close-grained, abounding in fragrant resin, durable, easily worked, light yellow, or almost white, with hardly distinguishable sapwood; largely manufactured into lumber used for the interior finish and flooring of buildings, railway-ties, fence-posts, and boat and shipbuilding, and on the Pacific coast almost exclusively for matches. The resin is a powerful diuretic.

Distribution. Usually scattered in small groves from the shores of Coos Bay, southwestern Oregon, south to the mouth of the Klamath River, California, ranging inland usually for about thirty miles; near Waldorf, in Josephine County, Oregon, on the slopes of the Siskiyou Mountains, and on the southern flanks of Mt. Shasta, California; most abundant north of Rogue River on the Oregon coast and attaining its largest size on the western slopes of the Coast Range foothills, forming between Point Gregory and the mouth of the Coquille River a nearly continuous forest belt twenty miles long.

Often cultivated with the innumerable forms originated in nurseries, in the middle Atlantic states and California, in all the temperate countries of Europe, and in New Zealand.

13. JUNIPERUS L. Juniper.

Pungent aromatic trees or shrubs, with usually thin shreddy bark, soft close-grained durable wood, slender branches, and scaly or naked buds. Leaves sessile, in whorls of 3, persistent for many years, convex on the lower side, concave and stomatiferous above, linear-subulate, sharp-pointed, without glands (Oxycedrus); or scale-like, ovate, opposite in pairs or ternate, closely imbricated, appressed and adnate to the branch, glandular or eglandular on the back, becoming brown and woody on the branch, but on young plants and vigorous shoots often free and awl-shaped (Sabina). Flowers minute, diœcious, axillary or terminal on short axillary branches from buds formed the previous autumn on branches of the year; the male solitary, oblong-ovoid, with numerous stamens decussate or in 3’s, their filaments enlarged into ovate or peltate yellow scale-like connectives bearing near the base 2—6 globose pollen-sacs; the female ovoid, surrounded at the base by many minute scale-like bracts persistent and unchanged under the fruit, composed of 2—6 opposite or ternate pointed scales alternate with or bearing on their inner face at the base on a minute ovuliferous scale 1 or 2 ovules. Fruit a berry-like succulent fleshy blue, blue-black, [79] or red strobile formed by the coalition of the flower-scales, inclosed in a membranaceous skin covered with a glaucous bloom, ripening during the first, second, or rarely during the third season, smooth or marked by the ends of the flower-scales, or by the pointed tips of the ovules, closed, or open at the top and exposing the apex of the seeds. Seeds 1—12, ovoid, acute or obtuse, terete or variously angled, often longitudinally grooved by depressions caused by the pressure of resin-cells in the flesh of the fruit, smooth or roughened and tuberculate, chestnut-brown, marked below by the large conspicuous usually 2-lobed hilum; seed-coat of 2 layers, the outer thick and bony, the inner thin, membranaceous or crustaceous; cotyledons 2, or 4—6, about as long as the superior radicle.

Juniperus is widely scattered over the northern hemisphere from the Arctic Circle to the highlands of Mexico, Lower California, and the West Indies in the New World, and to the Azores and Canary Islands, northern Africa, Abyssinia, the mountains of east tropical Africa, Sikkim, central China, Formosa, Japan and the Bonin Islands in the Old World. About thirty-five species are now distinguished. Of the exotic species cultivated in the United States the most common are European forms of Juniperus communis L. with fastigiate branches, and dwarf forms of the European Juniperus Sabina L., and of Juniperus chinensis L.

CONSPECTUS OF THE NORTH AMERICAN SPECIES.

1. Juniperus communis L. Juniper.

Leaves spreading nearly at right angles to the branchlets, ⅓′—½′ long, about ¹/₃₂′ wide, turning during winter a deep rich bronze color on the lower surface, persistent for many years. Flowers: male composed of 5 or 6 whorls each of 3 stamens, with broadly ovate acute and short-pointed connectives, bearing at the very base 3 or 4 globose anther-cells; female surrounded by 5 or 6 whorls of ternate leaf-like scales, composed of 3 slightly spreading ovules abruptly enlarged and open at the apex, with 3 minute obtuse fleshy scales below and alternate with them. Fruit maturing in the third season, subglobose or short-oblong, about ¼′ in diameter, with soft mealy resinous sweet flesh and 1—3 seeds; often persistent on the branches one or two years after ripening; seeds ovoid, acute, irregularly angled or flattened, deeply penetrated by numerous prominent thin-walled resin-glands, about ⅛′ long, the outer coat thick and bony, the inner membranaceous.

In America only occasionally tree-like and 10°—20° tall, with a short eccentric irregularly lobed trunk rarely a foot in diameter, erect branches forming an irregular open head, slender branchlets, smooth, lustrous, and conspicuously 3-angled between the short nodes during their first and second years, light yellow tinged with red, gradually growing darker, their dark red-brown bark separating in the third season into small thin scales, and ovoid acute buds about ⅛′ long and loosely covered with scale-like leaves; more often a shrub, with many short slender stems prostrate at the base and turning upward and forming a broad mass sometimes 20° across and 3° or 4° high (var. depressa Pursh.); at high elevations and in the extreme north prostrate, with long decumbent stems and shorter and more crowded leaves (var. montana Ait.) passing into the var. Jackii Rehdr. with long trailing branches and broader incurved leaves. Bark about ¹/₁₆′ thick, dark reddish brown, separating [81] irregularly into many loose papery persistent scales. Wood hard, close-grained, very durable in contact with the soil, light brown, with pale sapwood. In northern Europe the sweet aromatic fruit of this tree is used in large quantities to impart its peculiar flavor to gin; occasionally employed in medicine.

Distribution. Occasionally arborescent in New England, eastern Pennsylvania, and on the high mountains of North Carolina; the var. depressa, common in poor rocky soil, Newfoundland to southern New England, and to the shores of the Great Lakes and northwestward; the var. montana from the coast of Greenland to northern New England, on the high Appalachian Mountains, North Carolina, and to northern Nebraska, along the Rocky Mountains from Alberta to western Texas, and on the Pacific coast from Alaska, southward along mountain ranges to the high Sierras of central California, extending eastward to the mountains of eastern Washington and Oregon, and on the high peaks of northern Arizona up to altitudes of 10,000°—11,500° (P. Lowell); the var. Jackii on the coast mountains from northern California to Vancouver Island; in the Old World widely distributed in many forms through all the northern hemisphere from arctic Asia and Europe to Japan, the Himalayas and the mountains of the Mediterranean Basin.

Often planted, especially in several of its pyramidal and dwarf forms, in the eastern United States and in the countries of western, central, and northern Europe.

2. Juniperus Pinchotii Sudw.

Leaves ternate, obtusely pointed, rounded and glandular-pitted on the back, ¹/₁₆′ long, dark yellow-green, turning light red-brown before falling; on vigorous shoots and seedling plants linear-lanceolate, thin, acuminate, eglandular, ¼′—⅓′ in length. Fruit ripening in one season, subglobose, bright red, ¼′ in diameter, with a thin skin and thick dry mealy resinous flesh and 1 seed; seed ovoid, bluntly pointed, deeply grooved, irregularly marked by the usually two-lobed hilum, ⅙′—¼′ long and 2 cotyledons.

A tree rarely 20 feet high, with a trunk 1 foot in diameter, stout wide-spreading branches forming an open irregular head and thick branchlets covered with dark gray-brown scaly bark, their ultimate divisions about ¹/₁₂′ in diameter; more often a shrub with several stems 1° to 12° tall. Bark thin, light brown, separating into long narrow persistent scales.

Distribution. Dry rocky slopes and the rocky sides of cañons, Panhandle of western [82] Texas (Armstrong, Potter and Hartley Counties), and in Hardaman, Garza, Tom Green, Kemble, Valverde and Menard Counties; on Comanche Peak near Granbury, Hood County, Texas; in central and on the mountains of southern Arizona.

3. Juniperus californica Carr. Desert White Cedar. Sweet-berried Cedar.

Leaves usually in 3’s, closely appressed, thickened, slightly keeled and conspicuously glandular-pitted on the back, pointed at apex, cartilaginously fringed on the margins, light yellow-green, about ⅛′ long, dying and turning brown on the branch at the end of two or three years; on vigorous shoots linear-lanceolate, rigid, sharp-pointed, ¼′—½′ long, whitish on the upper surface. Flowers from January to March; male of 18—20 stamens, disposed in 3’s, with rhomboidal short-pointed connectives; scales of the female flower usually 6, ovate, acute, spreading, obliterated or minute on the fruit. Fruit short-oblong or ovoid, ½′—¾′ long, reddish brown, with a membranaceous loose skin covered with a thick glaucous bloom, thick fibrous dry sweet flesh, and 1 or 2 seeds; seeds ovoid, obtusely pointed, irregularly lobed and angled, and 4—6 cotyledons.

A conical tree, occasionally 40° high, with a straight, large-lobed unsymmetrical trunk 1°—2° in diameter; more often shrubby, with many stout irregular usually contorted stems forming a broad open head. Bark thin and divided into long loose plate-like scales ashy gray on the outer surface and persistent for many years. Wood soft, close-grained, durable in contact with the soil, light brown slightly tinged with red, with thin nearly white sapwood; used for fencing and fuel. The fruit is eaten by Indians fresh or ground into flour.

Distribution. Dry mountain slopes and hills at altitudes between 400° and 4000°, from Moraga Pass and Mt. Diabolo, Contra Costa County, California, southward on the coast ranges, spreading inland to their union with the Sierra Nevada, and northward at low altitudes along the western slopes of the Sierras to Kern and Mariposa Counties; on the desert slopes of the Tehachapi Mountains, the northern foothills of the San Bernardino Mountains, on the western slopes of the San Jacinto and Cayamaca Ranges, and southward in Lower California to Agua Dulce; arborescent and probably of its largest size on the Mohave Desert.

4. Juniperus utahensis Lemm. Juniper.

Leaves opposite or in 3’s, rounded, usually glandular, acute or often acuminate, light yellow-green, rather less than ⅛′ long, persistent for many years. Flowers: male with 18—24 opposite or ternate stamens, their connectives rhomboidal; scales of the female flower acute, spreading, often in pairs. Fruit ripening during the autumn of the second season, subglobose or short-oblong, marked by the more or less prominent tips of the flower-scales, reddish brown, with a thick firm skin covered with a glaucous bloom and closely investing [83] the thin dry sweet flesh, ¼′—⅓′ long, with 1 or rarely 2 seeds; seeds ovoid, acute, obtusely angled, marked to the middle by the hilum, with a hard bony shell, and 4—6 cotyledons.

A bushy tree, rarely exceeding 20° in height, with a short usually eccentric trunk sometimes 2° in diameter, generally divided near the ground by irregular deep fissures into broad rounded ridges, many erect contorted branches forming a broad open head, slender light yellow-green branchlets covered after the falling of the leaves with thin light red-brown scaly bark; more often with numerous stems spreading from the ground and frequently not more than 8°—10° high. Bark about ¼′ thick, ashy gray or sometimes nearly white, and broken into long thin persistent scales. Wood light brown, slightly fragrant, with thick nearly white sapwood; largely used locally for fuel and fencing. The fruit is eaten by Indians fresh, or ground and baked into cakes.

Distribution. Southwestern Wyoming (J. Knightii A. Nels.), southwestern Idaho (Pocatello, Bannock County), western Colorado, eastern Utah, and western New Mexico to northern Arizona and southeastern California at altitudes from 5000° to 8000°; the most abundant and generally distributed tree of the Great Basin, forming in the valleys open forests of stunted trees and shrubs, and on arid slopes more numerous and of larger size in dense nearly pure forests.

A variety (var. megalancocarpa Sarg.) occurs in eastern New Mexico and northern Arizona, with fruit sometimes ¾′ in diameter. A tree often 40° high with a single erect stem sometimes 3° in diameter.

5. Juniperus flaccida Schlecht. Juniper.

Leaves opposite, acuminate and long-pointed, spreading at the apex, glandular or eglandular on the back, light yellow-green, about ⅛′ long, turning cinnamon-red and dying on the branch; on vigorous young shoots ovate-lanceolate, sometimes ½′ long, with elongated rigid callous tips. Flowers: male slender, composed of 16—20 stamens, with ovate pointed connectives prominently keeled on the back; female with acute or acuminate spreading scales. Fruit subglobose, dull red-brown, more or less covered with a glaucous bloom, ⅓′—½′ in diameter, with a close firm skin and thick resinous flesh; seeds 4—12, pointed at apex, slightly ridged, often abortive and distorted, ⅛′—¼′ long, with 2 cotyledons.

A tree, occasionally 30° high, with gracefully spreading branches and long slender drooping branchlets, covered after the leaves fall with thin bright cinnamon-brown bark separating [84] into thin loose papery scales; often a shrub. Bark about ½′ thick, reddish brown, separating into long narrow loosely attached scales.

Distribution. In the United States only on the slopes of the Chisos Mountains, in Brewster County, southern Texas; common in northeastern Mexico, growing at elevations of 6000°—8000° on the hills east of the Mexican table-lands.

6. Juniperus pachyphlæa Torr. Juniper. Checkered-bark Juniper.

Leaves appressed, acute and apiculate at apex, thickened, obscurely keeled and glandular on the back, bluish green, rather less than ⅛′ long; on vigorous shoots and young branchlets linear-lanceolate, tipped with slender elongated points, and pale blue-green like the young branchlets. Flowers opening in February and March: the male stout, ⅛′ long, with 10 or 12 stamens, their connectives broadly ovate, obscurely keeled on the back, short-pointed: scales of the female flower, ovate, acuminate, and spreading. Fruit ripening in the autumn of its second season, subglobose to short-oblong, irregularly tuberculate, ⅓′—½′ in diameter, usually marked with the short tips of the flower-scales, occasionally opening and discharging the seeds at the apex, dark red-brown, more or less covered with [85] a glaucous bloom, especially during the first season and then occasionally bluish in color, with a thin skin closely investing the thick dry mealy flesh, and usually 4 seeds; seeds acute or obtusely pointed, conspicuously ridged and gibbous on the back, with a thick shell and 2 cotyledons.

A tree, often 50°—60° high, with a short trunk 3°—5° in diameter, long stout spreading branches forming a broad-based pyramidal or ultimately a compact round-topped head, and slender branchlets covered after the disappearance of the leaves with thin light red-brown usually smooth close bark occasionally broken into large thin scales. Bark ¾′—4′ thick, on young stems reddish brown becoming on old trunks whitish, deeply fissured and divided into nearly square plates 1′—2′ long, and separating on the surface into small thin closely appressed scales. Wood light, soft, not strong, brittle, close-grained, clear light red often streaked with yellow, with thin nearly white sapwood; often producing vigorous shoots from the base of the trunk or from the stumps of felled trees.

Distribution. Dry arid mountain slopes usually at elevations of 4000°—6000° above the sea, from the Eagle and Limpio mountains in southwestern Texas, westward along the desert ranges of New Mexico and Arizona, extending northward to the lower slopes of many of the high mountains of northern Arizona, and southward into Mexico.

7. Juniperus occidentalis Hook. Juniper.

Leaves opposite or ternate, closely appressed, acute or acuminate, rounded and conspicuously glandular on the back, denticulately fringed, gray-green, about ⅛′ long. Flowers: male stout, obtuse, with 12—18 stamens, their connectives broadly ovoid, rounded, acute or apiculate and scarious or slightly ciliate on the margins: scales of the female flower ovate, acute, spreading, mostly obliterated from the fruit. Fruit subglobose or short-oblong, ¼′—⅓′ in diameter, with a thick firm blue-black skin coated with a glaucous bloom, thin dry flesh filled with large resin-glands, and 2 or 3 seeds; seeds ovoid, acute, rounded and deeply grooved or pitted on the back, flattened on the inner surface, about ⅛′ long, with a thick bony shell, a thin brown inner seed-coat, and 2 cotyledons.

A tree, occasionally 60° high, with a tall straight trunk 2°—3° in diameter, more often not more than 20° in height, with a short trunk sometimes 10° in diameter, enormous branches, spreading at nearly right angles and forming a broad low head, and stout branchlets covered after the leaves fall with thin bright red-brown bark broken into loose papery scales; frequently when growing on dry rocky slopes and toward the northern limits of its range a shrub, with many short erect or semiprostrate stems. Bark about [86] ½′ thick, bright cinnamon-red, divided by broad shallow fissures into wide flat irregularly connected ridges separating on the surface into thin lustrous scales. Wood light, soft, very close-grained, exceedingly durable, light red or brown, with thick nearly white sapwood; used for fencing and fuel. The fruit is gathered and eaten by the California Indians.

Distribution. Mountain slopes and high prairies of western Idaho and of eastern Washington to the eastern slopes of the Cascade Mountains; eastern and southern Oregon up to altitudes of 4500°; along the summits and upper slopes of the Sierra Nevada of California, and southward to the San Bernardino Mountains, here abundant in Bear and Holcomb valleys; attaining its greatest trunk diameter on the wind-swept peaks of the California sierras, usually at altitudes between 6000° and 10,000° above the sea.

8. Juniperus monosperma Sarg. Juniper.

Leaves opposite or ternate, often slightly spreading at apex, acute or occasionally acuminate, much thickened and rounded on the back, usually glandular, denticulately fringed, gray-green, rather less than ⅛′ long, turning bright red-brown before falling; on vigorous shoots and young plants ovate, acute, tipped with long rigid points, thin, conspicuously glandular on the back, often ½′ long. Flowers: male with 8—10 stamens, their broadly ovate, rounded or pointed connectives slightly erose on the margins: female with spreading pointed scales. Fruit subglobose or short-oblong, ⅛′—¼′ long, dark blue or perhaps occasionally light chestnut-brown with a thick firm skin covered with a thin glaucous bloom, thin flesh, and 1 or rarely 2 seeds; seeds often protruding from the top of the fruit, ovoid, often 4-angled, somewhat obtuse at apex, with a small hilum, and 2 cotyledons.

A tree, occasionally 40°—50° high, with a stout much-lobed and buttressed trunk sometimes 3° in diameter, short stout branches forming an open very irregular head, and slender branchlets covered after the falling of the leaves with light red-brown bark spreading freely into thin loose scales; more often a much branched shrub sometimes only a few feet high. Bark ashy gray, divided into irregularly connected ridges, separating into long narrow persistent shreddy scales. Wood heavy, slightly fragrant, light reddish brown, with nearly white sapwood and eccentric layers of annual growth; largely used for fencing and fuel. The fruit is ground into flour and baked by the Indians, who use the thin strips of fibrous bark in making saddles, breechcloths, and sleeping-mats.

Distribution. Along the eastern base of the Rocky Mountains from the valley of the Platte River, Wyoming (near Alcova, Natrona County) and the divide between the [87] Platte and Arkansas rivers in Colorado; western Oklahoma (near Kenton, Cimarron County, common) and western Texas; on the Colorado plateau, northern Arizona; over the mountain ranges of southwestern Wyoming, Nevada, southern New Mexico and Arizona, and southward into northern Mexico; often covering, with the Nut Pine, in southern Colorado and Utah, and in northern and central New Mexico and Arizona, great areas of rolling hills 6000°—7000° above the sea-level; reaching its largest size in northern Arizona.

9. Juniperus mexicana Spreng. Cedar. Rock Cedar.

Leaves usually opposite or ternate, thickened and keeled on the back, obtuse or acute at apex, mostly without glands, denticulately fringed, rather more than ¹/₁₆′ long, dark blue-green, on vigorous young shoots and seedling plants lanceolate, long-pointed, rigid, ¼′—½′ long. Flowers: male with 12—18 stamens, their connectives ovoid, obtuse, or slightly cuspidate: scales of the female flower ovate, acute, and spreading, very conspicuous when the fruit is half grown, becoming obliterated at its maturity. Fruit short-oblong to subglobose, ¼′—½′ in diameter, dark blue, with a thin skin covered with a glaucous bloom, sweet resinous flesh, and 1 or 2 seeds; seeds ovoid, acute, slightly ridged, rarely tuberculate, dark chestnut-brown, with a small hilum, a thin outer seed-coat, a membranaceous dark brown inner coat, and 2 cotyledons.

A tree, occasionally 100° but generally not more than 20°—30° high, with a short or elongated slightly lobed trunk seldom exceeding a foot in diameter, small spreading branches forming a wide round-topped open and irregular or a narrow pyramidal head, slender sharply 4-angled branchlets becoming terete after the falling of the leaves, light reddish brown or ashy gray, with smooth or slightly scaly bark; often a shrub, with numerous spreading stems. Bark on old trees ¼′—½′ thick, brown tinged with red, and divided into long narrow slightly attached scales persistent for many years and clothing the trunk with a loose thatch-like covering. Wood light, hard, not strong, slightly fragrant, brown streaked with red; largely used for fencing, fuel, telegraph-poles, and railway-ties.

Distribution. From Brazos County over the low limestone hills of western and southern Texas, and southward into Mexico; forming great thickets and growing to its largest size on the San Bernardo River; much smaller farther westward, and usually shrubby at the limits of vegetation on the high mountains of central Mexico.

10. Juniperus virginiana L. Red Cedar. Savin.

Leaves usually opposite, acute or acuminate or occasionally obtuse, rounded and glandular or eglandular on the back, about ¹/₁₆′ long, dark blue-green or glaucous (var. glauca Carr.), at the north turning russet or yellow-brown during the winter, beginning in their third season to grow hard and woody, and remaining two or three years longer on the branches, on young plants and vigorous branchlets linear-lanceolate, long-pointed, light yellow-green, without glands, ½′—¾′ long. Flowers: diœcious or very rarely monœcious: male with 10 or 12 stamens, their connectives rounded and entire, with 4 or occasionally 5 or 6 pollen-sacs; scales of the female flower violet color, acute and spreading, becoming obliterated from the fruit. Fruit subglobose, ¼′—⅓′ in diameter, pale green when fully grown, dark blue and covered with a glaucous bloom at maturity, with a firm skin, thin sweetish resinous flesh, and 1 or 2 or rarely 3 or 4 seeds; seeds acute and occasionally apiculate at apex, ⅙′—⅛′ long, with a comparatively small 2-lobed hilum, and 2 cotyledons.

A tree, occasionally 100° high, with a trunk 3°—4° in diameter, often lobed and eccentric, and frequently buttressed toward the base, generally not more than 40°—50° tall, with short slender branches horizontal on the lower part of the tree, erect above, forming a narrow compact pyramidal head, in old age usually becoming broad and round-topped or irregular, and slender branchlets terete after the disappearance of the leaves and covered with close dark brown bark tinged with red or gray; on exposed cliffs on the coast of Maine, sometimes only a few inches high with long branches forming broad dense mats. Bark ⅛′—¼′ thick, light brown tinged with red, and separated into long narrow scales fringed on the margins, and persistent for many years. Wood light, close-grained, brittle, not strong, dull red, with thin nearly white sapwood, very fragrant, easily worked; largely used for posts, the sills of buildings, the interior finish of houses, the lining of closets and chests for the preservation of woolens against the attacks of moths, and largely for pails and other small articles of wooden ware. A decoction of the fruit and leaves is used in medicine, and oil of red cedar distilled from the leaves and wood as a perfume.

Distribution. Dry gravelly slopes and rocky ridges, often immediately on the seacoast, from southern Nova Scotia and New Brunswick to the coast of Georgia, the interior of southern Alabama and Mississippi, and westward to the valley of the lower Ottawa River, southern Michigan, eastern North and South Dakota, Nebraska and Kansas, and eastern Texas, not ascending the mountains of New England and New York nor the high southern Alleghanies; in middle Kentucky and Tennessee, and northern Alabama and Mississippi, [89] covering great areas of low rolling limestone hills with nearly pure forests of small bushy trees.

Often cultivated, in several forms, in the northern and eastern states as an ornamental tree and occasionally in the gardens of western and central Europe.

11. Juniperus lucayana Britt. Red Cedar.

Leaves usually opposite, narrow, acute, or gradually narrowed above the middle and acuminate, marked on the back by conspicuous oblong glands. Flowers opening in early March: male elongated, ⅛′ to nearly ¼′ long, with 10 or 12 stamens, their connectives rounded, entire, and bearing usually 3 pollen-sacs: female with scales gradually narrowed above the middle, acute at apex, and obliterated from the ripe fruit. Fruit subglobose to short-oblong, dark blue, covered when ripe with a glaucous bloom, about ¹/₂₄′ in diameter, with a thin skin, sweet resinous flesh, and 1 or 2 seeds; seeds acute, prominently ridged.

A tree, sometimes 50° high, with a trunk occasionally 2° in diameter, small branches erect when the tree is crowded in the forest, spreading when it has grown in open ground and forming a broad flat-topped head often 30° or 40° in diameter, long thin secondary branches erect at the top of the tree and pendulous below, and pendulous branchlets about ¹/₂₄′ in diameter, becoming light red-brown or ashy gray at the end of four or five years after the disappearance of the leaves. Bark thin, light red-brown, separating into long thin scales. Wood light, close, straight-grained, fragrant, dull red; formerly exclusively used in the manufacture of the best lead pencils.

Distribution. Inundated river swamps from southern Georgia, southward to the shores of the Indian River, Florida, and on the west coast of Florida from the northern shores of Charlotte Harbor to the valley of the Apalachicola River, often forming great thickets under the shade of larger trees; along streams and creeks in low woods near Houston, Harris County, and Milano, Milano County, Texas (E. J. Palmer); common in the Bahamas, San Domingo, eastern Cuba, and on the mountains of Jamaica and Antigua.

Often planted for the decoration of squares and cemeteries in the cities and towns in the neighborhood of the coast from Florida to western Louisiana, and now often naturalized beyond the limits of its natural range on the Gulf coast; occasionally cultivated in the temperate countries of Europe, and in cultivation the most beautiful of the Junipers.

12. Juniperus scopulorum Sarg. Red Cedar.

Leaves usually opposite, closely appressed, acute or acuminate, generally marked on the back by obscure elongated glands, dark green, or often pale and very glaucous. Flowers: male with about 6 stamens, their connectives rounded and entire, bearing 4 or 5 anther-sacs: scales of the female flower spreading, acute or acuminate, and obliterated from the mature fruit. Fruit ripening at the end of the second season, nearly globose, ¼′—⅓′ in diameter, bright blue, with a thin skin covered with a glaucous bloom, sweet resinous flesh, and 1 or usually 2 seeds; seeds acute, prominently grooved and angled, about ³/₁₆′ long, with a thick bony outer coat and a small 2-lobed hilum.

A tree, 30°—40° high, with a short stout trunk sometimes 3° in diameter, often divided near the ground into a number of stout spreading stems, thick spreading and ascending branches covered with scaly bark, forming an irregular round-topped head, and slender 4-angled branchlets becoming at the end of three or four years terete and clothed with smooth pale bark separating later into thin scales. Bark dark reddish brown or gray tinged with red, divided-by shallow fissures into narrow flat connected ridges broken on the surface into persistent shredded scales.

Distribution. Scattered often singly over dry rocky ridges, usually at altitudes of 5000° or 6000° but occasionally ascending in Colorado to 9000° above the sea, from the eastern foothill region of the Rocky Mountains from Alberta to the Black Hills of South Dakota, the valley of the Niobrara River, Sheridan County, northwestern Nebraska (J. M. Bates) and to western Texas and eastern and northern New Mexico, and westward to eastern Oregon, Nevada, and northern Arizona; descending to the sea-level in Washington on the shores of the northern part of Puget Sound and on the islands and mainland about the Gulf of Georgia, British Columbia.

II. TAXACEÆ.

Slightly resinous trees and shrubs, producing when cut vigorous stump shoots, with fissured or scaly bark, light-colored durable close-grained wood, slender branchlets, linear-lanceolate entire rigid acuminate spirally disposed leaves, usually appearing 2-ranked by a twist in their short compressed petioles and persistent for many years, and small ovoid acute buds. Flowers opening in early spring from buds formed the previous autumn, diœcious or monœcious, axillary and solitary, surrounded by the persistent decussate scales of the buds, the male composed of numerous filaments united into a column, [91] each filament surmounted by several more or less united pendant pollen-cells; the female of a single erect ovule, becoming at maturity a seed with a hard bony shell, raised upon or more or less surrounded by the enlarged and fleshy aril-like disk of the flower; embryo axile, in fleshy ruminate or uniform albumen; cotyledons 2, shorter than the superior radicle. Of the ten genera widely distributed over the two hemispheres, two occur in North America.

CONSPECTUS OF THE NORTH AMERICAN GENERA.

1. TORREYA ARN.

Glabrous fœtid or pungent aromatic trees, with fissured bark and verticillate or opposite spreading or drooping branches. Leaves thin, long-pointed, abruptly contracted at base, dark green, lustrous and slightly rounded above, thickened and revolute on the margins, with pale bands of stomata on each side of the midvein on the lower surface. Flowers diœcious; the male crowded in the axils of adjacent leaves, on shoots of the previous year, oval or oblong, composed of 6 or 8 close whorls each of 4 stamens, subverticillately arranged on a slender axis; filaments stout and expanded above into 4 globose yellow pollen-sacs united into a half ring, their connectives produced above the cells; the female on shoots of the year less numerous and scattered, sessile, the ovule surrounded by and finally inclosed in an ovoid urn-shaped fleshy sac, and becoming at the end of the second season an oblong-ovate yellow-brown seed, rounded and apiculate at apex, acute and marked at base by the large dark hilum; seed-coat thick and woody, its inner layer folded into the thick white albumen, surrounded and finally inclosed in the thick green or purple enlarged disk of the flower composed of thin flat easily separable fibers, splitting longitudinally when ripe into two parts and separating from the basal scales persistent on the short stout stalk of the seed.

Torreya is now confined to Florida and Georgia, western California, Japan, the island of Quelpart, and central and northern China. Four species are recognized. Of the exotic species the Japanese Torreya nucifera S. & Z. is occasionally cultivated in the eastern states.

The genus is named in honor of Dr. John Torrey, the distinguished American botanist.

CONSPECTUS OF THE NORTH AMERICAN SPECIES.

1. Torreya taxifolia Arn. Stinking Cedar. Torreya.

Leaves slightly falcate, 1½′ long, about ⅛′ wide, somewhat rounded, dark green and lustrous above, paler and marked below with broad bands of stomata. Flowers appearing in March and April; male with pale yellow anthers; female broadly ovoid, with a dark purple fleshy covering to the ovule, ⅛′ long, and inclosed at the base by broad thin rounded scales. Seed fully grown at midsummer, slightly obovoid, dark purple, 1′—1¼′ long, ¾′ thick, with a thin leathery covering, a light red-brown seed-coat furnished on the inner surface with 2 opposite [92] longitudinal thin ridges extending from the base toward the apex, and conspicuously ruminate albumen.

A tree, occasionally 40° high, with a short trunk 1°—2° in diameter, whorls of spreading slightly pendulous branches forming a rather open pyramidal head tapering from a broad base. Bark ½′ thick, brown faintly tinged with orange color, and irregularly divided by broad shallow fissures into wide low ridges slightly rounded on the back and covered with thin closely appressed scales. Wood hard, strong, clear bright yellow, with thin lighter colored sapwood; largely used for fence-posts.

Distribution. On bluffs along the eastern bank of the Apalachicola River, Florida, from River Junction to the neighborhood of Bristol, Liberty County, and in the southwestern corner of Decatur County, Georgia (R. M. Harper). Rare and local.

2. Torreya californica Torr. California Nutmeg.

Leaves slightly falcate, nearly flat, dark green and lustrous on the upper, somewhat paler and marked below with a narrow band of stomata, tipped with slender callous [93] points, 1′—3½′ long, ¹/₁₆′—⅛′ wide. Flowers appearing in March and April; male with broadly ovate acute scales; female nearly ¼′ long, with oblong-ovate rounded scales. Seed ovoid or oblong-ovoid, 1′—1½′ long, light green more or less streaked with purple.

A tree, 50°—70° but occasionally 100° high, with a trunk 1°—2° or rarely 4° in diameter, and whorls of spreading slender slightly pendulous branches forming a handsome pyramidal and in old age a round-topped head. Bark ⅓′—½′ thick, gray-brown tinged with orange color, deeply and irregularly divided by broad fissures into narrow ridges covered with elongated loosely appressed plate-like scales. Wood light, soft, close-grained, clear light yellow, with thin nearly white sapwood; occasionally used for fence-posts.

Distribution. Borders of mountain streams, California, nowhere common but widely distributed from Mendocino County to the Santa Cruz Mountains in the coast region and along the western slopes of the Sierra Nevada from Eldorado to Tulare Counties at altitudes of 3000°—5000° above the sea; most abundant and of its largest size on the northern coast ranges.

2. TAXUS L. Yew.

Trees or shrubs, with brown or dark purple scaly bark, and spreading usually horizontal branches. Leaves flat, often falcate, gradually narrowed at the base, dark green, smooth and keeled on the upper surface, paler, papillate, and stomatiferous on the lower surface, their margins slightly thickened and revolute. Flowers diœcious or monœcious: the male composed of a slender stipe bearing at the apex a globular head of 4—8 pale yellow stamens consisting of 4—6 conic pendant pollen-sacs peltately connate from the end of a short filament; the female sessile in the axils of the upper scale-like bracts of a short axillary branch, the ovule erect, sessile on a ring-like disk, ripening in the autumn into an ovoid-oblong seed gradually narrowed and short-pointed at apex, marked at base by the much-depressed hilum, about ⅓′ long, entirely or nearly surrounded by but free from the now thickened succulent translucent sweet scarlet aril-like disk of the flower open at apex; seed-coat thick, of two layers, the outer thin and membranaceous or fleshy, the inner much thicker and somewhat woody; albumen uniform.

Taxus with six or seven species, which can be distinguished only by their leaf characters and habit, is widely distributed through the northern hemisphere, and is found in eastern North America where two species occur, in Pacific North America, Mexico, Europe, northern Africa, western and southern Asia, China, and Japan. Of the exotic species the European, African, and Asiatic Taxus baccata L., and its numerous varieties, is often cultivated in the United States, especially in the more temperate parts of the country, and is replaced with advantage by the hardier Taxus cuspidata S. & Z., of eastern Asia in the northern states, where the native shrubby Taxus canadensis Marsh, with monœcious flowers is sometimes cultivated.

CONSPECTUS OF THE NORTH AMERICAN ARBORESCENT SPECIES.

1. Taxus brevifolia Nutt. Yew.

Leaves ½′—1′ long, about ¹/₁₆′ wide, dark yellow-green above, rather paler below, with stout midribs, and slender yellow petioles ¹/₁₂′ long, persistent for 5—12 years. Flowers and fruit as in the genus.

A tree, usually 40°—50° but occasionally 70°—80° high, with a tall straight trunk 1°—2° or rarely 4½° in diameter, frequently unsymmetrical, with one diameter much exceeding the other, and irregularly lobed, with broad rounded lobes, and long slender horizontal or slightly pendulous branches forming a broad open conical head. Bark about ¼′ thick [94] and covered with small thin dark red-purple scales. Wood heavy, hard, strong, bright red, with thin light yellow sapwood; used for fence-posts and by the Indians of the northwest coast for paddles, spear-handles, bows, and other small articles.

Distribution. Banks of mountain streams, deep gorges, and damp ravines, growing usually under large coniferous trees; nowhere abundant, but widely distributed usually in single individuals or in small clumps from the extreme southern part of Alaska, southward along the coast ranges of British Columbia, Washington, and Oregon, where it attains its greatest size; along the coast ranges of California as far south as the Bay of Monterey, and along the western slopes of the Sierra Nevada to Tulare County at altitudes between 5000° and 8000° above the sea-level, ranging eastward in British Columbia to the Selkirk Mountains, and over the mountains of Washington and Oregon to the western slopes of the continental divide in northern Montana; in the interior much smaller than near the coast and often shrubby in habit.

2. Taxus floridana Chapm. Yew.

Leaves usually conspicuously falcate, ¾′ to nearly 1′ long, ¹/₁₆′—¹/₁₂′ wide, dark green above, pale below, with obscure midribs and slender petioles about ¹/₁₆′ in length. Flowers appearing in March. Fruit ripening in October.

A bushy tree, rarely 25° high, with a short trunk occasionally 1° in diameter, and numerous stout spreading branches; more often shrubby in habit and 12°—15° tall. Bark ⅛′ thick, dark purple-brown, smooth, compact, occasionally separating into large thin irregular plate-like scales. Wood heavy, hard, very close-grained, dark brown tinged with red, with thin nearly white sapwood.

Distribution. River bluffs and ravines on the eastern bank of the Apalachicola River, in Gadsden County, Florida, from Aspalaga to the neighborhood of Bristol.

Class 2. ANGIOSPERMÆ.

Carpels or pistils consisting of a closed cavity containing the ovules and becoming the fruit.

Division 1. Monocotyledons.

Stems with woody fibres distributed irregularly through them, but without pith or annual layers of growth. Parts of the flower in 3’s; ovary superior; embryo with a single cotyledon. Leaves parallel-veined, alternate, long-persistent, without stipules.

III. PALMÆ.

Trees, growing by a single terminal bud, with stems covered with a thick rind, usually marked below by the ring-like scars of fallen leaf-stalks, and clothed above by their long-persistent sheaths; occasionally stemless. Leaves clustered at the top of the stem, plaited in the bud, fan-shaped or pinnate, their rachis sometimes reduced to a narrow border, long-stalked, with petioles dilated into clasping sheaths of tough fibres (vaginas); on fan-shaped leaves, furnished at the apex on the upper side with a thickened concave body (ligule). Flowers minute, perfect or unisexual, in the axils of small thin mostly deciduous bracts, in large compound clusters (spadix) surrounded by boat-shaped bracts (spathes); sepals and petals free or more or less united; stamens usually 6; anthers 2-celled, introrse, opening longitudinally; ovary 3-celled, with a single ovule in each cell; styles 1—3. Fruit a drupe or berry; embryo cylindric in a cavity of the hard albumen near the circumference of the seed. Of the 130 genera now usually recognized and chiefly inhabitants of the tropics, seven have arborescent representatives in the United States.

CONSPECTUS OF THE NORTH AMERICAN ARBORESCENT GENERA.

1. THRINAX Sw.

Small unarmed trees, with stems covered with pale gray rind. Leaves orbicular, or truncate at the base, thick and firm, usually silvery white on the lower surface, divided [97] to below the middle into narrow acuminate parted segments with thickened margins and midribs; rachis a narrow border, with thin usually undulate margins; ligule thick, concave, pointed, lined while young with hoary tomentum; petioles compressed, rounded above and below, thin and smooth on the margins, with large clasping bright mahogany-red sheaths of slender matted fibres covered with thick hoary tomentum. Spadix interfoliar, stalked, its primary branches short, alternate, flattened, incurved, with numerous slender rounded flower-bearing branchlets; spathes numerous, tubular, coriaceous, cleft and more or less tomentose at the apex. Flowers opening in May and June, and occasionally irregularly in the autumn, solitary, perfect; perianth 6-lobed; stamens inserted on the base of the perianth, with subulate filaments thickened and only slightly united at the base, or nearly triangular and united into a cup adnate to the perianth, and oblong anthers; ovary 1-celled, gradually narrowed into a stout columnar style crowned by a large funnel-formed flat or oblique stigma; ovule basilar, erect. Fruit a globose drupe with juicy bitter ivory-white flesh easily separable from the thin-shelled tawny brown nut. Seed free, erect, slightly flattened at the ends, with an oblong pale conspicuous subbasilar hilum, a short-branched raphe, a thin coat, and uniform albumen more or less deeply penetrated by a broad basal cavity; embryo lateral.

Thrinax is confined to the tropics of the New World and is distributed from southern Florida through the West Indies to the shores of Central America. Seven or eight species are now generally recognized.

The wood of the Florida species is light and soft, with numerous small fibro-vascular bundles, the exterior of the stem being much harder than the spongy interior. The stems are used for the piles of small wharves and turtle-crawls, and the leaves for thatch, and in making hats, baskets, and small ropes.

CONSPECTUS OF THE NORTH AMERICAN SPECIES.

1. Thrinax floridana Sarg. Thatch.

Leaves 2½°—3° in diameter, rather longer than broad, yellow-green and lustrous on the upper surface, silvery white on the lower surface, with a long-pointed, bright orange-colored ligule ¾′ long and broad; petioles 4°—4½° long, pale yellow-green or orange color toward the apex, coated at first with hoary deciduous tomentum, much thickened and tomentose toward the base. Flowers: spadix 3°—3½° long, the primary branches 6′—8′ long and ivory-white, flower-bearing branches 1½′—2′ in length; flowers on slender pedicels nearly ⅛′ long, ivory-white, very fragrant, with an obscurely-lobed perianth, much exserted stamens barely united at the base, and a large stigma. Fruit ⅜′ in diameter, somewhat depressed at the ends; seed from ⅛′ to nearly ¼′ in diameter, dark chestnut-brown.

A tree, with a slightly tapering stem 20°—30° high and 4′—6′ in diameter, clothed to the middle and occasionally almost to the ground with the sheaths of dead leaf-stalks.

Distribution. Florida, dry coral ridges and sandy shores of keys from Long Key to Torch Key, and on the mainland from Cape Romano to Cape Sable.

2. Thrinax Wendlandiana Becc. Thatch.

Leaves 2½°—3° in diameter, orbicular, pale yellow-green, lustrous above, with a thick concave ligule, acuminate or rarely rounded at apex; petioles 2°—4° long, much thickened and tomentose toward the base. Flowers: spadix stalked, 2°—4° long, its primary branches short, flattened, incurved, with numerous terete flower-bearing branchlets; flowers on slender pedicels ¹/₁₀′—⅛′ long, with a deeply lobed perianth, the lobes nearly [99] triangular, acuminate, and a small stigma. Fruit ¼′—⅜′ in diameter, globose; seed from ⅛′—¼′ in diameter, dark chestnut-brown.

A tree, in Florida, with a smooth pale trunk 20°—25° high and 3′—4′ in diameter.

Distribution. Florida: Dade County, Madeira Hummock, Pumpkin Key, Flamingo, and northwest of Cape Sable; also in Cuba and on Mugueres Island, Gulf of Honduras.

3. Thrinax keyensis Sarg. Thatch.

Leaves rather longer than broad, 3°—4° long, the lowest segments parallel with the petiole or spreading from it nearly at right angles, light yellow-green and lustrous on the upper surface, with bright orange-colored margins, below coated while young with deciduous hoary tomentum and pale blue-green and more or less covered with silvery white pubescence at maturity, with a thick pointed ligule 1′ long and wide, lined at first with hoary tomentum; petioles flattened above, obscurely ridged on the lower surface, tomentose while young, pale blue-green, 3°—4° long. Flowers: spadix usually about 6° long, spreading and gracefully incurved, with spathes more or less coated with hoary tomentum, large compressed primary branches, and short bright orange-colored flower-bearing branches; flowers on short thick disk-like pedicels, about ⅛′ long, white, slightly fragrant, with a tubular perianth, the lobes broadly ovate and acute, stamens with nearly triangular filaments united at the base, and a flat stigma. Fruit ¹/₁₆′ to nearly ¼′ in diameter; seed brown, ³/₁₆′ in diameter.

A tree, with a stem often 25° high and 10′—14′ in diameter, raised on a base of thick matted roots 2°—3° high and 18′—20′ in diameter, and a broad head of leaves, the upper erect, the lower pendulous and closely pressed against the stem.

Distribution. Dry, sandy soil close to the beach on the north side of the largest of the Marquesas Keys, and on Crab Key, a small island to the westward of Torch Key, one of the Bahia Honda group, Florida; on the Bahamas.

4. Thrinax microcarpa Sarg. Silvertop Palmetto. Brittle Thatch.

Leaves 2°—3° across, pale green above, silvery white below, more or less thickly coated while young with hoary tomentum, especially on the lower surface, divided near the base almost to the rachis, with an orbicular thick concave ligule lined with a thick coat of white tomentum; petioles thin and flexuose. Flowers: spadix elongated, with short, compressed erect branches slightly spreading below, numerous slender pendulous flower-bearing branches, and long acute spathes deeply parted at the apex, coriaceous and coated above the middle with thick hoary tomentum; flowers on short thick disk-like pedicels, with a [100] cupular perianth, the lobes broadly ovate and acute, stamens with thin nearly triangular exserted filaments slightly united at base and oblong anthers becoming reversed and extrorse at maturity, and a deep orange-colored ovary narrowed above into a short thick style dilated into a large funnel-formed stigma. Fruit globose, ⅛′ in diameter; seed subglobose, bright to dark chestnut-brown, depressed.

Distribution. Dry coral soil, on the shores of Sugar Loaf Sound, and on No Name and Bahia Honda keys, Florida; in Cuba.

2. COCCOTHRINAX Sarg.

Small unarmed trees, with simple or clustered stems or rarely stemless. Leaves orbicular, or truncate at base, pale or silvery white on the lower surface, divided into narrow obliquely folded segments acuminate and divided at apex; rachis narrow; ligules thin, free, erect, concave, pointed at the apex; petioles compressed, slightly rounded and ridged above and below, thin and smooth on the margins, gradually enlarged below into elongated sheaths of coarse fibres forming an open network covered while young by thick hoary tomentum. Spadix interfoliar, paniculate, shorter than the leaf-stalks, its primary branches furnished with numerous short slender pendulous flower-bearing secondary branches; spathes numerous, papery, cleft at the apex. Flowers solitary, perfect, jointed on elongated slender pedicels; perianth cup-shaped, obscurely lobed; stamens 9, inserted on the base of the perianth, with subulate filaments enlarged and barely united at the base, and oblong anthers; ovary 1-celled, narrowed into a slender style crowned by a funnel-formed oblique stigma; ovule basilar, erect. Fruit a subglobose berry raised on the thickened torus of the flower, with thick juicy black flesh. Seed free, erect, depressed-globose, with a thick hard vertically grooved shell deeply infolded in the bony albumen; hilum subbasilar, minute; raphe hidden in the folds of the seed-coat; embryo lateral.

Coccothrinax is confined to the tropics of the New World. Two species, of which one is stemless, inhabit southern Florida, and at least two other species are scattered over several of the West Indian islands.

Coccothrinax, from κόκκος and Thrinax, is in allusion to the berry-like fruit.

1. Coccothrinax jucunda Sarg. Brittle Thatch.

Leaves nearly orbicular, the lower segments usually parallel with the petiole, thin and brittle, 18′—24′ in diameter, divided below the middle of the leaf or toward its base nearly [101] to the ligule, with much-thickened bright orange-colored midribs and margins, pale yellow-green and lustrous on the upper surface, bright silvery white and coated at first on the lower surface with hoary deciduous pubescence, with a thin undulate obtusely short-pointed dark orange-colored rachis, and a thin concave crescent-shaped often oblique slightly undulate short-pointed and light or dark orange-colored ligule ¾′ wide, ⅓′ deep; petioles slender, pale yellow-green, 2½°—3° long. Flowers: spadix 18′—24′ long, with flattened stalks, slender much-flattened primary branches 8′—10′ long, light orange-colored slender terete flower-bearing branches 1½′—3′ long, and pale reddish brown spathes coated toward the ends with pale pubescence; flowers opening in June and irregularly also in the autumn on ridged spreading pedicels ⅛′ long, with an orange-colored ovary surmounted by an elongated style dilated into a rose-colored stigma. Fruit ripening at the end of six months, from ½′—¾′ in diameter, bright green at first when fully grown, becoming deep violet color, with succulent very juicy flesh, ultimately black and lustrous; seed light tawny brown.

A tree, with a stem slightly enlarged from the ground upward, 15°—25° high, 4′—6′ thick, covered with pale blue rind, and surmounted by a broad head of leaves at first erect, then spreading and ultimately pendulous. Wood used for the piles of small wharves and turtle-crawls. The soft tough young leaves are made into hats and baskets.

Distribution. Dry coral ridges and sandy flats from the shores of Bay Biscayne along many of the southern keys to the Marquesas group (var. marquesensis Becc.) Florida; and on the Bahamas (var. macrosperma Becc.).

3. SABAL Adans. Palmetto.

Unarmed trees, with stout columnar stems covered with red-brown rind. Leaves flabellate, tough and coriaceous, divided into many narrow long-pointed parted segments plicately folded at base, often separating on the margins into narrow threads; rachis extending nearly to the middle of the leaves, rounded and broadly winged toward the base on the lower side, thin and acute on the upper side; ligule adnate to the rachis, acute, concave, with thin incurved entire margins; petioles rounded and concave on the lower side, conspicuously ridged on the upper side, acute and entire on the margins, with elongated chestnut-brown shining sheaths of stout fibres. Spadix interfoliar, stalked, decompound, with a flattened stem, short branches, slender densely flowered ultimate branches, and numerous acuminate spathes, the outer persistent and becoming broad and [102] woody. Flowers solitary, perfect; calyx tabular, unequally lobed, the lobes slightly imbricated in the bud; corolla deeply lobed, with narrow ovate-oblong concave acute lobes valvate at the apex in the bud; stamens 6, those opposite the corolla lobes rather longer than the others, with subulate filaments united below into a shallow cup adnate to the tube of the corolla, and ovoid anthers, their cells free and spreading at the base; ovary of 3 carpels, 3-lobed, 3-celled, gradually narrowed into an elongated 3-lobed style truncate and stigmatic at the apex; ovule basilar, erect. Fruit a small black 1 or 2 or 3-lobed short-stemmed berry with thin sweet dry flesh. Seed depressed-globose, marked on the side by the prominent micropyle, with a shallow pit near the minute basal hilum, a thin seed-coat, and a ventral raphe; embryo minute, dorsal, in horny uniform albumen penetrated by a hard shallow basal cavity filled by the thickening of the seed-coat.

Sabal belongs to the New World, and is distributed from the Bermuda Islands and the South Atlantic and Gulf states of North America through the West Indies to Venezuela and Mexico.

Of the eight species now recognized four inhabit the United States; of these two are small stemless plants.

CONSPECTUS OF THE NORTH AMERICAN ARBORESCENT SPECIES.

1. Sabal Palmetto R. & S. Cabbage Tree. Cabbage Palmetto.

Leaves 5°—6° long and 7°—8° broad, dark green and lustrous, deeply divided into narrow parted recurved segments, with ligules 4′ long and more or less unsymmetrical at apex; petioles 6°—7° long and 1½′ wide at apex. Flowers: spadix 2°—2½° long, with slender incurved branches, slender ultimate divisions, and thin secondary spathes flushed with red at apex and conspicuously marked by pale slender longitudinal veins; flowers in the axils of minute deciduous bracts much shorter than the perianth, opening in June. Fruit ripening late in the autumn, subglobose or slightly obovoid, gradually narrowed at base, 1-seeded, about ⅓′ in diameter; seed light bright chestnut-colored, ¼′ broad.

A tree, often 40°—50° and occasionally 80°—90° high, with a tall clear trunk often 2° in diameter, sometimes branched by the destruction of the terminal bud, divided by shallow [103] irregular interrupted fissures into broad ridges, with a short pointed knob-like underground stem surrounded by a dense mass of contorted roots often 4° or 5° in diameter and 5° or 6° deep, from which tough light orange-colored roots often nearly ½′ in diameter penetrate the soil for a distance of 15° or 20°, and a broad crown of leaves at first upright, then spreading nearly at right angles with the stem, and finally pendulous. Wood light, soft, pale brown, or occasionally nearly black, with numerous hard fibro-vascular bundles, the outer rim about 2′ thick and much lighter and softer than the interior. In the southern states the trunks are used for wharf-piles, and polished cross sections of the stem sometimes serve for the tops of small tables; the wood is largely manufactured into canes. From the sheaths of young leaves the bristles of scrubbing-brushes are made. The large succulent leaf-buds are cooked and eaten as a vegetable, and coarse hats, mats, and baskets are made from the leaves. Pieces of the spongy bark of the stem are used as a substitute for scrubbing-brushes.

Distribution. Sandy soil in the immediate neighborhood of the coast from the neighborhood of Cape Hatteras and Smith Island at the mouth of Cape Fear River, North Carolina, southward near the coast to northern Florida; in Florida extending across the peninsula and south to Upper Metacomb Key, and along the west coast to Saint Andrews Bay; most abundant and of its largest size on the west coast of the Florida peninsula.

2. Sabal texana Becc. Palmetto.

Leaves dark yellow-green and lustrous, 5°—6° long, often 7° wide, divided nearly to the middle into narrow divided segments, with thickened pale margins separating into long thin fibres, with ligules about 6′ long; petioles 7°—8° long, 1½′ wide at the apex. Flowers: spadix 7°—8° long, with stout ultimate divisions; flowers in Texas appearing in March or April in the axils of persistent bracts half as long as the perianth. Fruit ripening early in the summer, globose, often 2 or 3-lobed; seeds nearly ½′ broad and ¼′ wide, dark chestnut-brown, with a broad shallow basal cavity, and a conspicuous orange-colored hilum.

A tree, with a trunk 30°—50° high, often 2½° in diameter, and a broad head of erect ultimately pendulous leaves. Wood light, soft, pale brown tinged with red, with thick light-colored rather inconspicuous fibro-vascular bundles, the outer rim 1′ thick, soft, and light colored. On the Gulf coast the trunks are used for wharf-piles, and on the lower Rio Grande the leaves for the thatch of houses.

Distribution. Rich soil of the bottom-lands on the Bernado River, Cameron County, and near the mouth of the Rio Grande, Texas, and southward in Mexico in the neighborhood of the coast.

Frequently planted as a street tree in the towns in the lower Rio Grande valley.

4. WASHINGTONIA H. Wendl.

Trees, with stout columnar stems and broad crowns of erect and spreading finally pendulous leaves. Leaves flabellate, divided nearly to the middle into many narrow deeply parted recurved segments separating on the margins into numerous slender pale fibres; rachis short, slightly rounded on the back, gradually narrowed from a broad base, with concave margins furnished below with narrow erect wings, and slender and acute above; ligule elongated, oblong, thin and laciniate on the margins; petioles elongated, broad and thin, flattened or slightly concave on the upper side, rounded on the lower, armed irregularly with broad thin large and small straight or hooked spines confluent into a thin bright orange-colored cartilaginous margin, gradually enlarged at base into thick broad concave bright chestnut-brown sheaths composed of a network of thin strong fibres. Spadix interfoliar, stalked, elongated, paniculate, with pendulous flower-bearing ultimate divisions and numerous long spathes. Flowers perfect, jointed on thick disk-like pedicels; calyx tubular, scarious, thickened at base, gradually enlarged and slightly lobed at apex, the lobes imbricated in the bud; corolla funnel-formed, with a fleshy tube inclosed in the calyx and about half as long as the lanceolate lobes thickened and glandular on the inner surface at the base, imbricated in the bud; stamens inserted on the tube of the corolla, with free filaments thickened near the middle and linear-oblong anthers; ovary 3-lobed, 3-celled, with slender elongated flexuose styles stigmatic at apex; ovules lateral, erect. Fruit a small ellipsoidal short-stalked black berry with thin dry flesh. Seed free, erect, oblong-ovoid, concave above, with a flat base depressed in the centre, a minute sublateral hilum, a broad conspicuous rachis, a minute lateral micropyle, and a thin pale chestnut-brown inner coat closely investing the simple horny albumen; embryo minute, lateral, with the radicle turned toward the base of the fruit.

Three species of Washingtonia are known: one inhabits the interior dry region of southern California and the adjacent parts of Lower California, and the others the mountain cañons of western Sonora and southern Lower California.

1. Washingtonia filamentosa O. Kuntze. Desert Palm. Fan Palm.

Leaves 5°—6° long and 4°—5° wide, light green, slightly tomentose on the folds; petioles [105] 4°—6° long and about 2′ broad at apex, with sheaths 16′—18′ long and 12′—14′ wide, and ligules 4′ long and cut irregularly into long narrow lobes. Flowers: spadix 10°—12° long, 3 or 4 being produced each year from the axils of upper leaves, the outer spathe inclosing the bud, narrow, elongated, and glabrous, those of the secondary branches coriaceous, yellow tinged with brown, and laciniate at apex; flowers slightly fragrant, opening late in May or early in June. Fruit produced in great profusion, ripening in September, ⅓′ long; seed ¼′ long, ⅛′ thick.

A tree, occasionally 75° high, with a trunk sometimes 50°—60° tall and 2°—3° in diameter, covered with a thick light red-brown scaly rind and clothed with a thick thatch of dead pendant leaves descending in a regular cone from the broad crown of living leaves sometimes nearly to the ground. Wood light and soft, with numerous conspicuous dark orange-colored fibro-vascular bundles. The fruit is gathered and used as food by the Indians.

Distribution. Often forming extensive groves or small isolated clumps in wet usually alkali soil in depressions along the northern and northwestern margins of the Colorado Desert in southern California, sometimes extending for several miles up the cañons of the San Bernardino and San Jacinto mountains; and in Lower California.

Now largely cultivated in southern California, New Orleans, southern Europe, and other temperate regions.

5. ACŒLORRAPHE H. Wendl.

Trees, with tall slender often clustered stems clothed for many years with the sheathing bases of the petioles of fallen leaves. Leaves suborbicular, divided into numerous two-parted segments plicately folded at the base; rachis short, acute; ligule thin, concave, furnished with a broad membranaceous dark red-brown deciduous border; petioles slender, flat or slightly concave on the upper side, rounded and ridged on the lower side, with a broad high rounded ridge, thickened and cartilaginous on the margins, more or less furnished with stout or slender flattened teeth; vagina thin and firm, bright mahogany red, lustrous, closely infolding the stem, its fibres thin and tough. Spadix paniculate, interpetiolar, its rachis slender, compressed, ultimate branches, numerous, slender, elongated, gracefully drooping, hoary-tomentose, the primary branches flattened, the secondary terete in the axils of ovate acute chestnut-brown bracts; spathes flattened, thick and firm, deeply two-cleft and furnished at apex with a red-brown membranaceous border, inclosing the rachis of the panicle, each primary branch with its spathe and the node of the rachis below it inclosed in a separate spathe, the whole surrounded by the larger spathe of the node next below. Flowers perfect, minute, sessile on the ultimate branches of the spadix, in the axils of ovate acute chestnut-brown caducous bracts, solitary toward the end of the branches and in two- or three-flowered clusters near their base; calyx truncate at base, divided into three broadly ovate sepals dentate on the margins, valvate in æstivation, enlarged and persistent under the fruit; corolla three-parted nearly to the base, its divisions valvate in æstivation, oblong-ovate, thick, concave and thickened at apex, deciduous; stamens six, included; filaments nearly triangular, united below into a cup adnate to the short tube of the corolla; anthers short-oblong, attached on the back below the middle, introrse, two-celled, the cells opening longitudinally; ovary obovoid, of three carpels, each with two deep depressions on their outer face, united into a slender style; stigma minute, terminal, persistent on the fruit; ovule solitary, erect from the bottom of the cell, anatropous. Fruit drupaceous, subglobose, one-seeded, black and lustrous; exocarp thin and fleshy; endocarp thin, crustaceous; seed erect, free, subglobose, light chestnut-brown; testa thin and hard; hilum small, suborbicular; raphe ventral, oblong, elongated, black, slightly prominent, without ramifications; embryo lateral; albumen homogeneous.

Two species of Acœlorraphe have been distinguished; they inhabit southern Florida, and one species occurs also in Cuba and on the Bahama Islands.

The generic name, from ἀ priv., Κοῖλος and ῥαφη, refers to the character of the seed.

CONSPECTUS OF THE NORTH AMERICAN SPECIES.

1. Acœlorraphe Wrightii Becc.

Leaves 30′—36′ in diameter, thin, light green, divided only to the middle, the divisions of the primary lobes 3½′—9′ long; petioles thin, gradually tapering from the base, 40′—60′ in length, armed throughout with stout straight or incurved teeth. Flowers: spadix 4°—6° long; flowers ⅛′—⅙′ long, with a light chestnut-brown calyx and a pale yellow-green corolla. Fruit ¼′ in diameter.

A tree with numerous stems, in Florida sometimes 10 metres high, forming great thickets.

Distribution. Dade County, Florida, from the rear of Madeira Hummock to Cape Sable, in swamps of fresh or brackish water at some distance from the coast; also in Cuba and on the Bahamas.

2. Acœlorraphe arborescens Becc.

Leaves about 2° in diameter, light yellow-green on the upper surface, blue-green or glaucescent on the lower surface, divided nearly to the base into numerous lobes slightly thickened at the pale yellow midribs and margins; petioles 18′—24′ long, armed, except toward the apex, with stout flattened curved orange-colored teeth. Flowers: spadix [107] 3°—4° long, with a slender much-flattened stalk, panicled lower branches 18′—20′ in length, and 6—8 thick firm pale green conspicuously ribbed spathes dilated at apex into a narrow border; flowers with a light chestnut-brown calyx and a pale yellow-green corolla. Fruit globose, ⅓′ in diameter; seed somewhat flattened below, with a pale vertical mark on the lower side, and a hilum joined to the micropyle by a pale band.

A tree, from 30°—40° high, with 1 or several clustered erect inclining or occasionally semiprostrate stems 3′—4′ in diameter, covered almost to the ground by the closely clasping bases of the leaf-stalks and below with a thick pale rind.

Distribution. Low undrained soil covered for many months of every year in water from 1′—18′ deep, occasionally occupying almost exclusively areas of several acres in extent or more often scattered among Cypress-trees or Royal Palms, in the swamps and along the hummocks adjacent to the Chokoloskee River and its tributaries and at the head of East River, Whitewater Bay, in southwestern Florida.

6. ROYSTONEA Cook. Royal Palm.

Unarmed trees, with massive stems enlarged near the middle, and terminating in long slender bright green cylinders formed by the densely imbricated sheaths of the leaf-stalks. Leaves equally pinnate, with linear-lanceolate long-pointed unequally cleft plicately-folded pinnæ inserted obliquely on the upper side of the rachis, folded together at the base, with thin midribs and margins; rachis convex on the back, broad toward the base of the leaf and acute toward its apex; petioles semicylindric, gradually enlarged into thick elongated green sheaths. Spadix large, decompound, produced near the base of the green part of the stem, with long pendulous branches and 2 spathes, the outer semicylindric and as long as the spadix, the inner splitting ventrally and inclosing the branches of the spadix. Flowers monœcious, in a loose spiral, toward the base of the branch in 3-flowered clusters, with a central staminate and smaller lateral pistillate flowers, higher on the branch the staminate in 2-flowered clusters; calyx of the staminate flower of minute broadly ovate obtuse scarious sepals imbricated in the bud, much shorter than the corolla; petals nearly equal, valvate in the bud, ovate or obovate, acute, slightly united at the base, coriaceous; stamens 6, 9, or 12, with subulate filaments united below and adnate to the base of the corolla, and large ovate-sagittate anthers, the cells free below; ovary rudimentary, subglobose or 3-lobed; pistillate flowers much smaller, ovoid-conic; sepals obtuse; corolla erect, divided to the middle into acute erect lobes incurved at apex; staminodia 6, scale-like, united into a cup adnate to the corolla; ovary subglobose, obscurely 2 or 3-lobed, [108] 2 or 3-celled, gibbous, the cells crowned with a 3-lobed stigma becoming subbasilar on the fruit; ovule ascending. Fruit a short-stalked drupe with thin crustaceous flesh. Seed oblong-reniform, marked by the conspicuous fibrous reticulate branches of the raphe radiating from the narrow basal hilum, and covered with a thin crustaceous coat; embryo minute, cylindric, lateral, in uniform albumen.

Roystonea is confined to the tropics of the New World, where two or three species occur.

The genus as here limited was named for General Roy Stone of the United States army.

1. Roystonea regia Cook. Royal Palm.

Leaves 10°—12° long, closely pinnate, the pinnæ, 2½°—3° long, 1½′ wide near the base of the leaf, and gradually decreasing in size toward its apex, deep green with slender conspicuous veins, and covered below with minute pale glandular dots; petioles almost terete, concave near the base, with thin edges separating irregularly into pale fibres, and enlarged into bright green cylindrical clasping bases 8° or 9° long and more or less covered with dark chaffy scales. Flowers: spadix about 2° long, with a nearly terete stem and slightly ridged primary and secondary branches compressed above, abruptly enlarged at the base, and simple slender flexuose long-pointed flower-bearing branchlets 3′—6′ long, pendant and closely pressed against the secondary branches; flowers opening in Florida in January and February, the staminate nearly ¼′ long and rather more than twice as long as the pistillate. Fruit oblong-obovoid, full and rounded at apex, narrowed at base, violet-blue, about ½′ long, with a thin outer coat and a light red-brown inner coat, loose and fibrous on the outer surface, and closely investing the thin light brown seed.

A tree, 80°—100° high, with a trunk rising from an abruptly enlarged base, gradually tapering from the middle to the ends and often 2° in diameter, covered with light gray rind tinged with orange color, marked with dark blotches and irregularly broken into minute plates, the green upper portion 8°—10° long, and a broad head of gracefully drooping leaves. Wood of the interior of the stem spongy, pale brown, much lighter than the hard exterior rim, containing numerous dark conspicuous fibro-vascular bundles. The outer portion of the stem is made into canes, and the trunks are sometimes used for wharf-piles and in construction.

Distribution. Florida, hummocks on Rogue River twenty miles east of Caximbas Bay, on some of the Everglades Keys, Long’s Key, and formerly on the shores of Bay Biscayne near the mouth of Little River; common in the West Indies and Central America.

Largely cultivated as an ornamental tree in tropical countries, and often planted to form avenues, for which its tall pale columnar stems and noble heads of graceful foliage make it valuable.

7. PSEUDOPHŒNIX H. Wendl.

A tree, with a slender stem abruptly enlarged at the base or tapering from the middle to the ends, covered with thin pale blue or nearly white rind, and conspicuously marked by the dark scars of fallen leaf-stalks. Leaves erect, abruptly pinnate, with crowded linear-lanceolate acuminate leaflets increasing in length and width from the ends to the middle of the leaf, thick and firm in texture, dark yellow-green above, pale and glaucous below; rachis convex on the lower side, concave on the upper side near the base of the leaf, with thin margins, becoming toward the apex of the leaf flat and narrowed below and acute above, marked on the sides at the base with dark gland-like excrescences; petioles short, concave above, with thin entire margins separating into slender fibres, gradually enlarged into broad thick sheaths of short brittle fibres. Spadix interfoliar, compound, pendulous, stalked, much shorter than the leaves, with spreading primary branches, stout and much flattened toward the base, slender and rounded above the middle, furnished at the base with a thickened ear-like body, slender secondary branches, short thin rigid densely flowered ultimate divisions, and compressed light green double spathes erose on their thin dark brown margins. Flowers on slender pedicels articulate by an expanded base, widely scattered on the ultimate branches of the spadix, staminate and bisexual in the same inflorescence; calyx reduced to the saucer-like rim of the thickened receptacle, undulate on the margin, the rounded angles alternating with the petals; petals 3, valvate in the bud, oblong, rounded at apex, thick conspicuously longitudinally veined, persistent; stamens 6, with short flattened nearly triangular filaments slightly united at the base into a narrow fleshy disk, and triangular cordate anthers attached at the base in a cavity on their outer face, 2-celled, the cells opening by lateral slits; styles of the perfect flower 3-lobed at the apex with obtuse appressed lobes, that of the sterile flower as long or longer than that of the perfect flower, more slender and tapering into a narrow 3-pointed apex. Fruit a stalked globose 2 or 3-lobed orange-scarlet thin-fleshed drupe marked by the lateral style and surrounded below by the withered remnants of the flower; pedicel abruptly enlarged at base, articulate from a persistent cushion-like body furnished in the centre with a minute point penetrating a cavity in the base of the pedicel. Seed subglobose, free, erect, with a basal hilum and a thin light red-brown coat marked by the pale conspicuous ascending 2 or 3-branched raphe; embryo minute, basal, in uniform horny albumen.

Pseudophœnix with a single species inhabits the keys of southern Florida, and the Bahamas.

The generic name is in allusion to a fancied resemblance to Phœnix, a genus of Palms.

1. Pseudophœnix vinifera Becc.

Leaves 5°—6° long, with pinnæ often 18′ long and 1′ wide near the middle of the leaf, [110] becoming at its extremities not more than half as long and wide; petioles 6′—8′ in length. Flowers: spadix 3° long and 2½° wide. Fruit ripening in May and June, ½′—¾′ in diameter on a peduncle ¼′ long; seed ¼′ in diameter.

Distribution. Florida, east end of Elliot’s Key, and east end of Key Largo near the southern shore, here forming a grove of 200 or 300 plants; more common on the Bahamas.

IV. LILIACEÆ.

YUCCÆ.

Leaves, alternate, linear-lanceolate. Flowers in terminal panicles; sepals and petals nearly similar, subequal, withering-persistent; ovary with more or less deeply introduced dorsal partitions; ovules numerous, 2-ranked in each cell; embryo subulate, obliquely placed across the seed; cotyledon arched in germination.

Yuccæ as here limited consists of two American genera, Hesperaloe, with two species, low plants of Texas and Mexico, and Yucca.

1. YUCCA L.

Trees with simple or branched stems prolonged by axillary naked buds, dark thick corky bark, light fibrous wood in concentric layers, and large stout horizontal roots; or often stemless. Leaves involute in the bud, at first erect, usually becoming reflexed, abruptly narrowed above the broad thickened clasping base, usually widest near the middle, concave on the upper surface, involute toward the horny usually sharp-pointed apex, convex and often slightly keeled toward the base on the lower surface, the margins serrulate or filamentose, light or dull green. Flowers fertilized by insects and opening for a single night, on slender pedicels in 2 or 3-flowered clusters or singly at the base of the large compound panicle furnished with conspicuous leathery white or slightly colored bracts, those at the base of the pedicels thin and scarious; perianth cup-shaped, with thick ovate-lanceolate creamy white segments more or less united at base, usually furnished with small tufts of white hairs at the apex, those of the outer rank narrower, shorter, and more colored than the more delicate petal-like segments of the inner rank; stamens 6, in 2 series, free, shorter than the ovary (as long in 1), white, with club-shaped fleshy filaments, obtuse and slightly 3-lobed at the apex, and cordate emarginate anthers attached on the back, the cells opening longitudinally, curling backward and expelling the large globose powdery pollen-grains; ovary oblong, 6-sided, sessile or stalked, with nectar-glands within the partitions, dull greenish white, 3-celled, gradually narrowed into a short or elongated 3-lobed ivory-white style forming a triangular stigmatic tube. Fruit oblong or oval, more or less distinctly 6-angled, 6-celled, usually beaked at the apex, baccate and indehiscent or capsular and 3-valved, the valves finally separating at the apex; pericarp of 2 coats, the outer at maturity thick, succulent and juicy, thin, dry and leathery, or thin and woody. Seeds compressed, triangular, obovoid, or obliquely ovoid or orbicular, thick, with a narrow 2-edged rim, or thin, with a wide or narrow brittle margin; seed-coat thin, black, slightly rugose or smooth; embryo in plain or rarely ruminate hard farinaceous oily albumen; cotyledon much longer than the short radicle turned toward the small oblong white hilum.

Yucca is confined to the New World and is distributed from Bermuda and the eastern Antilles, through the south Atlantic and Gulf states to Oklahoma and Arkansas, and through New Mexico and northward along the eastern base of the Rocky Mountains to South Dakota, westward to middle California, and southward through Arizona, Mexico, and Lower California to Central America. About thirty species with many varieties and probable hybrids are recognized. Of the species which inhabit the territory of the United States nine assume the habit and attain the size of small trees. The root-stalks of Yucca are used as a substitute for soap, and ropes, baskets, and mats are made from the tough fibres of the leaves. Many of the species are cultivated, especially in countries of scanty rainfall, for their great clusters of beautiful flowers, or in hedges to protect gardens from cattle.

CONSPECTUS OF THE ARBORESCENT SPECIES OF THE UNITED STATES.

1. Yucca aloifolia L. Spanish Bayonet.

Leaves 18′—32′ long, 1¼′—2½′ wide, erect, rigid, conspicuously narrowed above the light green base, widest above the middle, slightly concave on the upper surface, smooth, dark rich green, with a stiff dark red-brown tip, and horny finely and irregularly serrate margins; long-persistent. Flowers from June until August on stout pedicels, in nearly sessile glabrous or slightly pubescent panicles 18′—24′ long; perianth 1′—1½′ in length and 3′ or 4′ across when fully expanded, the segments ovate, thick and tumid toward the base, those of the outer rank rounded and often marked with purple at apex, the inner acuminate [112] and short-pointed; stamens as long or sometimes a little longer than the light green ovary raised on a short stout stipe. Fruit ripening from August to October, elongated, ellipsoidal, hexagonal, 3′—4′ long, 1¼′—1½′ thick, light green when fully grown, and in ripening turning dark purple, the outer and inner coats forming a thick succulent mass of bitter-sweet juicy flesh, finally becoming black and drying on its stalk; seeds ¼′—⅓′ wide, about ¹/₁₆′ thick, with a thin narrow ring-like border to the rim.

A tree, occasionally 25° high, usually much smaller, with an erect or more or less inclining simple or branched trunk slightly swollen at base, and rarely more than 6′ in diameter; sometimes with numerous clustered stems. Bark near the base of the trunk thick, rough, dark brown, marked above by scars left by falling leaves.

Distribution. Sand dunes of the coast from North Carolina to eastern Louisiana; west of the Apalachicola River attaining its largest size and sometimes ranging inland through Pine-forests for thirty or forty miles; and in Yucatan (var. yucatana Trel.).

A common garden plant in all countries with a temperate climate, and long naturalized in the southern states far beyond the limits of its natural range, in some of the West Indian islands and on the Gulf coast of Mexico. Forms with leaves variously striped with white, yellow, and red or with recurving leaves are frequent in cultivation.

2. Yucca Treculeana Carr. Spanish Bayonet. Spanish Dagger.

Leaves 2½°—4° long, 2′—3¼′ wide, slightly or not at all contracted above the dark red lustrous base, concave, stiff, rigid, dark blue-green, rough on the lower surface, nearly smooth on the upper, with a short stout dark red-brown tip, and dark brown margins roughened by minute deciduous teeth and ultimately separating into slender dark fibres; persistent for many years, the dead leaves hanging closely appressed against the trunk below the terminal crown of closely imbricated living leaves. Flowers in March and April on slender pedicels, in dense many-flowered glabrous or puberulous panicles 2°—4° long and raised on short stout stalks; perianth 1′—2′ long, 2′—4′ in diameter when fully expanded, with narrow elongated ovate-lanceolate to ovate segments, ¼′ wide, acute, thin and delicate, furnished at apex with a conspicuous tuft of short pale hairs; filaments slightly papillose, about as long as the prismatic ovary gradually narrowed above and crowned by the deeply divided stigmatic lobes. Fruit ripening in the summer, 3′—4′ long, about 1′ thick, dark reddish brown or ultimately black, with thin succulent sweetish flesh; seeds about ⅛′ wide, nearly ¹/₁₆′ thick, with a narrow border to the rim.

A tree, occasionally 25°—30° high, with a trunk sometimes 2° in diameter and numerous stout wide-spreading branches; usually smaller and often forming broad low thickets 4°—5° [113] tall. Bark on old trunks ¼′—½′ thick, dark red-brown and broken into thin oblong plates covered by small irregular closely appressed scales. Wood light brown, fibrous, spongy, heavy, difficult to cut and work.

Distribution. Shores of Matagorda Bay, southward through western Texas into Nuovo Leon, and through the valley of the Rio Grande to the eastern base of the mountains of western Texas; forming open stunted forests on the coast dunes at the mouth of the Rio Grande; farther from the coast often spreading into great impenetrable thickets.

Cultivated as an ornamental plant in the gardens of central and western Texas and in other southern States, and occasionally in those of southern Europe.

3. Yucca macrocarpa Coville. Spanish Dagger.

Leaves 1½°—2° long, 1′—2′ wide, gradually narrowed from the dark red lustrous base to above the middle, rigid, concave, yellow-green, rough on the lower surface and frequently also on the upper surface, with a stout elongated dark tip, and thickened margins separated into stout gray filaments. Flowers in March and April in densely flowered sessile or short-stalked glabrous or occasionally pubescent panicles; perianth usually about 2′ long, with acuminate segments, those of the outer and inner rows nearly of the same size; stamens shorter than the elongated style. Fruit 3′—4′ long, about 1½′ thick, abruptly contracted at apex into a stout point, nearly black when fully ripe, with sweet succulent flesh; seeds about ⅓′ wide, ⅛′ thick, with a narrow border to the rim.

A tree, rarely exceeding 15° in height, with a usually simple stem 6′—8′ in diameter, and often clothed to the ground with living leaves. Bark dark brown and scaly.

Distribution. Arid plains from western Texas to eastern Arizona and southward in Chihuahua.

4. Yucca mohavensis Sarg. Spanish Dagger.

Leaves 18′—20′ long, about 1½′ wide, abruptly contracted above the dark red lustrous base, gradually narrowed upward to above the middle, thin and concave except toward the slightly thickened base of the blade, dark green, smooth on both surfaces, with a stout rigid sharp-pointed tip, and entire bright red-brown margins soon separating into numerous pale filaments. Flowers from March to May on slender erect ultimately drooping pedicels 1′—1½′ long, in densely flowered sessile or short-stemmed panicles 12′—18′ in length; perianth 1′—2′ long, the segments united at the base into a short tube, thickened and hood-shaped at the apex, those of the outer rank often deeply flushed with purple, but little longer than the less prominently ribbed usually wider and thinner segments of the inner rank; stamens [114] with more or less pilose filaments nearly as long as the short style. Fruit ripening in August and September, 3′—4′ long, about 1½′ thick, usually much constricted near the middle, abruptly contracted at apex into a short stout point, dark dull brown or nearly black, with flesh often nearly ½′ thick; seeds ⅓′ wide, rather less than ⅛′ thick, with a narrow border to the rim.

A tree, rarely exceeding 15° in height, with a trunk usually simple or occasionally furnished with short spreading branches, and 6′—8′ in diameter, usually surrounded by a cluster of shorter more or less spreading stems and often clothed to the ground with living leaves. Bark dark brown and scaly. Wood soft, spongy, light brown.

Distribution. Southern Nevada and northwestern Arizona across the Mohave Desert to the California coast, extending northward to the neighborhood of Monterey, California, and southward into northern Lower California; common and attaining its largest size on the Mohave Desert, and sometimes ascending arid mountain slopes to altitudes of 4000° above the sea.

5. Yucca Schottii Engelm. Spanish Dagger.

Leaves 2½°—3° long, about 1½′ wide, gradually narrowed upward from the comparatively thin lustrous red base to above the middle, flat except toward the apex, smooth, light [115] yellow-green, with a long rigid sharp light red tip, and thick entire red-brown margins finally separating into short thin brittle threads. Flowers from July to September in erect stalked tomentose panicles; perianth 1′—1¾′ long, the broad oval or oblong-obovate thin segments pubescent on the outer surface toward the base and furnished at the apex with conspicuous clusters of white tomentum; stamens about two thirds as long as the ovary, with filaments pilose at the base, and only slightly enlarged at the apex. Fruit ripening in October and November, obscurely angled, 3½′—4′ long, about 1¼′ thick, often narrowed above the middle, with a stout thick point, and thin succulent flesh; seeds ¼′ wide, about ⅛′ thick, with a thin conspicuous marginal rim.

A tree, in Arizona rarely 18°—20° high, with a trunk often crooked or slightly inclining and simple or furnished with 2 or 3 short erect branches, covered below with dark brown scaly bark, roughened for many years by persistent scars of fallen leaves, and clothed above by the pendant dead leaves of many seasons.

Distribution. Dry slopes of the mountain ranges of Arizona near the Mexican boundary usually at altitudes between 5000° and 6000°, and southward into Sonora.

6. Yucca Faxoniana Sarg. Spanish Dagger.

Leaves 2½°—4° long, 2½′—3′ wide, abruptly contracted above the conspicuously thickened lustrous base, widest above the middle, flat on the upper surface, thickened and rounded on the lower surface toward the base, rigid, smooth and clear dark green, with a short stout dark tip, and brown entire margins breaking into numerous stout gray or brown fibres short and spreading near the apex of the leaf, longer, more remote, and forming a thick cobweb-like mass at their base. Flowers appearing in April on thin drooping pedicels, in dense many-flowered glabrous panicles 3°—4° long, with elongated pendulous branches; perianth 2½′ long, the segments thin, concave, widest above the middle, narrowed at the ends, united at base into a short tube, those of the outer rank being about half as wide as those of the inner rank and two thirds as long; stamens much shorter than the ovary, with slender filaments pilose above the middle and abruptly dilated at apex; ovary conspicuously ridged, light yellow marked with large pale raised lenticels, and gradually narrowed into an elongated slender style. Fruit ripening in early summer, slightly or not at all angled, abruptly contracted at apex into a long or short hooked beak, 3′—4′ long, 1′—1½′ thick, light orange-colored and lustrous when first ripe, becoming nearly black, with thick succulent bitter-sweet flesh; seeds ¼′ long, about ⅛′ thick, with a narrow nearly obsolete margin to the rim.

A tree, often 40° high, with a trunk sometimes 2° in diameter above the broad abruptly [116] enlarged base, unbranched or divided into several short branches, and covered above by a thick thatch of the pendant dead leaves of many seasons; frequently smaller and until ten or twelve years old clothed from the ground with erect living leaves. Bark near the base of old trees dark reddish brown, ⅓′—½′ thick, broken on the surface into small thin loose scales.

7. Yucca brevifolia Engelm. Joshua Tree.

Leaves 5′—8′ or on young plants rarely 10′—12′ long, ¼′—½′ wide, rigid, crowded in dense clusters, lanceolate, gradually tapering from the bright red-brown lustrous base, bluish green and glaucous, smooth or slightly roughened, concave above the middle, with a sharp dark brown tip, and thin yellow margins armed with sharp minute teeth; persistent for many years. Flowers appearing from March until the beginning of May, the creamy white closely imbricated bracts of the nearly sessile pubescent panicle forming before its appearance a conspicuous cone-like bud 8′ or 10′ long; perianth globose to oblong, 1′—2′ long, greenish white, waxy, dull or lustrous, its segments slightly united at the base, keeled on the back, thin below the middle, gradually thickened upward into the concave incurved rounded tip, those of the outer rank rather broader, thicker, and more prominently keeled than those of the inner rank, glabrous or pubescent; stamens about half as long as the ovary, with filaments villose-papillate from the base; ovary conic, 3-lobed above the middle, bright green, with narrow slightly developed septal nectar-glands, and a sessile nearly equally 6-lobed stigma. Fruit ripening in May or June, spreading or more or less pendant at maturity, oblong-ovoid, acute, slightly 3-angled, 2′—4′ long, 1½′—2′ thick, light red or yellow-brown, the outer coat becoming dry and spongy at maturity; seeds nearly ½′ long, rather less than ¹/₁₆′ thick, with a broad well-developed margin to the rim, and a large conspicuous hilum.

A tree, 30°—40° high, with a trunk 2°—3° in diameter, rising abruptly from a broad thick basal disk, thick tough roots descending deeply into the soil, and stout branches spreading into a broad, often symmetrical head formed by the continued forking of the branches at the base of the terminal flower-clusters; the stem until 8°—10° high simple and clothed to the ground with leaves erect until after the appearance of the first flowers, then spreading at right angles and finally becoming reflexed. Bark 1′—1½′ thick, deeply divided into oblong plates frequently 2° long. Wood light, soft, spongy, difficult to work, light brown or nearly white; sometimes cut into thin layers and used as wrapping material or manufactured into boxes and other small articles. The seeds are gathered and eaten by Indians.

Distribution. Southwestern Utah to the western and northern rim of the Mohave Desert in California; most abundant and of its largest size on the foothills on the desert slope of the Tehachapi Mountains, California.

8. Yucca gloriosa L. Spanish Dagger.

Leaves 2°—2½° long, gradually narrowed above the broad base and then gradually broadened to above the middle, thin, flat or slightly concave toward the apex, frequently longitudinally folded, dull often glaucous green, roughened on the under surface especially above the middle, with a stout dark red tip, and pale margins serrulate toward the base of the leaf, with minute early deciduous teeth, or occasionally separating into thin fibres. Flowers in October, in pubescent or glabrate panicles, 2°—4° long, on stout stalks sometimes 3°—4° in length, their large creamy white bracts forming before the panicle emerges a conspicuous egg-shaped bud 4′—6′ long; perianth when fully expanded 3½′—4′ across, its segments thin, ovate, acute, or lance-ovate, often tinged with green or purple, slightly united at the base, pubescent at apex; stamens about as long as the ovary, with hispid or slightly papillose filaments and deeply emarginate anthers; ovary slightly lobed, 6-sided, light green, gradually narrowed into the elongated spreading stigmatic lobes. Fruit very rarely produced, prominently 6-ridged, pendulous, 3′ long, 1′ in diameter, cuspidate, raised on a short stout stipe, with a thin leathery almost black outer coat; seeds ¼′ wide and about ¹/₃₆′ thick, with a smooth coat and a narrow marginal rim.

A tree, with a trunk occasionally 6°—8° high and 4′—6′ in diameter, simple or rarely furnished with a few short branches and usually clothed to the base with pendant dead leaves; in cultivation often becoming much larger, with a stout trunk covered with smooth light gray bark, and erect or in one form (var. recurvifolia Engelm.) pendulous leaves.

Distribution. Sand dunes and the borders of beaches of the seacoast from North Carolina to northern Florida.

Often cultivated with many forms in the gardens and pleasure-grounds of all temperate countries.

9. Yucca elata Engelm. Spanish Dagger.

Leaves 20′—30′ long, ¼′—½′ wide, rigid, gradually narrowed from the thin base, tapering toward the apex, or sometimes somewhat broadest at the middle, thin, flat on the upper surface, slightly thickened and rounded on the lower surface toward the base, smooth, pale [118] yellow-green, with a slender stiff red-brown tip, and thickened entire pale margins soon splitting into long slender filaments. Flowers in May and June on slender spreading more or less recurved pedicels, in glabrous much-branched panicles 4°—6° long, raised on stout naked stem 3°—7° in length; perianth ovoid and acute in the bud, when fully expanded 3½′—4′ across, its segments united at the base into a short slender distinct tube, ovate or slightly obovate, those of the outer rank usually acute, not more than half as broad as those of the inner rank; stamens as long or a little longer than the ovary, with slender nearly terete filaments; ovary sessile, almost terete, pale green, abruptly contracted into the stout elongated style. Fruit an erect oblong capsule rounded and obtuse at the ends, tipped by a short stout mucro, conspicuously 3-ribbed, with rounded ridges on the back of the carpels, 1½′—2′ long, 1′—1½′ wide, with a thin firm light brown ligneous outer coat closely adherent to the lustrous light yellow inner coat, in ripening splitting from the top to the bottom between the carpels, and through their backs at the apex; seeds ⅓′ wide and about ¹/₃₂′ thick, with a smooth coat and a thin brittle wide margin to the rim.

A tree, with a rough much-branched underground stem penetrating deep into the soil and a trunk often 15°—20° high and 7′—8′ in diameter, covered above with a thick thatch of the pendant dead leaves of many years, simple, or branched at the top with a few short stout branches densely covered with leaves at first erect, then spreading nearly at right angles, and finally pendulous. Bark dark brown, irregularly fissured, broken into thin plates, about ¼′ thick. Wood light, soft, spongy, pale brown or yellow.

Distribution. High desert plateaus from southwestern Texas to southern Arizona; southward into northern Mexico; most abundant and of its largest size on the eastern slope of the continental divide in southern New Mexico and along the northern rim of the Tucson Desert in Arizona.

Division II. Dicotyledons.

Stems formed of bark, wood, or pith, and increasing by the addition of an annual layer of wood inside the bark. Parts of the flower mostly in 4’s and 5’s; embryo with a pair of opposite cotyledons. Leaves netted-veined.

Subdivision 1. Apetalæ. Flowers without a corolla and sometimes without a calyx (with a corolla in Olacaceæ).

Section 1. Flowers in unisexual aments (female flowers of Juglans and Quercus solitary or in spikes); ovary inferior (superior in Leitneriaceæ) when calyx is present.

V. SALICACEÆ.

Trees or shrubs, with watery juice, alternate simple stalked deciduous leaves with stipules, soft light usually pale wood, astringent bark, scaly buds, and often stoloniferous roots. Flowers appearing in early spring usually before the leaves, solitary in the axils of the scales of unisexual aments from buds in the axils of leaves of the previous year, the male and female on different plants; perianth 0; stamens 1, 2 or many, their anthers introrse, 2-celled, the cells opening longitudinally; styles usually short or none; stigmas 2—4, often 2-lobed. Fruit a 1-celled 2—4-valved capsule, with 2—4 placentas bearing below their middle numerous ascending anatropous seeds without albumen and surrounded by tufts of long white silky hairs attached to the short stalks of the seeds and deciduous with them; embryo straight, filling the cavity of the seed; cotyledons flattened, much longer than the short radicle turned toward the minute hilum.

The two genera of this family are widely scattered but most abundant in the northern hemisphere, with many species, and are often conspicuous features of vegetation.

CONSPECTUS OF THE GENERA.

1. POPULUS L. Poplar.

Large fast-growing trees, with pale furrowed bark, terete or angled branchlets, resinous winter-buds covered by several thin scales, those of the first pair small and opposite, the others imbricated, increasing in size from below upward, accrescent and marking the base of the branchlet with persistent ring-like scars, and thick roots. Leaves involute in the bud, usually ovate or ovate-lanceolate, entire, dentate with usually glandular teeth, or lobed, penniveined, turning yellow in the autumn; petioles long, often laterally compressed, sometimes furnished at the apex on the upper side with 2 nectariferous glands, leaving in falling oblong often obcordate, elliptic, arcuate, or shield-shaped leaf-scars displaying the ends of 3 nearly equidistant fibro-vascular bundles; stipules caducous, those of the first leaves resembling the bud-scales, smaller higher on the branch, and linear-lanceolate and scarious on the last leaves. Flowers in pendulous stalked aments, the pistillate lengthening and rarely becoming erect before maturity; scales obovate, gradually narrowed into slender stipes, dilated and lobed, palmately cleft or fimbriate at apex, membranaceous, glabrous or villose, more crowded on the staminate than on the pistillate ament, usually caducous; disk of the flower broadly cup-shaped, often oblique, entire, dentate or irregularly lobed, fleshy or membranaceous, stipitate, usually persistent under the fruit; stamens 4—12 or 12—60 or more, inserted on the disk, their filaments free, short, light yellow; anthers ovoid or oblong, purple or red; ovary sessile in the bottom of the disk, oblong-conical subglobose or ovoid-oblong, cylindric or slightly lobed, with 2 or 3 or rarely 4 placentas; styles usually short; stigmas as many as the placentas, divided into filiform lobes or broad, dilated, 2-parted or lobed. Fruit ripening before the full growth of the leaves, greenish, reddish brown, or buff color, oblong-conic, subglobose or ovoid-oblong, separating at maturity into 2—4 recurved valves. Seeds broadly obovoid or ovoid, rounded or acute at the apex, light chestnut-brown; cotyledons elliptic.

Populus in the extreme north often forms great forests, and is common on the alluvial bottom-lands of streams and on high mountain slopes, ranging from the Arctic Circle to northern Mexico and Lower California and from the Atlantic to the Pacific in the New World, and to northern Africa, the southern slopes of the Himalayas, central China, and [120] Japan in the Old World. Of the thirty-four species now generally recognized fifteen are found in North America. The wood of many of the American species is employed in large quantities for paper-making, and several species furnish wood used in construction and in the manufacture of small articles of wooden ware. The bark contains tannic acid and is used in tanning leather and occasionally as a tonic, and the fragrant balsam contained in the buds of some species is occasionally used in medicine. The rapidity of their growth, their hardiness and the ease with which they can be propagated by cuttings, make many of the species useful as ornamental trees or in wind-breaks, although planted trees often suffer severely from the attacks of insects boring into the trunks and branches. Of the exotic species, the Abele, or White Poplar, Populus alba L., of Europe and western Asia, and its fastigiate form, and the so-called Lombardy Poplar, a tree of pyramidal habit and a form of the European and Asiatic Populus nigra L., and one of its hybrids, have been largely planted in the United States.

CONSPECTUS OF THE NORTH AMERICAN SPECIES.

1. Populus tremuloides Michx. Aspen. Quaking Asp.

Leaves ovate to broad-ovate or rarely reniform (var. reniformis Tidestrom) abruptly short-pointed or acuminate at apex rounded or rarely cuneate at the wide base, closely crenately serrate with glandular teeth, thin, green and lustrous above, dull green or rarely pale below, up to 4½′ long and broad with a prominent midrib, slender primary veins and conspicuous reticulate veinlets; petioles slender, compressed laterally, 1½′—3′ long. Flowers: aments 1½′—2½′ long, the pistillate becoming 4′ in length at maturity; scales deeply divided into 3—5 linear acute lobes fringed with long soft gray hairs; disk oblique, the staminate entire, the pistillate slightly crenate; stamens 6—12; ovary conic, with a short thick style and erect stigmas thickened and club-shaped below and divided into linear diverging lobes. Fruit maturing in May and June, oblong-conic, light green, thin-walled, nearly ¼′ long; seeds obovoid, light brown, about ¹/₃₂′ in length.

A tree, 20°—40° high, with a trunk 18′—20′ in diameter, slender remote and often contorted branches somewhat pendulous toward the ends, forming a narrow symmetrical round-topped head, and slender branchlets covered with scattered oblong orange-colored lenticels, bright red-brown and very lustrous during their first season, gradually turning light gray tinged with red, ultimately dark gray, and much roughened for two or three years by the elevated leaf-scars. Winter-buds slightly resinous, conic, acute, often incurved, about ¼′ long, narrower than the more obtuse flower-buds, with 6 or 7 lustrous glabrous red-brown scales scarious on the margins. Bark thin, pale yellow-brown or orange-green, often roughened by horizontal bands of circular wart-like excrescences, frequently marked below the branches by large rows of lunate dark scars. Wood light brown, with nearly white sapwood of 25—30 layers of annual growth.

Distribution. Southern Labrador to the southern shores of Hudson’s Bay and northwesterly to the mouth of the Mackenzie River, through the northern states to the mountains of Pennsylvania, northern Ohio, Indiana and Illinois, eastern and central Iowa and northeastern Missouri; common and generally distributed usually on moist sandy soil and gravelly hillsides; most valuable in the power of its seeds to germinate quickly in soil made infertile by fire and of its seedlings to grow rapidly in exposed situations; westward passing into the var. aurea Daniels, with thicker rhombic to semiorbicular or broad-ovate generally smaller leaves, usually pale on the lower surface, rounded or acute and minutely short-pointed at apex, rounded or cuneate at base, often entire with slightly [122] thickened margins, or occasionally coarsely crenately serrate, with inconspicuous reticulate veinlets, turning bright golden yellow in the autumn before falling.

A tree occasionally 100° high with a trunk up to 3° in diameter, with pale often white bark, becoming near the base of old stems 2′ thick, nearly black, and deeply divided into broad flat ridges broken on the surface into small appressed plate-like scales.

Distribution. Valley of the Yukon River to Saskatchewan, and southward through the mountain ranges of the Rocky Mountain region to southern New Mexico, the San Francisco Mountains of Arizona, and westward to the valley of the Skeena River, British Columbia, western Washington and Oregon, the western slopes of the Sierra Nevada and the high mountains of southern California, and eastward to North and South Dakota and western Nebraska; on the mountains of Chihuahua, and on the Sierra de Laguna, Lower California.

Populus tremuloides var. vancouveriana Sarg.

Leaves broadly ovate to semiorbicular, abruptly short-pointed or rounded at apex, rounded or slightly cordate at the broad base, coarsely crenately serrate and sometimes obscurely crispate on the margins, when they unfold covered below and on the petioles with a thick coat of long matted pale hairs, and slightly villose, glabrous or nearly glabrous above, soon glabrous, and at maturity thick dark green, lustrous and scabrate on the upper surface, paler on the lower surface, 3′—4½′ long and broad, with a prominent midrib and primary veins; petioles slender, compressed, becoming glabrous, 2′—3′ in length. Flowers: staminate aments slightly villose; pedicels pubescent; disk of the flower puberulous toward the base; flowers as in the species; pistillate aments 2′—2¼′ long, becoming 3′—3½′ in length at maturity; the rachis, pedicels and slightly lobed disk of the flower densely villose-pubescent; ovary conic, pubescent, with a short style and stigma divided into narrow divergent lobes. Fruit on pedicel not more than ¹/₂₄′ in length, oblong-conic, pubescent or glabrous, ¼′ long.

A tree 30°—36° high, with a trunk 12′—16′ in diameter, stout spreading branches forming a round-topped head, stout, reddish brown pubescent or puberulous branchlets often becoming glabrous during their first summer. Winter-buds acute, tomentose, pubescent or glabrous.

Coast of Vancouver Island, British Columbia and shores of Puget Sound; Tualitin, Washington County, and valley of the Willamette River at Corvallis, Benton County, Oregon.

2. Populus grandidentata Michex. Poplar.

Leaves semiorbicular to broad-ovate, short-pointed at apex, rounded, abruptly cuneate or rarely truncate at the broad entire base, coarsely repand-dentate above with few stout incurved teeth, covered like the petioles early in the season with white tomentum, soon glabrous, thin and firm in texture, dark green above, paler on the lower surface, 2′—3′ long, 2′—2½′ wide, with a prominent yellow midrib, conspicuously forked veins, and reticulate veinlets; petioles slender, laterally compressed, 1½′—2½′ long. Flowers: aments pubescent, 1½′—2½′ long, the pistillate becoming 4′—5′ long at maturity; scales pale and scarious below, divided above into 5 or 6 small irregular acute lobes covered with soft pale hairs; disk shallow, oblique, the staminate entire, the pistillate slightly crenate; stamens 6—12, with short slender filaments and light red anthers; ovary oblong-conic, bright green, puberulous, with a short style, and spreading stigmas divided nearly to the base into elongated filiform lobes. Fruit ripening before the leaves are fully grown, often more or less curved above the middle, light green and puberulous, thin-walled, 2-valved, about ⅛′ long; pedicel slender, pubescent, about ¹/₁₂′ in length; seeds minute dark brown.

A tree, often 60°—70° high, with a trunk occasionally 2° in diameter, slender rather rigid branches forming a narrow round-topped head, and stout branchlets marked by scattered oblong orange-colored lenticels, coated when they first appear with thick hoary deciduous tomentum, becoming during their first year dark red-brown or dark orange-colored, glabrous, lustrous, or covered with a delicate gray pubescence, and in their second year dark gray sometimes slightly tinged with green and much roughened by the elevated 3-lobed leaf-scars; generally smaller, and usually not more than 30°—40° tall. Winter-buds terete, broadly ovoid, acute, with light bright chestnut-brown scales, pubescent during the winter especially on their thin scarious margins, about ⅛′ long and not more than half the size of the flower-buds. Bark thin, smooth, light gray tinged with green, becoming near the base of old trunks ¾′—1′ thick, dark brown tinged with red, irregularly fissured and divided into broad flat ridges roughened on the surface by small thick closely appressed scales. Wood light brown, with thin nearly white sapwood of 20—30 layers of annual growth.

Distribution. Rich moist sandy soil near the borders of swamps and streams; Nova Scotia, through New Brunswick, southern Quebec and Ontario to northern Minnesota, [124] southward through the northern states to Pennsylvania, northern Ohio, and eastern (Muscatine County) and central Iowa, and westward to central Kentucky and Tennessee; passing into the var. meridionalis Tidestrom with broad-ovate acuminate leaves with more numerous teeth, often 4′—5′ long and 3′ wide; the common form in Maryland, northern Delaware, the piedmont region of Virginia and North Carolina, southern Ohio, and southern Indiana and Illinois; rare northward to northern New England.

3. Populus heterophylla L. Swamp Cottonwood. Black Cottonwood.

Leaves broadly ovate, gradually narrowed and acute, short-pointed or rounded at apex, slightly cordate or truncate or rounded at the wide base, usually furnished with a narrow deep sinus, finely or coarsely crenately serrate with incurved glandular teeth, covered as they unfold with thick hoary deciduous tomentum, becoming thin and firm in texture, dark deep green above, pale and glabrous below, with a stout yellow midrib, forked veins and conspicuous reticulate veinlets, 4′—7′ long, 3′—6′ wide; petioles slender terete tomentose or nearly glabrous 2½′—3½′ in length. Flowers: staminate aments broad, densely flowered, 1′ long, erect when the flowers first open, becoming pendulous and 2′—2½′ long; scales narrowly oblong-obovate, brown, scarious and glabrous below, divided into numerous elongated filiform light red-brown lobes; disk oblique, slightly concave; stamens 12—20, with slender filaments about as long as the large dark red anthers; pistillate aments slender, pendulous, few-flowered, 1′—2′ long, becoming erect and 4′—6′ long before maturing, their scales concave and infolding the flowers, linear-obovate, brown and scarious, laterally lobed, fimbriate above the middle, caducous; disk thin, irregularly divided in numerous triangular acute teeth, long-stalked; ovary ovoid, terete or obtusely 3-angled, with a short stout elongated style and 2 or 3 much-thickened dilated 2 or 3-lobed stigmas. Fruit on elongated pedicels, ripening when the leaves are about one third grown, ovoid, acute, dark red-brown, rather thick-walled, 2 or 3-valved, about ½′ long; seeds obovoid, minute, dark red-brown.

A tree, 80°—90° high, with a tall trunk 2°—3° in diameter, short rather slender branches forming a comparatively narrow round-topped head, and stout branchlets, marked by small elongated pale lenticels, coated at first with hoary caducous tomentum, becoming dark brown and rather lustrous or ashy gray, or rarely pale orange color and glabrous or slightly puberulous, or covered with a glaucous bloom in their first winter, growing darker in their second year and much roughened by the large thickened leaf-scars; usually much smaller and at the north rarely more than 40° tall. Winter-buds slightly resinous, broadly ovoid, acute, with bright red-brown scales, about ¼′ long and about one half the size of the [125] flower-buds. Bark on young trunks divided by shallow fissures into broad flat ridges separating on the surface into thick plate-like scales, becoming on old trunks ¾′—1′ thick, light brown tinged with red, and broken into long narrow plates attached only at the middle and sometimes persistent for many years. Wood dull brown, with thin lighter brown sapwood of 12—15 layers of annual growth; now often manufactured into lumber in the valley of the Mississippi River and in the Gulf states, and as black poplar used in the interior finish of buildings.

Distribution. Southington, Connecticut, and Northport, Long Island, southward near the coast to southern Georgia, and the valley of the lower Apalachicola River, Florida, through the Gulf states to western Louisiana, and through Arkansas to southeastern Missouri, western Kentucky and Tennessee, southern Illinois and Indiana, and in central and northern Ohio (Williams, Ottawa and Lake Counties); in the north Atlantic states in low wet swamps, rare and local; more common south and west on the borders of river swamps; very abundant and of its largest size in the valley of the lower Ohio and in southeastern Missouri, eastern Arkansas, and western Mississippi.

4. Populus tacamahacca Mill. Balsam. Tacamahac.

Leaves ovate-lanceolate, gradually narrowed and acuminate at apex, cordate or rounded at base, or narrow-elliptic and acute or acuminate at the ends, finely crenately serrate, with slightly thickened revolute margins, coated when they unfold with the gummy secretions of the bud, glabrous, or puberulous on the under side of the midrib, becoming thin and firm in texture, deep dark green and lustrous above, pale green or glaucous and more or less rusty and conspicuously reticulate-venulose below, 3′—5′ long, 1½′—3′ wide, with thin veins running obliquely almost to the margins; petioles slender, terete, 1½′ long, glabrous or rarely puberulous. Flowers: aments long-stalked, the pistillate becoming 4′—5′ long before the fruit ripens, glabrous or pubescent; scales broadly obovate, light brown and scarious, often irregularly 3-parted at apex, cut into short thread-like brown lobes; disk of the staminate flower oblique, short-stalked; stamens 20—30, with short filaments and large light red anthers; disk of the pistillate flower cup-shaped; ovary ovoid, slightly 2-lobed, with two nearly sessile large oblique dilated crenulate stigmas. Fruit ovoid-oblong, acute and often curved at apex, 2-valved, light brown, about ¼′—⅓′ long, nearly sessile or short-stalked, ½′—⅛′ in length; seeds oblong-obovoid, pointed at apex, narrowed and truncate at base, light brown, about ¹/₁₂′ long.

A tree, often 100° high, with a tall trunk 6°—7° in diameter, stout erect branches usually [126] more or less contorted near the end, forming a comparatively narrow open head, and glabrous or occasionally pubescent branchlets marked by oblong bright orange-colored lenticels, much roughened by the thickened leaf-scars, at first red-brown and glabrous or pubescent, becoming bright and lustrous in their first winter, dark orange-colored in their second year, and finally gray tinged with yellow-green; usually much smaller toward the southern limits of its range. Winter-buds saturated with a yellow balsamic sticky exudation, ovoid, terete, long-pointed; terminal 1′ long, ⅓′ broad; axillary about ¾′ long, ¹/₁₆′ broad, with 5 oblong pointed concave closely imbricated thick chestnut-brown lustrous scales. Bark light brown tinged with red, smooth or roughened by dark excrescences, becoming on old trunks ¾′—1′ thick, gray tinged with red, and divided into broad rounded ridges covered by small closely appressed scales. Wood light brown, with thick nearly white sapwood.

Distribution. Low often inundated river-bottom lands and swamp borders; Labrador to latitude 65° north in the valley of the Mackenzie River, and to the Alaskan coast, south to northern New England and New York, central Michigan, Minnesota (except in southern and southwestern counties), Turtle Mountains, Rolette County, North Dakota, the Black Hills of South Dakota, northwestern Nebraska (basin of Hat Creek), and in Colorado; the characteristic tree on the streams of the prairie region of British America, attaining its greatest size on the islands and banks of the Peace, Athabasca, and other tributaries of the Mackenzie; common in all the region near the northern boundary of the United States from Maine to the western limits of the Atlantic forests; the largest of the sub-Arctic American trees, and in the far north the most conspicuous feature of vegetation; passing into the variety Michauxii Farwell, with more cordate leaves, slightly pilose on the under side of the midrib and veins; common from Aroostook County, Maine, to the Province of Quebec, Newfoundland, and the shores of Hudson Bay.

Populus candicans Ait., the Balm of Gilead of which only the pistillate tree is known, has often been considered a variety of the North American Balsam Poplar. This tree has been long cultivated in the northeastern part of the country and has sometimes escaped from cultivation and formed groves of considerable extent, as on the banks of Cullasagee Creek on the western slope of the Blue Ridge in Macon County, North Carolina. The fact that only one sex is known suggests hybrid origin but of obscure and possibly partly of foreign origin.

5. Populus trichocarpa Hook. Black Cottonwood. Balsam Cottonwood.

Leaves broad-ovate, acute or acuminate at apex, rounded or abruptly cuneate at base, finely crenately serrate, glabrous, dark green above, pale and rusty or silvery white and conspicuously reticulate-venulose below, 3′—4′ long, 2′—2½′ wide; petioles slender, pubescent, puberulous, pilose or rarely glabrous, 1½′—2′ in length. Flowers: aments stalked, villose-pubescent, the staminate densely flowered, 1½′—2′ long, ⅓′ thick, the pistillate loosely flowered, 2½′—3′ long, becoming 4′—5′ long before the fruit ripens; scales dilated at the apex, irregularly cut into numerous filiform lobes, glabrous or slightly puberulous on the outer surface; disk of the staminate flower broad, slightly oblique; stamens 40—60, with slender elongated filaments longer than the large light purple anthers; disk of the pistillate flower deep cup-shaped, with irregularly crenate or nearly entire revolute margins; ovary subglobose, coated with thick hoary tomentum, with 3 nearly sessile broadly dilated deeply lobed stigmas. Fruit subglobose, nearly sessile, pubescent, thick-walled, 3-valved; seeds obovoid, apiculate at the gradually narrowed apex, light brown, puberulous toward the ends, ¹/₁₂′ long.

A tree, 30°—100° high, with a trunk 1°—3° in diameter, erect branches forming an open head, and slender branchlets terete or slightly angled while young, marked by many orange-colored lenticels, glabrous or when they first appear coated with deciduous rufous or pale pubescence, reddish brown during their first year, gradually becoming dark gray, and roughened by the greatly enlarged and thickened elevated leaf-scars. Winter-buds resinous, [127] fragrant, ovoid, long-pointed, frequently curved above the middle, ¾′ long and ¼′ thick, with 6 or 7 light orange-brown slightly puberulous scales scarious on the margins. Bark ½′—2½′ thick, ashy gray, deeply divided into broad rounded ridges broken on the surface into thick closely appressed scales. Wood light, dull brown, with thin nearly white sapwood.

Distribution. In California in small groves with widely scattered individuals on the coast ranges, the western slope of the Sierra Nevada up to elevations of 6000°—8000°, and on the southern mountains to Mt. Palomar in San Diego County; on the California islands, and on the western slopes of the San Pedro Mártir Mountains, Lower California.

On the high Sierra Nevada and in northern California passing into the var. hastata A. Henry, differing in its thicker leaves, usually longer in proportion to their width, often long-acuminate, rounded or cordate at base, frequently 5′ or 6′ long and 3′ or 4′ wide, with glabrous petioles and larger sometimes nearly glabrous capsules on glabrous or pubescent aments, sometimes 10′—12′ in length, and in its glabrous young branchlets.

A tree sometimes 200° high, with a trunk 7°—8° in diameter, and the largest deciduous-leaved tree of northwestern North America. The wood is largely used in Oregon and Washington for the staves of sugar barrels and in the manufacture of wooden ware.

Distribution. In open groves on rich bottom lands of streams from Siskiyou County, California, to southern Alaska; eastward in the United States through Oregon and Washington to western and southern Idaho; and to the mountains of western Nevada; in British Columbia to the valley of the Columbia River; on the banks of the east fork of the Kaweah River, Tulare County, California, at 10,000° above the sea.

6. Populus angustifolia James. Narrow-leaved Cottonwood.

Leaves lanceolate, ovate-lanceolate, elliptic or rarely obovate, narrowed to the tapering acute or rounded apex, gradually narrowed and cuneate or rounded at base, finely or on vigorous shoots coarsely serrate, thin and firm, bright yellow-green above, glabrous or rarely puberulous and paler below, 2′—3′ long, ½′—1′ wide, or on vigorous shoots occasionally 6′—7′ long, and 1½′ wide, with a stout yellow midrib and numerous slender-oblique primary veins arcuate and often united near the slightly thickened revolute margins; petioles slender, somewhat flattened on the upper side, and in falling leaving small nearly oval obcordate scars. Flowers: aments densely flowered, glabrous, short-stalked, ½′—2½′ long, the pistillate becoming 2½′—4′ long before the fruit ripens; scales broadly obovate, glabrous, thin, scarious, light brown, deeply and irregularly cut into numerous [128] dark red-brown filiform lobes; disk of the staminate flower cup-shaped, slightly oblique, short-stalked; stamens 12—20, with short filaments and large light red anthers; disk of the pistillate flower shallow, cup-shaped, slightly and irregularly lobed, short-stalked; ovary ovoid, more or less 2-lobed, with a short or elongated style and 2 oblique dilated irregularly lobed stigmas. Fruit broadly ovoid, often rather abruptly contracted above the middle, short-pointed, thin-walled, 2-valved; pedicels often ⅓′ long; seeds ovoid or obovoid, rather obtuse, light brown, nearly ⅛′ long.

A tree, 50°—60° high, with a trunk rarely more than 18′ in diameter, slender erect branches forming a narrow and usually pyramidal head, and slender glabrous or rarely puberulous branchlets marked by pale lenticels, at first light yellow-green, becoming bright or dark orange color in their first season, pale yellow in their second winter, and ultimately ashy gray. Winter-buds very resinous, ovoid, long-pointed, covered by usually 5 thin concave chestnut-brown scales; terminal ¼′—½′ long and nearly twice as large as the axillary buds. Bark ¾′—1′ thick, light yellow-green, divided near the base of old trees by shallow fissures into broad flat ridges, smooth and much thinner above. Wood light brown, with thin nearly white sapwood of 10—30 layers of annual growth.

Distribution. Banks of streams usually at altitudes of 5000°—10,000° above the sea; southern Alberta to the Black Hills of South Dakota and northwestern Nebraska (basin of Hat Creek) westward through Wyoming, Montana and Idaho to Yakima County, Washington, and southward to central Nevada, southwestern New Mexico (Silver City, Grant County) and northern Arizona; the common Cottonwood of northern Colorado, Utah, Wyoming, southern Montana, and eastern Idaho; on the mountains of Chihuahua.

7. Populus acuminata Rydb. Cottonwood.

Leaves rhombic-lanceolate to ovate, abruptly acuminate, gradually or abruptly narrowed and cuneate or concave-cuneate, or rarely broad and rounded at the mostly entire base, coarsely crenately serrate except near the apex, dark green and lustrous above, dull green below, 2′—4′ long, ¾′—2′ wide, with a slender yellow midrib, thin remote primary veins and obscure reticulate veinlets; petioles slender, nearly terete, 1′—3′ long. Flowers: aments slender, short-stalked, 2′—3′ long, the pistillate becoming 4′ or 5′ long before the fruit ripens; scales scarious, light brown, glabrous, dilated and irregularly divided into filiform lobes; disk of the staminate flower wide, oblique, and membranaceous; stamens numerous, with short filaments and dark red anthers; disk of the pistillate flower deep cup-shaped; ovary broad-ovoid, gradually narrowed above, with large laciniately lobed nearly sessile stigmas. Fruit pedicellate, oblong-ovoid, acute, thin-walled, slightly pitted, [129] about ⅓′ long, 3 or rarely 2-valved; seeds oblong-obovoid, rounded at the apex, light brown, about ¹/₁₂′ in length.

A tree, usually about 40° high, with a trunk 12′—18′ in diameter, stout spreading and ascending branches forming a compact round-topped head, and slender terete or slightly 4-angled pale yellow-brown branchlets roughened for two or three years by the elevated oval horizontal leaf-scars. Winter-buds acuminate, resinous, about ⅓′ long, with 6 or 7 light chestnut-brown lustrous scales. Bark on young stems and large branches smooth, nearly white, becoming on old trunks pale gray-brown, about ½′ thick, deeply divided into broad flat ridges.

Distribution. Banks of streams in the arid eastern foothill region of the Rocky Mountains; Assiniboia to the Black Hills of South Dakota, northwestern Nebraska, eastern Wyoming, southern Colorado, and southwestern New Mexico (Fort Bayard, Grant County); in Colorado crossing the Continental Divide to southeastern Utah; passing into the var. Rehderi Sarg. differing in the larger leaves on longer petioles, and in the pubescent branchlets and winter-buds. Borders of streams southeastern New Mexico.

Sometimes planted as a shade-tree in the streets of cities in the Rocky Mountain region.

× Populus Andrewsii Sarg. intermediate in its character between P. acuminata and P. Sargentii and believed to be a natural hybrid of these species has been found growing naturally near Boulder and Walsenburg, Colorado, and as a street tree in Montrose, Colorado.

8. Populus Fremontii S. Wats. Cottonwood.

Leaves deltoid or reniform, generally contracted into broad short entire points, or rarely rounded or emarginate at apex, truncate, slightly cordate or abruptly cuneate at the entire base, coarsely and irregularly serrate, with few or many incurved gland-tipped teeth, coated like the petioles when they unfold with short spreading caducous pubescence, at maturity thick and firm, glabrous bright green and lustrous, 2′—2½′ long, 2½′—3′ wide, with a thin yellow midrib and 4 or 5 pairs of slender veins; petioles flattened, yellow, 1½′—3′ long. Flowers: staminate aments densely flowered, 1½′—2′ long, nearly ½′ thick, with slender glabrous stems, the pistillate sparsely flowered, with stout glabrous or puberulous stems, becoming before the fruit ripens 4′ or 5′ long; scales light brown, thin and scarious, dilated and irregularly cut at apex into filiform lobes; disk of the staminate flower broad, oblique, slightly thickened on the entire revolute margin; stamens 60 or more, with large dark red [130] anthers; disk of the pistillate flower cup-shaped; ovary ovoid or ovoid-oblong, with 3 or rarely 4 broad irregularly crenately lobed stigmas. Fruit ovoid, acute or obtuse, slightly pitted, thick-walled, 3 or rarely 4-valved, ⅓′—½′ long; pedicel stout, from ¹/₂₀′—⅙′ long; seeds ovoid, acute, light brown, nearly ⅛′ in length.

A tree, occasionally 100° high, with a short trunk 5°—6° in diameter, stout spreading branches pendulous at the ends and forming a broad rather open graceful head, and slender terete branchlets light green and glabrous, becoming light yellow before winter, dark or light gray more or less tinged with yellow in their second year, and only slightly roughened by the small 3-lobed leaf-scars. Winter-buds ovoid, acute, with light green lustrous [131] scales, the terminal usually about ⅓′ long and usually two or three times as large as the lateral buds. Bark on young stems light gray-brown, thin, smooth or slightly fissured, becoming on old trees 1½′—2′ thick, dark brown slightly tinged with red, and deeply and irregularly divided into broad connected rounded ridges covered with small closely appressed scales. Wood light brown, with thin nearly white sapwood.

Distribution. Banks of streams; valley of the upper Sacramento River southward through western California to the San Pedro Mártir Mountains, Lower California; most abundant in the San Joaquin Valley, and ascending the western slopes of the southern Sierra Nevada to altitudes of 3000°.

Often planted in southern California as a shade-tree, and for the fuel produced quickly and abundantly from pollarded trees.

In San Bernardino and San Diego Counties, California, generally replaced by the var. pubescens Sarg., differing in its pubescent branchlets and ranging eastward to southwestern Nevada and southern Utah. In southern Arizona and near Silver City, Grant County, New Mexico, represented by the var. Thornberii Sarg., differing from the typical P. Fremontii in the more numerous serratures of the leaves, in the ellipsoidal not ovoid capsules with smaller disk and shorter pedicels, and by the var. Toumeyi Sarg., differing from the type in the shallow cordate base of the leaves, gradually narrowed and cuneate to the insertion of the petiole, and in the larger disk of the fruit (Fig. 124). The var. macrodisca Sarg. with a broad disc nearly inclosing the ellipsoidal fruit is known only in the neighborhood of Silver City.

× Populus Parryi Sarg., a probable hybrid of P. Fremontii and P. trichocarpa, with characters intermediate between those of its supposed parents, grows naturally along Cottonwood Creek on the west side of Owens Lake, Inyo County, and in the neighborhood of Fort Tejon, Kern County, and as a street tree is not rare in San Bernardino, California.

9. Populus arizonica Sarg. Cottonwood.

Leaves deltoid or reniform, gradually or abruptly long-pointed at the acuminate entire apex, truncate or broad-cuneate at the wide base, finely serrate with numerous teeth, as they unfold dark red covered below with pale pubescence, pubescent above, ciliate on the margins, thin, glandular with bright red caducous glands, soon becoming glabrous, at [132] maturity subcoriaceous, bright yellow-green, very lustrous, 1½′—2′ long and broad, with a slender yellow midrib and obscure primary veins; petioles laterally compressed, sparingly villose when they first appear, soon glabrous, 1½′—2′ long; leaves on vigorous leading shoots often rounded at apex, cuneate at base, and often 2′ long and 3′ wide, with petioles often 3′ in length. Flowers: staminate aments dense, cylindric, 1—1½′ long, the pistillate slender, many-flowered, 1½′—2′ long, becoming 3′—4′ long before the fruit ripens; disk of the staminate flower broad-oblong; stamens numerous; disk of the pistillate flower deep cup-shaped, nearly entire; ovary ovoid, rounded at apex, slightly 3 or 4-angled, short-stalked, nearly inclosed in the cup-shaped membranaceous disk. Fruit on short stout pedicels, round-ovoid, buff color, slightly 3 or 4-lobed, deeply pitted, thin-walled, about ¼′ long.

A tree, 50°—70° high, with a trunk occasionally 3° in diameter, gracefully spreading and ascending branches forming a broad open head of wide-spreading branches, and slender often pendulous branchlets, pale green and glabrous or puberulous when they first appear, soon becoming glabrous, and light yellow during their first season. Winter-buds narrow, acute, light orange-brown, puberulous toward the base of the outer scales, the terminal about ¼′ long, and two or three times as large as the much-compressed oblong lateral buds. Bark pale gray or rarely white, and deeply divided into broad flat ridges.

Distribution. Banks of mountain streams; southwestern California (Mill Creek, above Forest Home, San Bernardino Mountains) and southern and central Arizona; widely distributed through northern Mexico (var. Jonesii Sarg.); well distinguished from the other Cottonwoods of the United States by its small fruit.

10. Populus texana Sarg.

Leaves thin, glabrous, broadly ovate, gradually narrowed, long-pointed and acuminate at apex, truncate at base, coarsely crenately serrate below the middle, entire above, 3′—3¼′ long and 2¼′—2½′ wide; petioles slender, compressed, 1½′—2½′ in length. Flowers not seen. Fruit: aments slender, glabrous, 2½′—3′ long; fruit oblong-ovoid, acute, deeply pitted, glabrous, thin-walled, 3-valved, ⅓′ in length; disk slightly lobed; pedicel slender, ¹/₁₆′—⅛′ in length; seeds ovoid, acuminate, ¹/₁₆′ long.

A tree up to 60° high, with a trunk sometimes 3° in diameter, stout more or less pendulous branches and stout glabrous pale yellow-brown branchlets. Winter-buds acuminate, glabrous.

In cañons and along the streams of northwestern Texas, where it appears to be the only Cottonwood.

11. Populus McDougallii Rose.

Leaves broadly ovate, abruptly short-pointed or acute at apex, broadly or acutely cuneate or truncate, or on vigorous shoots rarely slightly cordate at base, finely or often coarsely crenately serrate, bluish green, thin, pubescent on the under sides of the midrib and primary veins early in the season, otherwise glabrous, 1½′—3′ long and broad, with slender midribs and veins; petioles slender, slightly compressed, pubescent early in the season, becoming glabrous, 1½′—2′ in length. Flowers not seen. Fruit: aments glabrous, short-stalked, 2′—2½′ long; fruit ovoid and acute at apex to ellipsoidal and acute or acuminate at ends, glabrous, slightly pitted, thin-walled, 3-valved, ⁵/₁₂′—½′ long; disk not more than ⅛′ in diameter; pedicels glabrous, ⅛′—⅕′ in length; seeds oblong-ovoid, acuminate, ⅛′ long.

A tree rarely 90°—110° high, usually much smaller, with erect branches and slender branchlets pubescent or puberulous when they first appear, sometimes becoming glabrous during their first season, and sometimes pubescent during two years.

Distribution. Banks of streams and springs, San Bernardino County, California (Cottonwood Springs, Meca, etc.), and eastward to the bottoms of the Colorado River from Clark County, Nevada, to Yuma, Arizona, and probably the only Cottonwood in this arid region.

Often planted as a street tree in the towns of southwestern California and of adjacent Nevada and Arizona.

12. Populus Wislizenii Sarg. Cottonwood.

Leaves broadly deltoid, abruptly short- or long-pointed at apex, truncate or sometimes cordate at the broad entire base, coarsely and irregularly crenately serrate except toward the entire apex, coriaceous, glabrous, yellow-green and lustrous, 2′—2½′ long, usually about 3′ wide, with a slender yellow midrib, thin remote primary veins and conspicuous reticulate veinlets; petioles slender, glabrous, 1½′—2′ long; on vigorous shoots often 3½′—4′ long and wide with petioles 3½′—4′ in length. Flowers: aments 2′—4′ long, the pistillate becoming 4′—5′ long before the fruit ripens; scales scarious, light red, divided at the apex into elongated filiform lobes; disk of the staminate flower broad and oblique; stamens numerous, with large oblong anthers and short filaments; disk of the pistillate flower cup-shaped, [134] irregularly dentate, inclosing to the middle the long stalked ovary full and rounded at apex, with 3 broad crenulate lobed stigmas raised on the short branches of the style. Fruit oblong-ovoid, thick-walled, acute, 3 or 4-valved, slightly ridged, buff color, ¼′ long; pedicels slender, ½′—¾′ in length and placed rather remotely on the slender glabrous rachis of the ament.

A large tree, with wide-spreading branches, and stout light orange-colored glabrous branchlets. Winter-buds acute lustrous, puberulous. Bark pale gray-brown, deeply divided into broad flat ridges. Wood used as fuel, for fence-posts and the rafters of Mexican houses.

Distribution. Western Texas through New Mexico to the valley of Grand River, western Colorado (Grand Junction, Mesa County); common in the valley of the Rio Grande in western Texas and New Mexico, and the adjacent parts of Mexico.

13. Populus Sargentii Dode.

Leaves ovate, usually longer than broad, abruptly narrowed into a long slender entire acuminate point or rarely rounded at apex, truncate or slightly cordate at base, and coarsely crenately serrate, as they unfold slightly villose above and tomentose on the margins, soon glabrous, light green and very lustrous, 3′—3½′ long, 3½′—4′ wide, with a thin midrib, slender primary veins and reticulate veinlets occasionally furnished on the upper side at the insertion of the petiole with one or two small glands; petioles slender, compressed laterally, 2½′—3½′ long. Flowers: aments short-stalked, glabrous, the staminate 2′—2½′ in length, the pistillate becoming 4′—8′ long before the fruit ripens; scales fimbriately divided at apex, scarious, light brown; disk of the staminate flower broad, oblique, slightly thickened on the margins; stamens 20 or more, with short filaments and yellow anthers; disk of the pistillate flower cup-shaped, slightly lobed on the margin; ovary subglobose, with 3 or 4 sessile dilated or laciniately lobed stigmas. Fruit oblong-ovoid, gradually or abruptly narrowed to the blunt apex, thin-walled, about ⅖′ long and three or four times longer than the pedicel; seeds oblong-obovoid, rounded at apex, about ¹/₁₆′ in length.

A tree 60°—90° tall with a trunk often 6° or 7° in diameter, erect and spreading branches forming a broad open head, and stout glabrous light yellow often angular branchlets conspicuously roughened by the elevated scars of fallen leaf-stalks. Winter-buds ovoid, [135] acute, with light orange-brown puberulous scales. Bark pale, thick, divided by deep fissures into broad rounded ridges broken into closely appressed scales.

Distribution. The common Cottonwood along the streams in the eastern foothill region of the Rocky Mountains from Saskatchewan to New Mexico, and ranging east to the Dakotas, western Nebraska, Kansas, Oklahoma and Texas.

Often planted as a shade and street tree in the Rocky Mountain states; hardy in Massachusetts.

14. Populus balsamifera L. Cottonwood.

Leaves ovate, longer than broad, abruptly acuminate and often long-pointed at apex, subcordate or rarely truncate at the wide base, finely crenately serrate with glandular teeth, furnished on the upper surface at the insertion of the petiole with two glands, thick, glabrous, green and lustrous on the upper surface, paler below, 5′—7′ long and 4′—5′ wide, with stout midribs and conspicuous primary veins sometimes sparingly pilose below early in the season; petioles much compressed laterally, often more or less tinged with red, 3′—4′ in length. Flowers: aments glabrous, short-stalked, the staminate densely flowered, 1½′—2′ long, ½′—¾′ in diameter, the pistillate slender, sparsely flowered, 3′—3½′ in length; scales scarious, light brown, glabrous, dilated and irregularly divided at apex into filiform lobes; disk of the staminate flower broad, oblique, slightly thickened and revolute on the margins; stamens 60 or more, with short filaments and large dark red anthers; disk of the pistillate flower broad, slightly crenate, inclosing about ⅓′ of the ovoid obtusely pointed ovary, with 3 or 4 sessile dilated laciniately lobed stigmas. Fruit on aments 8′—12′ in length, ellipsoidal, pointed, thin-walled, 3 or 4-valved, ⅓′ long, the disk little enlarged; pedicels ⅙′—¼′ in length; seeds oblong-obovoid, rounded at apex, light brown, about ¹/₁₂′ long.

A large tree with massive spreading branches and stout yellow-brown often angular branchlets. Winter-buds resinous, acute, ½′ long with light chestnut brown lustrous scales.

Distribution. Shores of Lake Champlain (Shelburne Point, Chittenden County), Vermont; western New York; Island of the Delaware River above Easton, Northampton County, Pennsylvania; Baltimore County, and Bare Hills, Maryland; northern banks of the Potomac River opposite Plummer’s Island near Washington, D.C.; Artisia, Lowndes County, and Starkville, Oktibbeha County, Mississippi; rare and local.

Populus balsamifera var. virginiana Sarg. Cottonwood.

Populus deltoidea Marsh. at least in part.
Populus nigra β virginiana Castiglioni.

Leaves deltoid to ovate-deltoid, acuminate with entire points, truncate, slightly cordate or occasionally abruptly cuneate at the entire base, crenately serrate above, with incurved glandular teeth, fragrant with a balsamic odor, glabrous, thick and firm, light bright green and lustrous, paler on the lower than on the upper surface, 3′—5′ long and broad, with a stout yellow midrib often tinged with red toward the base, raised and rounded on the upper side, and conspicuous primary veins; petioles slender, pilose at first, soon glabrous, compressed laterally, yellow often more or less tinged with red, 2½′—3½′ long. Flowers and Fruit: as on the type.

A tree, sometimes 100° high, with a trunk occasionally 7°—8° in diameter, divided often 20°—30° above the ground into several massive limbs spreading gradually and becoming pendulous toward the ends, and forming a graceful rather open head frequently 100° across, or on young trees nearly erect above and spreading below almost at right angles with the stem, and forming a symmetrical pyramidal head, and stout branchlets marked with long pale lenticels, terete, or, especially on vigorous trees, becoming angled in their second year, with thin more or less prominent wings extending downward from the two sides and from the base of the large 3-lobed leaf-scars. Winter-buds very resinous, ovoid, acute, the lateral much flattened, ½′ long, with 6 or 7 light chestnut-brown lustrous scales. Bark thin, smooth, light yellow tinged with green on young stems and branches, becoming on old trunks 1½′—2′ thick, ashy gray, and deeply divided into broad rounded ridges broken into closely appressed scales. Wood dark brown, with thick nearly white sapwood, warping badly in drying and difficult to season.

Distribution. Banks of streams, often forming extensive open groves, and toward the western limits of its range occasionally in upland ravines and on bluffs; Province of Quebec and the shores of Lake Champlain, through western New England, western New York, Pennsylvania west of the Allegheny Mountains, and westward to southern Minnesota, North and South Dakota, eastern Nebraska, eastern Kansas, Oklahoma and Texas, and southward through the Atlantic states from Delaware to western Florida, and through the Gulf states to western Texas (Brown County). In the south Atlantic states and the valley of the Lower Ohio River and southward sometimes replaced by a variety with leaves covered above when they unfold with soft white hairs and below with close pubescence more or less persistent during the season especially on the midribs and veins (var. pilosa Sarg.).

Often planted for shelter and ornament on the treeless plains and prairies between the Mississippi River and the Rocky Mountains, and as an ornamental tree in the eastern United States and largely in western and northern Europe.

× Populus canadensis Moench, believed to be a hybrid between the northern glabrous form of P. balsamifera and the European P. nigra L., with several varieties, is cultivated in Europe and occasionally in the United States. The best known of these varieties, × P. canadensis var. Eugenie Schelle, the Carolina Poplar of American nurseries, believed to be a hybrid of the northern Cottonwood with the Lombardy Poplar, has been planted in the United States in immense numbers.

× Populus Jackii Sarg., believed to be a hybrid of the northern Cottonwood with P. tacamahacca, with characters intermediate between those of its supposed parents, grows spontaneously near the mouth of the Chateaugay River and at Beauharnois, Province of Quebec, and at South Haven, Michigan, and is now occasionally cultivated.

15. Populus Palmeri Sarg.

Leaves thin, ovate, gradually or abruptly contracted at apex into a narrow acuminate entire point, cuneate or rounded at the broad base, finely serrate with incurved teeth, ciliate on the margins when they unfold, otherwise glabrous, 2½′—5′ long and 1½′—2¼′ wide; petioles slender, glabrous, 1½′—2½′ in length. Flowers not seen. Fruit: aments glabrous, 12—15 cm. long; fruit ovoid, obtuse, slightly pitted, puberulous, thin-walled, 4-valved, ¼′—⅓′ long, the disk deeply lobed; pedicel slender, ¼′—⅓′ in length.

A tree 60° tall, with a straight trunk 3° in diameter, erect smooth pale branches forming an open pyramidal head, the lower branches smaller, horizontal or pendulous, and slender glabrous branchlets light reddish brown early in the season, becoming pale grayish brown [138] in their second year. Bark pale, 3′—4′ thick, deeply divided by wide fissures into narrow ridges.

Distribution. In moist fertile soil near springs, at the base of high chalky bluffs of Nueces Cañon of the upper Nueces River, Uvalde County, growing with Salix nigra var. Lindheimeri, Carya pecan, Morus rubra and Ulmus crassifolia, and at Strawn, Palo Pinto County, Texas.

2. SALIX L. Willow.

Trees or shrubs, with watery juice, scaly bark, soft wood, slender terete tough branchlets often easily separated at the joints, and winter-buds covered by a single scale of 2 coats, the inner membranaceous, stipular, rarely separable from the outer, inclosing at its base 2 minute opposite lateral buds alternate with 2 small scale-like caducous leaves coated with long pale or rufous hairs. Leaves variously folded in the bud, alternate, simple, lanceolate, obovate, rotund or linear, penniveined; petioles sometimes glandular at the apex, and more or less covering the bud, in falling leaving U-shaped or arcuate elevated leaf-scars displaying the ends of 3 small equidistant fibro-vascular bundles; stipules oblique, serrate, small and deciduous, or foliaceous and often persistent, generally large and conspicuous on vigorous young branches, leaving in falling minute persistent scars. Flowers in sessile or stalked aments, terminal and axillary on leafy branchlets; scales of the ament lanceolate, concave, rotund or obovate, entire or glandular-dentate, of uniform color or dark-colored toward the apex, more or less hairy, deciduous or persistent; disk of the flower nectariferous, composed of an anterior and posterior or of a single posterior gland-like body; stamens 3—12 or 1 or 2, inserted on the base of the scale, with slender filaments free or rarely united and usually light yellow, glabrous, or hairy toward the base, and small ovoid or oblong anthers generally rose-colored before anthesis, becoming orange or purple; ovary sessile or stipitate, conic, obtuse to subulate-rostrate, glandular at the base, glabrous, tomentose or villose, with an abbreviated style divided into 2 short recurved retuse or 2-parted stigmas; ovules 4—8 on each of the 2 placentas. Fruit an acuminate 1-celled capsule separating at maturity into 2 recurved valves. Seeds minute, narrowed at the ends, dark chestnut-brown or nearly black; cotyledons oblong.

Salix inhabits the banks of streams and low moist ground, the alpine summits of mountains, [139] and the Arctic and sub-Arctic regions of the northern hemisphere, ranging south in the New World, with a few species, through the West Indies and Central America to Brazil, and the Andes of Chile, and in the Old World to Madagascar, southern Africa, the Himalayas, Burmah, the Malay peninsula, Java, and Sumatra. Of the 160 or 170 species which are now recognized about seventy are found in North America. Of these twenty-four attain the size and habit of trees, the others being small and sometimes prostrate shrubs. Of exotic species, Salix alba L., and Salix fragilis L., important European timber-trees, are now generally naturalized in the northeastern states. The flexible tough branches of several species are used in making baskets; the bark is rich in tannic acid and is used in tanning leather and yields salicin, a bitter principle valuable as a tonic. Many of the species are cultivated as ornamental trees.

CONSPECTUS OF THE NORTH AMERICAN ARBORESCENT SPECIES

1. Salix nigra Marsh. Black Willow.

Leaves lanceolate, long-acuminate, often falcate, gradually cuneate or rounded at base, finely serrate, thin bright light green, rather lustrous, with obscure reticulate veins, glabrous or often pubescent on the under side of the midribs and veins and on the short slender petioles, 3′—6′ long, ⅛′—¾′ wide; at the north turning light yellow before falling in the autumn; stipules semicordate, acuminate, foliaceous, persistent, or ovoid, minute, and deciduous. Flowers: aments terminal on leafy pubescent branches, narrowly cylindric, 1′—3′ long; scales yellow, elliptic to obovate, rounded at apex and coated on the inner surface with pale hairs; stamens 3—5, with filaments hairy toward the base; ovary ovoid, short-stalked, glabrous, gradually narrowed above the middle to the apex, with nearly sessile slightly divided stigmatic lobes. Fruit ovoid-conic, short-stalked, glabrous, about ⅛′ long, light reddish brown.

A tree, usually 30°—40° high, with usually several clustered stout stems, thick spreading upright branches forming a broad somewhat irregular open head, and reddish brown or gray-brown branchlets pubescent when they first appear, soon glabrous, and easily separated at the joints. Winter-buds acute, about ⅛′ long. Bark 1′—1¼′ thick, dark brown or nearly black and deeply divided into broad flat connected ridges separating freely into thick plate-like scales and becoming shaggy on old trunks. Wood light, soft, weak, light reddish brown, with thin nearly white sapwood; now sawed into lumber in the valley of the lower Mississippi River and largely used for packing cases, cellar and barn floors, in furniture, and in the manufacture of toys and other purposes where strength is not important as it does not warp, check or splinter.

Distribution. Low moist alluvial banks of streams and lakes; southern New Brunswick through southern Quebec and Ontario to the region north of Lake Superior, southward to northern and western North Carolina, through the Piedmont region of South Carolina and Georgia to eastern and central Alabama, and westward to southeastern North Dakota, eastern South Dakota, Nebraska, Kansas, the valley of Wichita River, Oklahoma, and central and western Texas to Valverde County.

In southern Arkansas, in Louisiana and in eastern Texas Salix nigra is often replaced by var. altissima Sarg., differing from the type in the more pubescent young branchlets, leaves and petioles, in the more acute base of the leaves and longer petioles, and in its later flowering. A tree sometimes 120 feet high and the tallest of American Willows.

Salix nigra var. Lindheimeri Schn.

Leaves lanceolate, often slightly falcate, long-pointed and acuminate at apex, cuneate at base, finely glandular-serrate, glabrous, light green on the upper surface, paler below, 4′—5′ long, ⅓′—½′ wide; petioles pubescent early in the season, becoming glabrous, ½′—¾′ in length. Flowers: aments slender, densely villose, 2′—3′ long; scales ovate, acute or rarely rounded at apex, covered with matted white hairs, more abundant on the inner surface; stamens 4 or 5; filaments villose below the middle; ovary ovoid, gradually narrowed to the apex, the 2-lobed stigmas nearly sessile. Fruit ovoid-conic; pedicels about ¼′ long.

A tree, 50°—70° high with a trunk often 3° in diameter, large erect spreading branches forming an open irregular head, and slender branchlets light green and slightly pubescent [142] when they first appear, becoming light orange or yellow-brown and lustrous. Bark thick, pale yellow-brown, deeply furrowed, the surface sometimes separating into long plate-like scales.

Distribution. River banks, central and western Texas from Grayson and Dallas Counties and the lower valley of the Brazos River to the valleys of the San Antonio and upper Guadalupe Rivers; in Coahuila, Nuevo Leon and Tamaulipas.

2. Salix Gooddingii Ball.

Leaves lanceolate to narrow elliptic-lanceolate, acute or acuminate, acutely cuneate at base, finely glandular-serrate, often slightly falcate, silky pubescent when they unfold especially below, glabrous and dull green at maturity, 1½′—3′ long, ¼′—½′ wide, or on vigorous shoots 5′ or 6′ long and ¾′ wide; petioles pubescent, usually becoming glabrous, ⅙′—¼′ in [143] length; stipules orbicular-cordate, coarsely glandular-serrate, pubescent. Flowers: aments pubescent terminal on leafy pubescent branchlets, narrow-cylindric, 1′—2′ long; scales linear-oblanceolate, acute, yellow, hoary tomentose; stamens 3—5; filaments villose toward the base; ovary ovoid-conic, gradually narrowed to the acuminate apex, pubescent or glabrous; style distinct, 2-lobed. Fruit ovoid, acute, light reddish brown, glabrous or pubescent, ¼′ long; pedicels glabrous or rarely pubescent, ¹/₁₆′—⅛′ in length.

A tree, 25°—50° high, with slender light orange-colored or grayish glabrous or pubescent easily separable branchlets. Bark rough, thick, deeply furrowed, sometimes nearly black.

Distribution. River banks; Reed Creek, Shasta County, and Red Bluff, Tehama County, California, southward in the interior valleys and on the western foothills of the Sierra Nevada to the mountain valleys in the southern part of the state, and to northern Lower California; eastward through central and southern Arizona; in southeastern Nevada; through southern New Mexico to western Texas (El Paso, El Paso County, and Fort Davis, Jeff Davis County); and southward into northern Mexico.

3. Salix Harbisonii Schn.

Leaves linear-lanceolate, narrow-elliptic or rarely obovate-lanceolate, acute or short-acuminate, obtusely or acutely cuneate at the base, and finely glandular dentate; when the flowers open more or less pubescent especially below or glabrous, and at maturity green on the upper surface, pale on the lower surface, glabrous, 4′ or 5′ long, ¾′ broad; petioles villose early in the season, becoming glabrous, ¼′ in length, minutely glandular at apex; stipules wanting or minute, semicordate, acute, pubescent on vigorous leading branches and sometimes ¼′ long. Flowers: aments terminal on leafy branchlets, 2½′—3′ in length, their rachis villose-pubescent; scales ovate or ovate-oblong, obtuse or acute; stamens usually 5—7, rarely 3—9; filaments densely villose; ovary ovoid, long-acuminate, glabrous, long-stalked; style short, distinct, 2-lobed. Fruit acuminate and long-pointed, acute at base, ¼′ long and about as long as its pedicel.

A tree, 30°—50° high, with a trunk 10′ or 12′ in diameter, with often pendulous branches, and slender branchlets more or less densely pubescent or tomentose or nearly glabrous when they first appear, becoming glabrous and dark reddish purple in their second season, [144] and easily separable at the joints; often only a large shrub. Bark thick, deeply furrowed, dark red-brown, separating on the surface into small appressed scales.

Distribution. River banks and the borders of swamps; Dismal Swamp, Norfolk County, Virginia; near Goldsboro, Wayne County, North Carolina; common in the coast region of South Carolina and Georgia, extending up the Savannah River at least as far as Augusta, Richmond County, and through southern Georgia to the valley of the Flint River; swamps near Jacksonville, Duval County, and in the neighborhood of Apalachicola, Florida.

4. Salix amygdaloides Anders. Peach Willow. Almond Willow.

Leaves lanceolate to ovate-lanceolate, frequently falcate, gradually or abruptly narrowed into a long slender point, cuneate or gradually rounded and often unequal at base, finely serrate, slightly puberulous when they unfold, becoming at maturity thin and firm in texture, light green and lustrous above, pale and glaucous below, 2½′—4′ long, ¾′—1¼′ wide, with a stout yellow or orange-colored midrib, prominent veins and reticulate veinlets; petioles slender, nearly terete ½′—¾′ in length; stipules reniform, serrate, often ½′ broad on vigorous shoots, usually caducous. Flowers: aments on leafy branchlets, elongated, cylindric, slender, arcuate, stalked, pubescent or tomentose, 2′—3′ long; scales yellow, sparingly villose on the outer, densely villose on the inner face, the staminate broadly ovate, rounded at the apex, the pistillate oblong-obovate, narrower, caducous; stamens 5—9, with free filaments slightly hairy at the base; ovary oblong-conic, long-stalked, glabrous, with a short style and emarginate stigmas. Fruit globose-conic, light reddish yellow, about ¼′ in length.

A tree, sometimes 60°—70° high, with a single straight or slightly inclining trunk rarely more than 2° in diameter, straight ascending branches, and slender glabrous or rarely pilose (f. pilosiuscula Schn.) branchlets marked with scattered pale lenticels, dark orange color or red-brown and lustrous, becoming in their first winter light orange-brown. Winter-buds broadly ovoid, gibbous, dark chestnut-brown, very lustrous above the middle, light orange-brown below, ⅛′ long. Bark ½′—¾′ thick, brown somewhat tinged with red, and divided by irregular fissures into flat connected ridges separating on the surface into thick plate-like scales. Wood light, soft, close-grained, light brown, with thick nearly white sapwood.

Distribution. Banks of streams; Province of Quebec from the neighborhood of Montreal to Winnipeg, and along the fiftieth degree of north latitude to southeastern British Columbia, and to central New York, along the southern shores of Lake Erie, and through northern Ohio to northern Indiana, southwestern Illinois, northern and central Missouri, and to [145] Kansas, northwestern Oklahoma and northwestern Texas; in Colorado, Utah and Nevada to central Oregon and southeastern Washington.

Salix amygdaloides var. Wrightii Schn.

Leaves lanceolate, gradually acuminate and long-pointed at apex, cuneate at base, finely serrate, occasionally slightly falcate, glabrous, yellow-green on the upper surface, pale on the lower surface, 1½′—2′ long, ¼′—⅓′ wide, and on vigorous summer shoots sometimes 4′ or 5′ long and ½′ wide; petioles slender, glabrous, ¼′—⅓′ in length. Flowers and Fruit as in the species.

A small or large tree best distinguished from S. amygdaloides by the distinctly yellow or yellowish brown glabrous branchlets.

Distribution. Barstow, Ward County, common along the Rio Grande near El Paso and at Belon, El Paso County, and on Amarillo Creek, Potter County, western Texas; through southern New Mexico to the Sacramento Mountains, Otero County.

5. Salix Bonplandiana var. Toumeyi Schn.

Leaves 4′—6′ long, ½′—¾′ wide, linear-lanceolate to oblong-lanceolate, acuminate with a long slender point at apex, gradually narrowed and often unequal at the cuneate base, obscurely serrate with glandular teeth, or entire with revolute margins, thick and firm, reticulate-venulose, yellow-green and lustrous above, silvery white below, with a broad yellow midrib; falling irregularly during the winter; petioles stout, grooved, reddish; stipules ovate, rounded, slightly undulate, thin and scarious, ⅛′—¼′ broad, often persistent during the summer. Flowers: aments on leafy branchlets, cylindric, erect, slender, short-stalked, the staminate 1′—1½′ long and somewhat longer than the pistillate; scales broadly obovate, rounded at the apex, light yellow, villose on the outer surface and glabrous or slightly hairy above the middle on the inner surface; stamens usually 3, with free filaments slightly hairy at the base; ovary slender, oblong-conic, short-stalked, glabrous, with nearly sessile much-thickened club-shaped stigmas, sometimes nearly encircled below by the large broad ventral gland. Fruit ovoid-conic, rounded at base, light reddish yellow.

A tree, rarely more than 30° high, with a trunk 12′—15′ in diameter, slender erect and spreading branches often pendulous at the ends, forming a broad round-topped head, and [146] slender glabrous branchlets marked with occasional pale lenticels, light yellow, becoming light or dark red-brown and lustrous, and paler orange-brown in their second year. Winter-buds narrowly ovoid, long-pointed, more or less falcate, bright red-brown, lustrous, ¼′ long. Bark ½′—¾′ thick, dark brown or nearly black, and deeply divided by narrow fissures into broad flat ridges separating on the surface into closely appressed scales.

Distribution. Banks of streams in the cañons of the mountains of central and southern Arizona (Sicamore Cañon near Flagstaff and Sabino Cañon, Santa Catalina Mountains); and southwestern New Mexico (cañon, Saint Louis Mountains, Grant County); in Chihuahua, Sonora and Lower California.

The typical S. Bonplandiana H. B. K. with broader and more coarsely serrate leaves, and flower-aments appearing from July to January from the axils of mature leaves is widely distributed in Mexico and ranges to Guatemala.

6. Salix lævigata Bebb. Red Willow.

Leaves obovate, narrowed and rounded or acute and mucronate at apex, cuneate at base, with slightly revolute obscurely serrate margins, on sterile branches lanceolate or oblong-lanceolate, [147] acute or acuminate, when they unfold light blue-green and coated on the lower surface with long pale or tawny deciduous hairs, at maturity glabrous, dark blue-green and lustrous above, paler and glaucous below, 3′—7′ long, ¾′—1½′ wide, with a broad flat yellow midrib; petioles broad, grooved, puberulous, rarely ½′ long; stipules ovate, acute, finely serrate, usually small and caducous. Flowers: aments cylindric, slender, lax, elongated, 2′—4′ long, on leafy branchlets; scales peltate, dentate at apex, covered with long pale hairs, the staminate obovate, rounded, the pistillate narrower and more or less truncate; stamens usually 5 or 6, with free filaments hairy at the base; ovary conic, acute, rounded below, short-stalked, glabrous, with broad spreading emarginate stigmatic lobes. Fruit elongated, conic, long-stalked, nearly ¼′ in length.

A tree, 40°—50° high, with a straight trunk 2° in diameter, slender spreading branches, and slender light or dark orange-colored or bright red-brown glabrous, or in one form tomentose or villose (f. araquipa Jeps.) branchlets; often much smaller, with an average height of 20°—30°. Winter-buds ovoid, somewhat obtuse, pale chestnut-brown, ⅛′—¼′ long. Bark ¾′—1′ thick, dark brown slightly tinged with red and deeply divided into irregular connected flat ridges broken on the surface into thick closely appressed scales. Wood light, soft, light brown tinged with red, with thick nearly white sapwood.

Distribution. Banks of streams; western California from the Oregon boundary to the southern borders of the state, ascending to altitudes of 4500° on the western slopes of the southern Sierra Nevada, and eastward to Mohave and Yavapai Counties, Arizona, southeastern Nevada and southwestern Utah.

7. Salix longipes Shuttl.

Leaves lanceolate, acuminate or on fertile branches occasionally rounded at the apex, rounded or cuneate at the base, finely serrate, hoary-tomentose early in the season, becoming glabrous above, and pale and glabrous or pubescent below, 2′—4′ long, ½′—¾′ wide; petioles hoary-tomentose, ¼′—½′ long; stipules minute, ovate, acute, hoary-tomentose, caducous, on vigorous shoots foliaceous, reniform, serrate above the middle, often ¾′ in diameter. Flowers: aments terminal on leafy tomentose or glabrous branchlets, narrow-cylindric, 3′ or 4′ long; scales ovate, rounded at the apex, yellow, densely villose-pubescent; stamens 3—7, usually 5 or 6, the filaments hairy toward the base; ovary ovoid-conic, acute, cuneate at the base with a short 2-lobed style, and pedicels up to ¼′ in length. Fruit ovoid, often rather abruptly contracted above the middle, ¼′ in length.

A tree, 20°—30°, high with a trunk occasionally 12′—18′ in diameter, spreading branches, and glabrous or pubescent red-brown or gray-brown branchlets; or more often a shrub. Bark dark, sometimes nearly black, deeply divided into broad ridges covered by small closely appressed scales.

Distribution. Borders of swamps and streams; coast of North Carolina southward to the Everglade Keys of Florida, ranging westward in Florida to the valley of the Saint Marks River, Wakulla County; in Cuba.

A variety with narrower summer leaves and longer petioles is var. venulosa Schn.

Distribution. Newbern, Craven County, North Carolina, southward near the coast to northern and western Florida, ranging inland in Georgia to the banks of the Savannah River near Augusta, Richmond County, and to Traders Hill, Charlton County; in the neighborhood of New Orleans, Louisiana (Drummond); in southwestern Oklahoma and in western Texas (Blanco, Kendall, Kerr, Bandera and Uvalde Counties).

A variety with obtuse stipules, usually glabrous branchlets and lanceolate or narrow elliptic-lanceolate leaves is distinguished as var. Wardii Schn.

Distribution. Banks of the Potomac River, District of Columbia, and Alleghany County, Maryland to Natural, Rockbridge, Fairfax and Elizabeth Counties, Virginia; northern Kentucky; northern Tennessee; northeastern Mississippi (near Iuka, Tishamingo County); St. Clair and Madison Counties, Illinois; more abundant in Missouri from Pike County southward to southwestern Kansas, western Arkansas and eastern Oklahoma.

8. Salix lasiandra Benth. Yellow Willow.

Leaves lanceolate to ovate-lanceolate, acuminate and long-pointed at apex, cuneate or rounded at base, often slightly falcate, finely serrate, glabrous, dark green and lustrous above, pale or glaucous below, 1½′—3′ long, about ½′ wide, on vigorous summer shoots often 6′ or 7′ long and 1½′ wide; petioles slender, glabrous, glandular at apex, ¼′ in length, or on summer shoots stout and 1′—1½′ long; stipules reniform, caducous. Flowers: aments terminal on leafy puberulous branchlets, narrow-cylindric, 2½′—3′ in length; scales pale pubescent, those of the staminate ament lanceolate-acuminate to obovate and rounded at apex and entire, those of the pistillate ament obovate and usually dentate near the apex; stamens 5—9; filaments hairy below the middle; ovary rather abruptly narrowed above the middle and acuminate, long-stalked; style short with slightly emarginate lobes. Fruit light red-brown, ¼′ long; pedicels about ¹/₁₆′ in length.

Distribution. Valley of the Yukon River near Dawson, Yukon, Vancouver Island, and southward near the coast of Washington and Oregon, and on the western slope of the Sierra Nevada and on the coast ranges to southern California, ranging from the sea-level to altitudes of 8500° on the southern Sierra Nevada; in New Mexico (Glenwood, Soccoro County, and Santa Fé, Santa Fé County); in Colorado (Buena Vista, Chaffee County, Alice Eastwood). Passing into var. caudata Sudw., distinguished by its caudate-acuminate leaves green on both surfaces, and by its bright yellow or orange-yellow branchlets, and ranging from northeastern Oregon and eastern Washington through Idaho, and from northern Wyoming to southern Colorado, Utah and Nevada.

A variety (var. lancifolia Bebb), differing from the typical S. lasiandra in the gray or rusty villose pubescence covering the branchlets during their first and sometimes their second season and the lower surface of the young leaves, is distributed from Dawson in the valley of the Yukon River southward to the valley of the upper Nesqually River, Washington, to the valley of the Willamette River (Salem, Oregon), to Santa Cruz, Santa Cruz County, and to the San Bernardino Mountains, California.

9. Salix lucida Muehl. Shining Willow.

Leaves ovate-lanceolate, or narrow lanceolate (f. angustifolia Anders.), acuminate and long-pointed at apex, cuneate or rounded at base, finely serrate, 3′—5′ long, 1′—1½′ wide, covered when they unfold with scattered pale caducous hairs, at maturity coriaceous, smooth and lustrous, dark green above, paler below, with a broad yellow midrib, and slender primary veins arcuate and united near the margins; petioles stout, yellow, puberulous, glandular at the apex, with several dark or yellow conspicuous glands, ¼′—½′ long; stipules nearly semicircular, glandular-serrate, membranaceous, ⅛′—¼′ wide, often persistent during the summer. Flowers: aments erect, tomentose, on stout puberulous peduncles terminal on short leafy branchlets, the staminate oblong-cylindric, densely flowered, about 1½′ in length, the pistillate slender, elongated, 1½′—2′ long, often persistent until late in the season; scales oblong or obovate, rounded, entire, erose or dentate at apex, light yellow, nearly glabrous or coated on the outer surface with pale hairs, often ciliate on the margins; stamens usually 5, with elongated free filaments slightly hairy at base; ovary narrowly cylindric, long-stalked, elongated, glabrous, with nearly sessile emarginate stigmas. Fruit: cylindric, lustrous, about ⅓′ long.

A tree, occasionally 25° high, with a short trunk 6′—8′ in diameter, erect branches forming a broad round-topped symmetrical head, and stout glabrous branchlets dark orange color [150] and lustrous in their first season, becoming darker and more or less tinged with red the following year; usually smaller and shrubby in habit. Winter-buds narrowly ovoid, acute, light orange-brown, lustrous, about ¼′ long. Bark thin, smooth, dark brown slightly tinged with red.

Distribution. Banks of streams and swamps; Newfoundland to the shores of Hudson’s Bay and northwestward to the valley of the Mackenzie River and the eastern base of the Rocky Mountains, southward to southern Pennsylvania, northeastern Iowa, the Turtle Mountains, North Dakota, and eastern Nebraska; very abundant at the north, rare southward; a variety from extreme northeastern New England and adjacent New Brunswick and Quebec (var. intonsa Fernald) is distinguished by its often linear leaves rufous pubescent during the season on the under side of the veins and by its pubescent branchlets; a shrub or tree up to 25°.

10. Salix taxifolia H. B. K.

Leaves linear-lanceolate, narrowed at the ends, acute, slightly falcate, mucronate at the apex, entire or rarely obscurely dentate above the middle, coated as they unfold with long soft white hairs, at maturity pale gray-green, slightly puberulous, ⅓′—1⅓′ long, ¹/₁₂′—⅛′ wide, with a slender midrib, thin arcuate veins, and thickened slightly revolute margins; petioles stout, puberulous, rarely ¹/₁₂′ long; stipules ovate, acute, scarious, minute, caducous. Flowers: aments densely flowered, oblong-cylindric or subglobose, ¼′—½′ long, terminal, or terminal and axillary on the staminate plant, on short leafy branchlets; scales oblong or obovate, rounded or acute and sometimes apiculate at apex, coated on the outer surface with hoary tomentum and pubescent or glabrous on the inner; stamens 2, with free filaments hairy below the middle; ovary ovoid-conic, short-stalked or subsessile, villose, with nearly sessile deeply emarginate stigmas. Fruit cylindric, long-pointed, bright red-brown, more or less villose, short-stalked, about ¼′ long.

A tree, often 40°—50° high, with a trunk 18′ in diameter, erect and drooping branches forming a broad open head, and slender branchlets covered during their first season with hoary tomentum, becoming light reddish or purplish brown and much roughened by the elevated persistent leaf-scars. Winter-buds ovoid, acute, dark chestnut-brown, puberulous, about ¹/₁₆′ long and nearly as broad as long. Bark of the trunk ¾′—1′ thick, light gray-brown, and divided by deep fissures into broad flat ridges covered by minute closely appressed scales.

Distribution. Near El Paso, Texas; southwestern New Mexico, and along mountain [151] streams in southern Arizona; southward through Mexico to Guatemala, and on the Sierra de la Victoria, Lower California.

11. Salix sessilifolia Nutt.

Leaves linear-lanceolate to elliptic-lanceolate, acute or acuminate at apex, cuneate at base, entire or furnished above the middle with a few remote apiculate glandular teeth, bluish green and thickly covered with silky white hairs most abundant on the lower side of the midrib, 1′—2′ long, ½′—¾′ wide, or on vigorous summer shoots often 4′ long and 1¼′ wide; petioles densely villose-pubescent, ¹/₁₆′—⅛′ in length; stipules ovate to lanceolate, acute, entire or denticulate. Flowers: aments appearing after the leaves, terminal on leafy branchlets, densely hoary-tomentose, 1½′—2½′ long; scales broadly elliptic, acute or rounded at apex, cuneate at base, densely villose-tomentose; stamens 2; filaments villose below the middle; ovary sessile, villose, the stigmas sessile, deeply 2-lobed. Fruit ovoid-acuminate, densely villose, pubescent.

A shrub or small tree occasionally 20° high, with short hairy tomentose branchlets.

Distribution. River banks, southwestern British Columbia; Whitcomb County, Washington, and on the Umpqua and Willamette Rivers, western Oregon. Southward passing into

Var. Hindsiana Anders., a large shrub with numerous stems often 20° high, differing in its more linear or narrow lanceolate usually entire leaves on longer petioles, smaller aments and pubescent, not tomentose, branchlets; and distributed from the valleys of central California to southwestern Oregon. A shrubby form of S. sessilifolia (var. leucodendroides Schn.) with longer and broader leaves is common on the banks of streams in southern California.

12. Salix exigua Nutt.

Leaves lanceolate to oblanceolate, acuminate at the ends, often slightly falcate, minutely glandular-serrate above the middle, bluish green and glabrous above, covered below with appressed silky white hairs, 1½′—3′ long, ⅛′—¼′ wide, or on summer shoots sometimes 4½′ long and 1½′ wide; petioles glabrous, ¹/₁₆′ long or less; stipules minute or wanting. Flowers: aments terminal and solitary or terminal and axillary, on leafy glabrous branchlets, 1′—2′ in length; scales hoary pubescent, lanceolate and acute on staminate aments, often wider, obovate and rounded at the apex on pistillate aments; stamens, 2, filaments hairy [152] below the middle; ovary sessile, villose, the stigmatic lobes sessile. Fruit ovoid, acuminate, glabrous.

A shrub with stems 10° or 12° tall, or rarely a tree 25° high, with a trunk 5′ or 6′ in diameter, thin spreading branches forming a round-topped head, and slender glabrous red-brown branchlets. Bark of the trunk thin, longitudinally fissured, grayish brown.

Distribution. Southern Alberta and valley of the Fraser River (Clinton), British Columbia, southward through western Washington and Oregon to San Diego County, California, and southeastern Nevada, and eastward to southern Idaho, central Nevada and western Wyoming (Yellowstone National Park).

Apparently only truly a tree on the banks of the Palouse and other streams of eastern Washington.

Several shrubby forms of S. exigua found in Nevada, Arizona, Colorado, western Nebraska and in Lower California are distinguished.

13. Salix longifolia Muehl. Sand Bar Willow.

Leaves linear-lanceolate, often somewhat falcate, gradually narrowed at the ends, long-pointed, dentate with small remote spreading callous glandular teeth, 2′—6′ long, ⅛′—⅓′ wide, when they unfold coated below with soft lustrous silky hairs, at maturity thin, glabrous, light yellow-green, darker on the upper than on the lower surface, with a yellow midrib, slender arcuate primary veins, and slender reticulate veinlets; petioles grooved, ⅛′—¼′ long; stipules ovate-lanceolate, foliaceous, about ¼′ long, deciduous. Flowers: aments cylindric on leafy branchlets, pubescent, the staminate about 1′ long, ⅓′ broad, terminal and axillary, the pistillate elongated, 2′ or 3′ long, about ¼′ broad; scales obovate-oblong, entire, erose or dentate above the middle, light yellow-green, densely villose on the outer surface, slightly hairy on the inner; stamens 2, with free filaments slightly hairy at the base; ovary oblong-cylindric, acute, short-stalked, glabrous or pubescent, with large sessile deeply lobed stigmas. Fruit light brown, glabrous or villose, about ¼′ long.

A tree, usually about 20° high, with a trunk only a few inches in diameter, spreading by stoloniferous roots into broad thickets, short slender erect branches, and slender glabrous light or dark orange-colored or purplish red branchlets, growing darker after their first season; occasionally 60°—70° high, with a trunk 2° in diameter; often a shrub not more than 5°—6° tall. Winter-buds narrowly ovoid, acute, chestnut-brown, about ⅛′ long. Bark [153] ⅛′—¼′ thick, smooth, dark brown slightly tinged with red and covered with small closely appressed irregularly shaped scales. Wood light, soft, light brown tinged with red, with thin light brown sapwood.

Distribution. River banks, sand bars and alluvial flats; shores of Lake St. John, Quebec to Manitoba, and southward through western New England to northeastern Virginia, southern Ohio, Indiana and Illinois, western Kentucky, south Tennessee, to the mouth of the Mississippi River, and westward to southwestern South Dakota, southwestern Wyoming, northeastern Colorado, western Kansas and Oklahoma, and northern Texas.

From central and northwestern Texas to northeastern Mexico and southern New Mexico represented by var. angustissima Anders., differing in the absence of a dorsal gland in the male flowers and in the silky pubescence of the young ovary.

In the northern Rocky Mountains region replaced by var. pedunculata Anders., differing from the type in its narrower linear leaves, glabrous ovaries and longer pedicels of the fruit, and ranging from western South Dakota and northwestern Wyoming, through eastern Montana, Saskatchewan, and Alberta, to the valley of the Yukon River in the neighborhood of Dawson.

A shrubby form with leaves densely covered with silky pubescence (var. Wheeleri Schn.) is distributed from New Brunswick to North Dakota, Nebraska and Beckham County, Oklahoma.

14. Salix lasiolepis Benth. Arroyo Willow.

Leaves oblanceolate to lanceolate-oblong, often inequilateral and occasionally falcate, acute or acuminate or rarely rounded at apex, gradually or abruptly cuneate or rounded at base, entire or remotely serrate, pilose above and coated below with thick hoary tomentum when they unfold, at maturity thick and subcoriaceous, conspicuously reticulate-venulose, dark green and glabrous above, pale or glaucous and pubescent or puberulous below, 3′—6′ long, ½′—1′ wide, with a broad yellow midrib and slender arcuate veins forked and united within the slightly thickened and revolute margins; petioles slender, ⅛′—½′ long; stipules ovate, acute, coated with hoary tomentum, minute and caducous, or sometimes foliaceous, semilunar, acute or acuminate, entire or denticulate, dark green above, pale below, persistent. Flowers: aments erect, cylindric, slightly flexuose, densely flowered, nearly sessile on short tomentose branchlets, 1½′ long, the staminate ½′ thick, and nearly twice as thick as the pistillate; scales oblong-obovate, rounded or acute at the apex, dark-colored, clothed with long crisp white hairs, persistent under the fruit; stamens 2, with elongated glabrous filaments more or less united below the middle; ovary narrow, cylindric [154] acute and long-pointed, dark green, glabrous, with a short style and broad nearly sessile stigmas. Fruit oblong-cylindric, light reddish brown, about ¼′ long.

A tree, 20°—35° high, with a trunk 3′—7′ in diameter, slender erect branches forming a loose open head, and stout branchlets coated at first with hoary tomentum, bright yellow or dark reddish brown and puberulous or pubescent during their first year, becoming darker and glabrous in their second season; or often at the north and at high altitudes a low shrub. Winter-buds ovoid, acute, compressed, contracted laterally into thin wing-like margins, light brownish yellow, glabrous or puberulous. Bark on young stems and on the branches thin, smooth, light gray-brown, becoming on old trunks dark, about ⅓′ thick, roughened by small lenticels and broken into broad flat irregularly connected ridges. Wood light, soft, close-grained, light brown, with thick nearly white sapwood; in southern California often used as fuel.

Distribution. Banks of streams in low moist ground; valley of the Klamath River, California, southward along the foothills of the Sierra Nevada, the central valley, and on the Coast Ranges to southern California; on Santa Catalina Island and on the mountains of southern Arizona; on the Sierra de Laguna, Lower California; occasionally ascending to altitudes of 4000° above the sea.

15. Salix Mackenzieana Barr.

Leaves lanceolate to oblanceolate, or elliptic, long-pointed at apex, cuneate or rounded at base, finely crenately serrate, reddish and pilose with caducous pale hairs when they unfold, at maturity thin and firm in texture, light green above, pale below, 1½′—2′ long, about ½′—¾′ wide, on summer shoots, often 4′ long and 1½′ wide, with a slender yellow midrib, arcuate veins, and obscure reticulate veinlets; petioles thin, yellow, about ⅓′ long; stipules reniform, conspicuously veined, about ¹/₁₆′ broad. Flowers: aments densely flowered, glabrous, erect, often more or less curved, about 1½′ long, terminal on short leafy branchlets; scales oblanceolate, acute, dark-colored; stamens 2, with elongated free glabrous filaments; ovary cylindric, long-stalked, elongated, gradually narrowed into a short style, with spreading emarginate stigmas. Fruit ovoid, acuminate, light brown, about ⅛′ long; pedicels about ⅙′ in length.

A small tree, with a slender trunk, upright branches forming a narrow shapely head, and slender branchlets marked with scattered lenticels, glabrous or slightly puberulous and often tinged with red when they first appear, soon becoming yellow and lustrous, growing lighter colored in their second year. Winter-buds ovoid, rounded on the back, compressed and acute at the apex, bright orange color, about ⅛′ long.

Distribution. Borders of streams and swamps; shores of Great Slave Lake southward through the region at the eastern base of the Rocky Mountains to Saskatchewan, northern Idaho, and northwestern Wyoming, and to western Nevada (Lake County; M. S. Bebb), and on the high Sierra Nevada in Calaveras and Mariposa Counties, California (W. L. Jepson).

16. Salix missouriensis Bebb.

Leaves lanceolate or oblanceolate, acuminate and long-pointed at apex, gradually narrowed from above the middle to the cuneate or rounded base, finely glandular-serrate, coated with pale hairs on the lower surface and pilose on the upper surface when they unfold, soon becoming nearly glabrous, at maturity thin and firm, dark green above, pale and often silvery white below, 4′—6′ long, 1′—1½′ wide, with slender veins often united near the margins and connected by coarse reticulate veinlets; petioles stout, pubescent or tomentose, ½′—¾′ long; stipules foliaceous, semicordate, pointed or rarely reniform and obtuse, serrate with incurved teeth, dark green and glabrous on the upper side, coated on the lower with hoary tomentum, reticulate-venulose, often ½′ long, deciduous or persistent during [156] the season. Flowers: aments oblong-cylindric, densely flowered, appearing early in February on short leafy branchlets, the staminate 1½′ long and nearly ½′ wide and rather longer than the more slender pistillate aments becoming at maturity lax and 3′—4′ long; scales oblong-obovate, light green, and covered on the outer surface with long straight white hairs; stamens 2, with elongated free glabrous filaments; ovary cylindric, short-stalked, beaked, glabrous, with a short style and spreading entire or slightly emarginate stigmas. Fruit narrow, long-pointed, light reddish brown, ⅓′ in length; pedicels about half the length of the scales.

A tree, 40°—50° high, with a tall straight trunk 10′—12′ or rarely 18′ in diameter, rather slender upright slightly spreading branches forming a narrow open symmetrical head, and slender branchlets marked by small scattered orange-colored lenticels, light green and coated during their first year with thick pale pubescence, becoming reddish brown and glabrous or puberulous in their second winter. Winter-buds ovoid, round, or flattened, acute at the apex, reddish brown, hoary-tomentose, nearly 1′ long. Bark thin, smooth, light gray, slightly tinged with red, and covered with minute closely appressed plate-like scales. Wood dark red-brown, with thin pale sapwood; durable, used for fence-posts.

Distribution. Deep sandy alluvial bottom-lands of the Missouri River in southwestern Nebraska to western Missouri; through northeastern Kansas and eastern Oklahoma to Cache Creek, Comanche County (G. W. Stevens); and from the neighborhood of St. Louis to southeastern and western Iowa.

17. Salix pyrifolia Anders.

Leaves ovate to oblong-lanceolate, acute at apex, broad and rounded and usually subcordate at base, finely glandular serrulate, balsamic particularly while young, when they unfold thin, pellucid, red and coated below with long slender caducous hairs, at maturity thin and firm, dark green above, pale and glaucous below, 2′—4′ long, 1′—1½′ wide, with a yellow midrib and conspicuous reticulate veinlets; petioles reddish or yellow, ⅓′—½′ long; stipules often wanting or on vigorous shoots foliaceous, broadly ovate and acute. Flowers: aments cylindric, 1′—1½′ long, on short leafy branchlets, the staminate 1′—1¼′ long and ¾′ in diameter and shorter and broader than the pistillate ament; scales obovate, rose-colored, coated with long white hairs; stamens 2, with free filaments and reddish ultimately yellow anthers; ovary narrow-ovoid, long-stalked, gradually contracted above the middle, with a short style and emarginate stigmas. Fruit ovoid-conic, ¼′ long, dark orange color; pedicels ⅙′ in length.

Usually a shrub, often making clumps of crowded slender erect stems generally destitute of branches except near the top, rarely arborescent, with a height of 25°, a trunk 12′—14′ in diameter, erect branches, and comparatively stout reddish brown branchlets becoming olive-green in their second year and marked with narrow slightly raised leaf-scars. Winter-buds acute, much-compressed, bright scarlet, very lustrous, about ¼′ long. Bark thin, smooth, dull gray.

Distribution. Cold wet bogs; Newfoundland and the coast of Labrador to the valley of the Saskatchewan and the Mackenzie, and British Columbia, and to northern Maine, New Hampshire, Vermont, New York, Michigan, and northeastern South Dakota; reported to become arborescent only near Fort Kent on the St. John River, Aroostook, Maine.

18. Salix amplifolia Cov.

Leaves oval to broadly obovate, rounded or broadly pointed at apex, gradually or abruptly narrowed at the cuneate base, dentate-serrulate or entire, densely villose when they unfold, with long matted white hairs, at maturity nearly glabrous, pale yellow-green above, slightly glaucous below, 2′—2½′ long, 1′—1½′ wide, with a midrib broad and hoary-tomentose toward the base of the leaf and thin and glabrous above the middle; petioles slender, tomentose. Flowers: aments appearing about the middle of June, stout, pedunculate, tomentose, on leafy branchlets, the staminate 1½′—2′ long and shorter than the pistillate; scales oblanceolate or lanceolate, dark brown or nearly black, covered with long pale hairs; stamens 2, with slender elongated glabrous filaments; ovary ovoid-lanceolate, short-stalked, glabrous or slightly pubescent, gradually narrowed into the elongated slender style crowned with a 2-lobed slender stigma. Fruit ovoid-lanceolate, glabrous, short-stalked, ¼′ long.

A tree, occasionally 25° high, with a trunk a foot in diameter, and stout branchlets conspicuously roughened by the large elevated U-shaped leaf-scars, and marked by occasional pale lenticels, coated at first with thick villose pubescence, becoming during their second and third years dark dull reddish purple.

Distribution. Sand dunes on the shores of Yakutat Bay and Disenchantment Bay, Alaska.

19. Salix alaxensis Cov. Feltleaf Willow.

Leaves elliptic-lanceolate to obovate, acute, acuminate or occasionally rounded at apex, gradually narrowed into a short thick petiole, coated above as they unfold with thin pale deciduous tomentum and covered below with a thick mass of snowy white lustrous [158] hairs persistent on the mature leaves, entire, often somewhat wrinkled, dull yellow-green above, 2′—4′ long, 1′—1½′ wide, with a broad yellow midrib; stipules linear-lanceolate to filiform, entire, ½′—¾′ long, usually persistent until midsummer. Flowers: aments appearing in June when the leaves are nearly fully grown, stout, erect, tomentose, stalked, on leafy branchlets, the staminate 1′—1½′ long, much shorter than the pistillate; scales oblong-ovate, rounded at apex, dark-colored, and coated with long silvery white soft hairs; stamens 2, with slender elongated filaments; ovary acuminate, short-stalked, covered with soft pale hairs, gradually narrowed into the elongated slender style, with 2-lobed stigmas. Fruit nearly sessile, ovoid, acuminate covered with close dense pale tomentum, ¼′ long.

A tree, sometimes 30° high, with a trunk 4′—6′ in diameter, and stout branchlets thickly coated at first with matted white hairs, becoming in their second year glabrous, dark purple, lustrous, marked by large elevated pale scattered lenticels and much roughened by large U-shaped leaf-scars; often shrubby, and in the most exposed situations frequently only a foot or two high, with semiprostrate stems.

Distribution. Coast of Alaska from the Alexander Archipelago to Cape Lisbourne, and eastward to the valley of the Mackenzie River and to the shores of Coronation Gulf; the only arborescent Willow in the coast region west and north of Kadiak Island; attaining its largest size from the Shumagin Islands eastward.

20. Salix Bebbiana Sarg.

Leaves oblong-obovate to oblong-elliptic or lanceolate, acuminate and short-pointed or acute at apex, gradually narrowed and cuneate or rounded at base, remotely and irregularly serrate usually only above the middle, or rarely entire, when they unfold pale gray-green, glabrous or villose, and often tinged with red on the upper surface and coated on the lower with pale tomentum or pubescence, at maturity thick and firm, dull green and glabrous or puberulous above, blue or silvery white and covered with pale rufous pubescence below, especially along the midrib, veins, and conspicuous reticulate veinlets, 1′—3′ long, ½′—1′ wide; petioles slender, often pubescent, reddish, ¼′—½′ long; stipules foliaceous, semicordate, glandular-dentate, sometimes nearly ½′ long on vigorous shoots, deciduous. Flowers: aments terminal on short leafy branchlets; scales ovate or oblong, rounded at apex, broader on the staminate than on the pistillate plant, yellow below, rose color at apex, villose with long pale silky hairs, persistent under the fruit; staminate aments cylindric, obovoid, narrowed at base, densely flowered, ¾′—1′ long, ½′—1′ thick; pistillate aments oblong-cylindric, loosely flowered, 1′—1¼′ long, ½′ thick; stamens 2, with free glabrous filaments; ovary [159] cylindric, villose; with long silky white hairs, gradually narrowed at apex, with broad sessile entire or emarginate spreading yellow stigmas; pedicel villose, about ¼′ in length, and about as long as the scale. Fruit elongated-cylindric, gradually narrowed into a long thin beak, and raised on a slender stalk sometimes ½′ long.

A bushy tree, occasionally 25° high, with a short trunk 6′—8′ in diameter, stout ascending branches forming a broad round head, and slender branchlets coated at first with hoary deciduous tomentum, varying during their first winter from reddish purple to dark orange-brown, marked by scattered raised lenticels and roughened by conspicuous elevated leaf-scars, growing lighter-colored and reddish brown in their second year; usually much smaller and often shrubby in habit. Bark thin, reddish or olive-green or gray tinged with red, and slightly divided by shallow fissures into appressed plate-like scales. Winter-buds oblong, gradually narrowed and rounded at apex, full and rounded on the back, bright light chestnut-brown, nearly ¼′ long.

Distribution. Borders of streams, swamps, and lakes, hillsides, open woods and forest margins, usually in moist rich soil; valley of the St. Lawrence River to the shores of Hudson’s Bay, the valley of the Mackenzie River within the Arctic Circle, Cook Inlet, Alaska, and the coast ranges of British Columbia, forming in the region west of Hudson’s Bay almost impenetrable thickets, with twisted and often inclining stems; common in all the northern states, ranging southward to Pennsylvania and westward to Minnesota and through the Rocky Mountain region from western Idaho and northern Montana to northern North Dakota, eastern South Dakota, northeastern and central Iowa, and western Nebraska, and southward through Colorado to northern Arizona; ascending as a low shrub in Colorado to an altitude of 10,000°.

21. Salix discolor Muehl. Glaucous Willow.

Leaves lanceolate to elliptic, gradually narrowed at the ends, remotely crenulate-serrate, as they unfold thin, light green often tinged with red, pubescent above and coated with a pale tomentum below, at maturity thick and firm, glabrous, conspicuously reticulate-venulose, bright green above, glaucous or silvery white below, 3′—5′ long, ¾′—1½′ wide, with a broad yellow midrib and slender arcuate primary veins; petioles slender, ½′—1′ long; stipules foliaceous, semilunar, acute, glandular-dentate, about ¼′ long, deciduous. Flowers: aments appearing late in winter or in very early spring, erect, terminal on short scale-bearing branchlets coated with thick white tomentum, oblong-cylindric, about 1′ long and ⅔′ thick, the staminate soft and silky before the flowers open and densely flowered; scales [160] oblong-obovate, dark reddish brown toward the apex, covered on the back with long silky silvery white hairs; stamens 2, with elongated glabrous filaments; ovary oblong-cylindric, narrowed above the middle, villose, with a short distinct style and broad spreading entire stigmas; pedicel glabrous, about twice the length of the scale. Fruit cylindric, more or less contracted above the middle, long-pointed, light brown, coated with pale pubescence.

A tree, rarely more than 25° high, with a trunk about 1° in diameter, stout ascending branches forming an open round-topped head, and stout branchlets marked by occasional orange-colored lenticels, dark reddish purple and coated at first with pale deciduous pubescence; more often shrubby, with numerous tall straggling stems. Winter-buds semiterete, flattened and acute at the apex, about ⅜′ long, dark reddish purple and lustrous. Bark ¼′ thick, light brown tinged with red, and divided by shallow fissures into thin plate-like oblong scales. Wood light, soft, close-grained, brown streaked with red, with lighter brown sapwood.

Distribution. Moist meadows and the banks of streams and lakes; Nova Scotia to Manitoba, and southward to Delaware, southern Indiana and Illinois, eastern and southwestern Iowa, the Black Hills of South Dakota, and northeastern Missouri; common.

A form of Salix discolor with more densely flowered and more silvery pubescent aments is described as var. eriocephala Schn. and a form with loosely flowered aments with less tomentose fruits with longer styles and with narrower leaves as var. prinoides Schn.

22. Salix Scouleriana Barr. Black Willow.

Leaves oblong-obovate to elliptic, acute or abruptly acuminate with a short or long-pointed apex, gradually narrowed and cuneate at the often unsymmetrical base, entire or remotely and irregularly crenately serrate, thin and firm, dark yellow-green and lustrous above, pale or glaucous and glabrous or pilose below, 1¼′—4′ long, ½′—1½′ wide, with a broad yellow pubescent midrib and slender veins forked and arcuate within the slightly thickened and revolute margins and connected by conspicuous reticulate veinlets; petioles slender, puberulous, ¼′—½′ in length; stipules foliaceous, semilunar, glandular-serrate, ⅛′—¼′ long, caducous. Flowers: aments appearing before the leaves, oblong-cylindric, erect, nearly sessile on short tomentose scale-bearing branchlets, the staminate about 1′ long and rather more than ½′ thick, the pistillate 1½′ long, about ⁵/₁₂′ thick; scales oblong, narrowed at the ends, dark-colored, covered with long white hairs, persistent under the fruit; stamens 2, with free [161] glabrous filaments; ovary cylindric, short-stalked, with a distinct style and broad emarginate stigmas; pedicels less than half the length of the scale, villose. Fruit oblong-ovoid, acuminate, light reddish brown, pale pubescent, about ⅓′ long.

A tree, occasionally 30° high, with a short trunk rarely exceeding 1° in diameter, slender pendulous branches forming a rather compact round-topped shapely head, and stout branchlets marked by scattered yellow lenticels, coated when they first appear with pale early deciduous pubescence, becoming bright yellow or dark orange color, and in their second year dark red-brown and much roughened by the conspicuous leaf-scars; or more often a shrub. Winter-buds ovoid, acute, nearly terete or slightly flattened, with narrow lateral wing-like margins, light or dark orange color, glabrous or pilose at the base, about ¼′ long. Bark thin, dark brown slightly tinged with red, and divided into broad flat ridges. Wood light, soft, close-grained, light brown tinged with red, with thick nearly white sapwood.

Distribution. Cook’s Inlet, coast of Alaska, and valley of the Yukon River near Dawson southward through western British Columbia to northern California, ranging eastward through Washington and northwestern Oregon to northern Idaho and Montana.

From central California to San Bernardino County represented by the variety crassijulis Andr. (S. brachystachys Benth.) with shorter and broader obovate leaves rounded at apex, pubescent and tomentose branchlets and larger pubescent winter-buds. A tree sometimes 70° high with a trunk often 2½° in diameter.

On the high Sierra Nevada eastward to the eastern ranges of the Rocky Mountains of Colorado and to northern New Mexico, northern Wyoming and the Black Hills of South Dakota represented by the var. flavescens Schn. A shrub or rarely a small tree with obovate rounded yellowish leaves and branchlets.

23. Salix Hookeriana Barr.

Leaves oblong to oblong-obovate, acute or abruptly acuminate, or rarely rounded and frequently apiculate at apex, gradually narrowed and cuneate or rounded at base, coarsely crenately serrate, especially those on vigorous shoots, or entire, when they unfold villose with pale hairs, or tomentose above and clothed below with silvery white tomentum, at maturity thin and firm, bright yellow-green and lustrous, nearly glabrous or tomentose on the upper surface, pale and glaucous and tomentose or pubescent on the lower surface, especially along the midrib and slender arcuate primary veins and conspicuous reticulate veinlets, 2′—6′ long, 1′—1½′ wide; petioles stout, tomentose, ¼′—⅓′ long. Flowers: aments [162] oblong-cylindric, erect, rather lax, often more or less curved, about 1½′ long, on short tomentose scale-bearing branchlets, the staminate ⅔′ thick and rather thicker than the pistillate; scales oblong-obovate, yellow, coated with long pale hairs, the staminate rounded above and rather shorter than the more acute scales of the pistillate ament persistent under the fruit; stamens 2, with free elongated glabrous filaments; ovary conic, glabrous, stalked, with a slender stalk about one third as long as the scale, gradually narrowed above, with a slender elongated bright red style and broad spreading entire stigmas. Fruit oblong-cylindric, narrowed above, about ¼′ long.

A tree, occasionally 30° high, with a trunk about 1° in diameter, and stout branchlets marked by large scattered orange-colored lenticels, covered during their first season with hoary tomentum and rather bright or dark red-brown and pubescent in their second summer; more often shrubby, with numerous stems 4′—8′ thick and 15°—20° high; frequently a low bush, with straggling almost prostrate stems. Winter-buds ovoid, acute, nearly terete, dark red, coated with pale pubescence, about ¼′ long. Bark nearly ⅛′ thick, light red-brown, slightly fissured and divided into closely appressed plate-like scales. Wood light, soft, close-grained, light brown tinged with red, with thin nearly white sapwood.

Distribution. Borders of salt marshes and ponds and sandy coast dunes; Vancouver Island southward along the shores of Puget Sound and the Pacific Ocean to southern Oregon.

24. Salix sitchensis Sanson.

Leaves oblong-obovate to oblanceolate, entire or minutely glandular dentate, acute or acuminate, or rounded and short-pointed, or rounded at apex, gradually narrowed and cuneate at base, when they unfold pubescent or tomentose on the upper surface, and coated on the lower with lustrous white silky pubescence or tomentum persistent during the season or sometimes deciduous from the leaves of vigorous young shoots, at maturity thin and firm, dark green, lustrous and glabrous above, with the exception of the pubescent midrib, 2′—5′ long, ¾′—1½′ wide, with conspicuous slender veins arcuate and united within the margins and prominent reticulate veinlets; petioles stout, pubescent, rarely ½′ long; stipules rarely produced, foliaceous, semilunar, acute or rounded at apex, glandular-dentate, coated below with hoary tomentum, often ½′ long, caducous. Flowers: aments cylindric, densely flowered, erect on short tomentose leafy branchlets, the staminate 1½′—2′ long and ½′ thick, the pistillate 2½′—3′ long, and ¼′ thick; scales yellow or tawny, the staminate oblong-obovate, rounded at the apex, covered with long white hairs, much longer than the more acute pubescent scales of the pistillate ament; stamen 1, with an elongated [163] glabrous filament, or very rarely 2, with filaments united below the middle or nearly to the apex; ovary short-stalked, ovoid, conic, acute, pubescent and gradually narrowed into the elongated style, with entire or slightly emarginate short stigmas. Fruit ovoid, narrowed above, light red-brown, pubescent about ¼′ long.

A much-branched tree, occasionally 25°—30° high, with a short contorted often inclining trunk sometimes 1° in diameter, and slender brittle branchlets coated at first with hoary tomentum, pubescent and tomentose and dark red-brown or orange color during their first winter, becoming darker, pubescent or glabrous, and sometimes covered with a glaucous bloom in their second season; more often shrubby and 6°—15° tall. Winter-buds acute, nearly terete, light red-brown, pubescent or puberulous, about ¼′ long. Bark about ⅛′ thick and broken into irregular closely appressed dark brown scales tinged with red. Wood light, soft, close-grained, pale red, with thick nearly white sapwood.

Distribution. Banks of streams and in low moist ground; Cook Inlet and Kadiak Island, Alaska, southward in the neighborhood of the coast to Santa Barbara, California; on the Marble Creek of the Kaweah River at 6900° altitude (f. Ralphiana Jeps.)

VI. MYRICACEÆ.

Aromatic resinous trees and shrubs, with watery juice, terete branches, and small scaly buds. Leaves alternate, revolute in the bud, serrate, resinous-punctate, persistent in our species, in falling leaving elevated semiorbicular leaf-scars showing the ends of three nearly equidistant fibro-vascular bundles. Flowers unisexual, diœcious or monœcious, usually subtended by minute bractlets, in the axils of the deciduous scales of unisexual or androgynous simple oblong aments from buds in the axils of the leaves of the year, opening in early spring, the staminate below the pistillate in androgynous aments; staminate, perianth 0; stamens 4 or many, inserted on the thickened base of the scales of the ament; filaments slender, united at the base into a short stipe; anthers ovoid, erect, 2-celled, introrse, opening longitudinally; ovary rudimentary or 0; pistillate flowers single or in pairs; ovary sessile, 1-celled; styles short, divided into 2 elongated filiform stigmas stigmatic on the inner face; ovule solitary, erect from the base of the cell, orthotropous, the micropyle superior. Fruit a globose or ovoid dry drupe usually covered with waxy exudations; nut hard, thick-walled. Seed erect, with a thin coat, without albumen; embryo straight; cotyledons plano-convex, fleshy; radicle short, superior, turned away from the minute basal hilum.

The family consists of the genus Myrica L., of about thirty or forty species of small [164] trees and shrubs, widely distributed through the temperate and warmer parts of both hemispheres. Of the seven North American species three are trees. Wax is obtained from the exudations of the fruit of several species. The bark is astringent, and sometimes used in medicine, in tanning, and as an aniline dye. Myrica rubra Sieb and Zacc., of southern Japan and China, is cultivated for its succulent aromatic red fruit.

The generic name is probably from the ancient name of some shrub, possibly the Tamarisk.

CONSPECTUS OF THE NORTH AMERICAN SPECIES.

1. Myrica cerifera L. Wax Myrtle.

Leaves oblanceolate or rarely oblong-lanceolate, acute or rarely gradually narrowed and rounded at apex, cuneate at base, decurrent on short stout petioles, coarsely serrate above the middle or entire, yellow-green, covered above by minute dark glands and below by bright orange-colored glands, 1½′—4′ long and ¼′—½′ wide, with a slender pale midrib often puberulous below, and few obscure arcuate veins, fragrant with a balsamic resinous odor; gradually deciduous at the end of their first year. Flowers in small oblong aments, with ovate acute ciliate scales, those of the staminate plant ½′—¾′ long, about twice as long as those of the pistillate plant; stamens few, with oblong slightly obcordate anthers at first tinged with red, becoming yellow; ovary gradually narrowed into 2 slender spreading stigmas longer than its scale. Fruit in short spikes, ripening in September and October and persistent on the branches during the winter, irregularly deciduous in the spring and early summer, globose, about ⅛′ in diameter, slightly papillose, light green, coated with thick pale blue wax; seed pale, minute.

A tree, occasionally 40° high, with a tall trunk 8′—10′ in diameter, slender upright or slightly spreading branches forming a narrow round-topped head, and slender branchlets [165] marked by small pale lenticels, coated at first with loose rufous tomentum and caducous orange-colored glands, bright red-brown or dark brown tinged with gray, usually lustrous and nearly glabrous during their first winter, finally becoming dark brown; generally smaller, frequently shrubby. Winter-buds oblong, acute, ¹/₁₆′—⅛′ long, with numerous ovate acute imbricated scales, the inner scales becoming nearly ½′ long, and often persistent until the young branch has completed its growth. Bark of the trunk ¼′ thick, compact, smooth, light gray. Wood light, soft and brittle, dark brown, with thin lighter-colored sapwood.

Distribution. In the neighborhood of the coast; Cape May, New Jersey, southern Delaware and Maryland to the keys of southern Florida, and through the Gulf states to the shores of Aranzas Pass, San Patricio County, Texas, ranging inland to the neighborhood of Natchez, Jackson County, Mississippi, the valley of the Red River (Natchitoches, Louisiana and Fulton, Arkansas), and to Cherokee County, Texas, and northward to the valley of the Washita River, Arkansas; on the Bermuda and Bahama Islands and on several of the Antilles; most abundant and of its largest size on the south Atlantic and Gulf coasts in sandy swamps and pond holes; the most common woody plant and forming great thickets on the Everglades east of Lake Okeechobee, Florida; in the sandy soil of Pine-barrens and on dry arid hills of the interior, often only a few inches in height, var. pumila Michx.

2. Myrica inodora W. Bartr. Wax Myrtle.

Leaves broadly oblong-obovate or rarely ovate, rounded or sometimes pointed and occasionally apiculate at apex, narrowed at base, decurrent on short stout petioles, entire or rarely obscurely toothed toward the apex, thick and coriaceous, glandular-punctate, dark green and very lustrous above, bright green below, 2′—4′ long, ¾′—1½′ wide, with a broad conspicuously glandular midrib slightly pubescent on the lower side, and few remote slender obscure primary veins forked and arcuate near the much-thickened and revolute margins; gradually deciduous from May until midsummer. Flowers in aments ¾′—1′ long, with ovate acute glandular scales; stamens numerous, with oblong slightly emarginate yellow anthers; pistillate flowers usually in pairs, with an ovate glabrous ovary and slender bright red styles. Fruit produced sparingly in elongated spikes, oblong, ⅓′—½′ long, papillose, black, and covered with a thin coat of white wax; seed oblong-oval, acute at apex, rounded at base, ⅛′ long, bright orange-brown, with a pale yellow hilum.

Usually a shrub, with numerous slender stems, occasionally arborescent and 18°—20° high, with a straight trunk 6°—8° tall and 2′—3′ in diameter, and stout branchlets roughened [166] by small scattered lenticels, coated at first with dense pale tomentum, soon becoming bright red-brown, scurfy, and glabrous or pubescent. Winter-buds ovoid, acute, nearly ⅛′ long, with numerous loosely imbricated lanceolate acute red-brown scurfy-pubescent scales. Bark thin, smooth, nearly white.

Distribution. Deep swamps, Round Lake, Jackson County, and Appalachicola, and Saint Andrews Bay, Florida; near Mobile and Stockton, Alabama; near Poplarville, Pearl County, Mississippi, and Bogalusa, Washington Parish, Louisiana.

3. Myrica californica Cham. Wax Myrtle.

Leaves oblanceolate to oblong-lanceolate, acute at apex, remotely serrate except at the gradually narrowed base with small incurved teeth, decurrent on a short stout petiole, thin and firm, dark green and lustrous above, yellow-green, glabrous or puberulous and marked by minute black glandular dots below, 2′—4′ long, ½′—¾′ wide, with a narrow yellow midrib and numerous obscure primary veins arcuate near the thickened and revolute margins, slightly fragrant, gradually deciduous after the end of their first year. Flowers subtended by conspicuous bractlets, those of the two sexes on the same plant; staminate in oblong simple aments often 1′ long, pistillate in shorter aments in the axils of upper leaves, androgynous aments occurring between the two with staminate flowers at their base and pistillate flowers above, or with staminate flowers also mixed with the pistillate at their apex; scales of the aments ovate, acute, coated with pale tomentum; stamens numerous, with oblong slightly emarginate dark red-purple anthers soon becoming yellow; ovary ovoid, with bright red exserted styles. Fruit in short crowded spikes ripening in the early autumn and usually falling during the winter, globose, papillose, dark purple, covered with a thin coat of grayish white wax; seed pale reddish brown, minute.

A tree, occasionally 40° high, with a trunk 14′—15′ in diameter, short slender branches forming a narrow compact round-topped head, and stout branchlets coated at first with loose tomentum, dark green or light or dark red-brown, glabrous or pubescent during their first season, becoming in their second year much roughened by the elevated leaf-scars, darker and ultimately ashy gray; usually smaller at the north and toward the northern and southern limits of its range reduced to a low shrub often only 3°—4° tall. Winter-buds ovoid, acute, about ⅓′ thick, with loosely imbricated ovate acute dark red-brown tomentose scales nearly ½′ long when fully grown and long-persistent on the branch. Bark smooth, compact, ¹/₁₆′—⅛′ thick, dark gray or light brown on the surface and dark red-brown internally. Wood heavy, very hard and strong, brittle, close-grained, light rose color, with thick lighter colored sapwood.

Distribution. Ocean sand-dunes and moist hillsides in the vicinity of the coast from the shores of Puget Sound to the neighborhood of Santa Monica, Los Angeles County, California; of its largest size on the shores of the Bay of San Francisco.

VII. LEITNERIACEÆ.

A tree or shrub, with pale slightly fissured bark, scaly buds, stout terete pithy branchlets marked by pale conspicuous nearly circular lenticels and by elevated crescent-shaped angled or obscurely 3-lobed leaf-scars, very light soft wood, and thick fleshy stoloniferous yellow roots. Leaves involute in the bud, lanceolate to elliptic-lanceolate, acuminate or acute and short-pointed at apex, gradually narrowed at base, entire, with slightly revolute undulate margins, penniveined with remote primary veins arcuate and united near the margins, and conspicuous reticulate veinlets, petiolate, at first coated on the lower surface and on the petioles with thick pale tomentum and puberulous on the upper surface, thick and firm at maturity, bright green and lustrous above, pale and villose-pubescent below, deciduous. Flowers in unisexual aments, with ovate acute concave tomentose scales, the male and female on different plants, opening in early spring from buds formed the previous autumn and covered with acute chestnut-brown hairy scales; the staminate clustered near the end of the branches, their scales bearing on the thickened stipe a ring of 3—12 stamens, with slender incurved filaments and oblong light yellow introrse 2-celled anthers opening longitudinally; perianth 0; pistillate aments scattered, shorter and more slender than the staminate, their scales bearing in their axils a short-stalked pistil surrounded by a rudimentary perianth of small gland-fringed scales, the 2 larger lateral, the others next the axis of the inflorescence; ovary superior, pubescent, 1-celled, with an elongated flattened style inserted obliquely, curving inward above the middle in anthesis, grooved and stigmatic on the inner face; ovule solitary, attached laterally, ascending, semianatropous; micropyle directed upward. Fruit an oblong compressed dry drupe thick and rounded on the ventral, narrowed on the dorsal edge, rounded at base, thin and pointed at apex, chestnut-brown, rugose, with a thick dry exocarp closely investing the thin-walled light brown crustaceous rugose nutlet. Seed flattened, rounded at the ends, light brown, marked on the thick edge with the oblong nearly black hilum; embryo erect, surrounded by thin fleshy albumen; cotyledons oblong, flattened; radicle superior, conical, short, and fleshy.

The family consists of a single genus, Leitneria Chapm., with one species of the southern United States, named for a German naturalist killed in Florida during the Seminole War.

1. Leitneria floridana Chapm. Cork Wood.

Leaves 4′—6′ long, 1½′—2½′ wide, with petioles 1′—2′ in length. Flowers opening at the end of February or early in March; staminate aments 1′—1¼′ long, ¼′ thick, and twice as long as the pistillate. Fruit solitary or in clusters of 2—4, ripening when the leaves are about half grown, ¾′ long, ¼′ wide.

A shrub or small tree, occasionally 20° high, with a slender straight trunk 4′—5′ in diameter above the swollen gradually tapering base, spreading branches forming a loose open head, and branchlets at first light reddish brown and thickly coated with gradually deciduous hairs, becoming in their first winter glabrous or puberulous, especially toward the ends, and dark red-brown. Winter-buds: terminal broad, conic, ⅛′ long, covered by 10 or 12 oblong nearly triangular closely imbricated scales coated with pale tomentum and long-persistent at the base of the branch; lateral scattered, ovoid, flattened. Bark about ¹/₁₆′ thick, dark gray faintly tinged with brown, divided by shallow fissures into narrow rounded ridges. Wood soft, exceedingly light, close-grained, the layers of annual growth hardly distinguishable, pale yellow, without trace of heartwood; occasionally used for the floats of fishing-nets.

Distribution. Borders of swamps of the lower Altamaha River, Georgia (C. L. Boynton); [168] muddy saline shores on the coast of the Gulf of Mexico near Apalachicola, Florida; swampy prairies, Velasco (E. J. Palmer), and swamps of the Brazos River near Columbia, Brazoria County, Texas; Varner, Lincoln County (B. F. Bush), and Moark, Clay County (E. J. Palmer) Arkansas; and in Butler and Dunklin Counties, southeastern Missouri, here sometimes occupying muddy sloughs of considerable extent to the exclusion of other woody plants.

VIII. JUGLANDACEÆ.

Aromatic trees, with watery juice, terete branchlets, scaly buds, the lateral buds often superposed, 2—4 together, and alternate unequally pinnate deciduous leaves with elongated grooved petioles and without stipules, the leaflets increasing in size from the lowest upward, penniveined, sessile, short-stalked or the terminal usually long-stalked. Flowers monœcious, opening after the unfolding of the leaves, the staminate in lateral aments and composed of a 3—6-lobed calyx in the axil of and adnate to an ovate acute bract, and numerous stamens inserted on the inner and lower face of the calyx in 2 or several rows, with short distinct filaments and oblong anthers opening longitudinally; the pistillate in a spike terminal on a branch of the year and composed of a 1—3-celled ovary subtended by an involucre free toward the apex and formed by the union of an anterior bract and 2 lateral bractlets, a 1 or 4-lobed calyx inserted on the ovary, a short style with 2 plumose stigmas stigmatic on the inner face, and a solitary erect orthotropous ovule. Fruit drupaceous, the exocarp (husk) indehiscent or 4-valved, inclosing a thick- or thin-shelled nut divided by partitions extending inward from the shell, and like the shell more or less penetrated by internal longitudinal cavities often filled with dry powder. Seed solitary, 2-lobed from the apex nearly to the middle, light brown, its coat thin, of 2 layers, without albumen; cotyledons fleshy and oily, sinuose or corrugated, 2-lobed; radicle short, superior, filling the apex of the nut. Of the six genera of the Walnut family two occur in North America.

CONSPECTUS OF THE NORTH AMERICAN GENERA.

1. JUGLANS L. Walnut.

Trees, with furrowed scaly bark, durable dark-colored wood, stout branchlets, laminate pith, terminal buds with 2 pairs of opposite more or less open scales often obscurely pinnate at apex, those of the inner pair more or less leaf-like, and obtuse slightly flattened axillary buds formed before midsummer and covered with 4 ovate rounded scales, closed or open during winter. Leaves with numerous leaflets, and terete petioles leaving in falling large conspicuous elevated obcordate 3-lobed leaf-scars displaying 3 equidistant U-shaped clusters of dark fibro-vascular bundle-scars; leaflets conduplicate in the bud, ovate, acute or acuminate, mostly unequal at base, with veins arcuate and united near the margins. Aments of the staminate flowers many-flowered, elongated, solitary or in pairs from lower axillary buds of upper nodes, appearing from between persistent bud-scales in the autumn and remaining during the winter as short cones covered by the closely imbricated bracts of the flowers; calyx 3—6-lobed, its bract free only at the apex; stamens 8—40, in 2 or several ranks, their anthers surmounted by a conspicuous dilated truncate or lobed connective; pistillate flowers in few-flowered spikes, their involucre villose, free only at the apex and variously cut into a laciniate border (corolla?) shorter than the erect calyx-lobes; ovary rarely of 3 carpels; stigmas club-shaped, elongated, fimbriately plumose. Fruit ovoid, globose or pyriform, round or obscurely 4-angled, with a fleshy indehiscent glabrate or hirsute husk; nut ovoid or globose, more or less flattened, hard, thick-walled, longitudinally and irregularly rugose, the valves alternate with the cotyledons, and more or less ribbed along the dorsal sutures and in some species also on the marginal sutures. Seed more or less compressed, gradually narrowed or broad and deeply lobed at base, with conspicuous dark veins radiating from the apex and from the minute basal hilum.

Juglans is confined to temperate North America, the West Indies, South America from Venezuela to Peru, western and northern China, Korea, Manchuria, Japan, and Formosa. Eleven species are known. Of exotic species Juglans regia L., an inhabitant probably originally of China, is cultivated in the middle Atlantic and southern states and largely in California for its edible nuts, which are an important article of commerce. The wood of several species is valued for the interior finish of houses and for furniture.

CONSPECTUS OF THE NORTH AMERICAN SPECIES.

1. Juglans cinerea L. Butternut.

Leaves 15′—30′ long, with stout pubescent petioles, and 11—17 oblong-lanceolate acute or acuminate leaflets 2′—3′ long, 1½′—2′ wide, finely serrate except at the unequal rounded [170] base, glandular and sticky as they unfold, at maturity thin, yellow-green and rugose above, pale and soft-pubescent below; turning yellow or brown and falling early in the autumn. Flowers: staminate in thick aments 3′—5′ long; calyx usually 6-lobed, light yellow-green, puberulous on the outer surface, ¼′ long, its bract rusty-pubescent, acute at apex; stamens 8—12, with nearly sessile dark brown anthers and slightly lobed connectives; pistillate in 6—8-flowered spikes, constricted above the middle, about ⅓′ long, its bract and bractlets coated with sticky white or pink glandular hairs and rather shorter than the linear-lanceolate calyx-lobes; stigmas bright red, ½′ long. Fruit in 3—5 fruited drooping clusters, obscurely 2 or rarely 4-ridged, ovoid-oblong, coated with rusty clammy matted hairs, 1½′—2½′ long with a thick husk; nut ovoid, abruptly contracted and acuminate at apex, with 4 prominent and 4 narrow less conspicuous ribs, light brown, deeply sculptured between the ribs into thin broad irregular longitudinal plates, 2-celled at the base and 1-celled above the middle; seed sweet, very oily, soon becoming rancid.

A tree, occasionally 100° high, with a tall straight trunk 2°—3° in diameter, and sometimes free of branches for half its height; more frequently divided 20° or 30° above the ground into many stout limbs spreading horizontally and forming a broad low symmetrical round-topped head, and dark orange-brown or bright green rather lustrous branchlets coated at first with rufous pubescence, covered more or less thickly with pale lenticels, gradually becoming puberulous, brown tinged with red or orange in their second year and marked by light gray leaf-scars with large black fibro-vascular bundle-scars and elevated bands of pale tomentum separating them from the lowest axillary bud. Winter-buds: terminal ½′—⅔′ long, ¼′ wide, flattened and obliquely truncate at apex, their outer scales coated with short pale pubescence; axillary buds ovoid, flattened, rounded at apex, ⅛′ long, covered with rusty brown or pale pubescence. Bark of young stems and of the branches smooth and light gray, becoming on old trees ¾′—1′ thick, light brown, deeply divided into broad ridges separating on the surface into small appressed plate-like scales. Wood light, soft, not strong, coarse-grained, light brown, turning darker with exposure, with thin light-colored sapwood composed of 5 or 6 layers of annual growth; largely employed in the interior finish of houses, and for furniture. The inner bark possesses mild cathartic properties. Sugar is made from the sap, and the green husks of the fruit are used to dye cloth yellow or orange color.

Distribution. Rich moist soil near the banks of streams and on low rocky hills, southern New Brunswick to the valley of the St. Lawrence River in Ontario, the northern peninsular of Michigan, southern Minnesota, eastern South Dakota, eastern Iowa, southeastern Nebraska, and southward to central Kansas, northern Arkansas, Delaware, eastern [171] Virginia, and on the Appalachian Mountains and their foothills to northern Georgia; in northern Alabama, southern Illinois and western Tennessee; most abundant northward.

× Juglans quadrangulata A. Rehd., a natural hybrid of J. cinerea and the so-called English Walnut (J. regia) is not uncommon in eastern Massachusetts, and a hybrid of J. cinerea with the Japanese J. Sieboldiana Maxm. has appeared in the United States.

2. Juglans nigra L. Black Walnut.

Leaves 1°—2° long, with pubescent petioles, and 15—23 ovate-lanceolate leaflets 3′—3½′ long, 1′—1¼′ wide, long-pointed, sharply serrate except at the more or less rounded often unequal base, thin, bright yellow-green, lustrous and glabrous above, soft-pubescent below, especially along the slender midrib and primary veins; turning bright clear yellow in the autumn before falling. Flowers: staminate in stout puberulous aments 3′—5′ long, calyx rotund, 6-lobed, with nearly orbicular lobes concave and pubescent on the outer surface, its bract ¼′ long, nearly triangular, coated with rusty brown or pale tomentum; stamens 20—30, arranged in many series, with nearly sessile purple and truncate connectives; pistillate in 2—5 flowered spikes, ovoid, gradually narrowed at the apex, ¼′ long, their bract and bractlets coated below with pale glandular hairs and green and puberulous above, sometimes irregularly cut into a laciniate border, or reduced to an obscure ring just below the apex of the ovary; calyx-lobes ovate, acute, light green, puberulous on the outer, glabrous or pilose on the inner surface; stigmas yellow-green tinged on the margins with red, ½′—¾′ long. Fruit solitary or in pairs, globose, oblong and pointed at apex, or slightly pyriform, light yellow-green, roughened by clusters of short pale articulate hairs, 1½′—2′ in diameter, with a thick husk; nut oval or oblong, slightly flattened, 1⅛′—1½′ in diameter, dark brown tinged with red, deeply divided on the outer surface into thin or thick often interrupted irregular ridges, 4-celled at base and slightly 2-celled at the apex; seed sweet, soon becoming rancid.

A tree, frequently 100° and occasionally 150° high, with a straight trunk often clear of branches for 50°—60° and 4°—6° in diameter, thick limbs spreading gradually and forming a comparatively narrow shapely round-topped head of mostly upright rigid branches, and stout branchlets covered at first with pale or rusty matted hairs, dull orange-brown and pilose or puberulous during their first winter, marked by raised conspicuous orange-colored lenticels and elevated pale leaf-scars, gradually growing darker and ultimately light brown. Winter-buds: terminal ovoid, slightly flattened, obliquely rounded at apex, coated with pale silky tomentum, ⅓′ long, with usually 4 obscurely pinnate scales; axillary [172] ⅛′ long, tomentose, their outer scales opening at the apex during the winter. Bark of young stems and branches light brown and covered with thin scales, becoming on old trees 2′—3′ thick, dark brown slightly tinged with red, and deeply divided into broad rounded ridges broken on the surface into thick appressed scales. Wood heavy, hard, strong, rather coarse-grained, very durable, rich dark brown, with thin lighter colored sapwood of 10—20 layers of annual growth; largely used in cabinet-making, the interior finish of houses, gun-stocks, air-planes, and in boat and shipbuilding.

Distribution. Rich bottom-lands and fertile hillsides, western Massachusetts to southern Ontario, southern Michigan, southeastern Minnesota, central and northern Nebraska, central Kansas, eastern Oklahoma, and southward to western Florida, central Alabama and Mississippi, and the valley of the San Antonio River, Texas; most abundant in the region west of the Alleghany Mountains, and of its largest size on the western slopes of the high mountains of North Carolina and Tennessee, and on the fertile river bottom-lands of southern Illinois and Indiana, southwestern Arkansas, and Oklahoma; largely destroyed for its valuable timber, and now rare.

Occasionally cultivated as an ornamental tree in the eastern United States, and in western and central Europe. × Juglans intermedia Carr., a natural hybrid, of J. nigra with the so-called English Walnut (J. regia) has appeared in the United States and Europe, and on the banks of the James River in Virginia has grown to a larger size than any other recorded Walnut-tree. In California a hybrid, known as “Royal,” between J. nigra and J. Hindsii has been artificially produced.

3. Juglans major Hell. Nogal.

Juglans rupestris var. major Torr.
Juglans rupestris Sarg., in part, not Engelm.

Leaves 8′—12′ long, with slender pubescent petioles and rachis, and 9—13 rarely 19 oblong-lanceolate to ovate acuminate often slightly falcate coarsely serrate leaflets cuneate or rounded at base, coated when they first appear with scurfy pubescence, soon becoming glabrous, or at maturity slightly pubescent on the midrib below, 3′—4′, or those of the lower pairs 1½′—2′ long, and 1′—1½′ wide, thin, yellow-green, with a thin conspicuous yellow midrib and primary veins. Flowers: staminate in slender puberulous or pubescent aments 8′—10′ [173] long; calyx nearly orbicular, long-stalked, pale yellow-green, 5 or 6-lobed, the lobes ovate, acute, hoary pubescent on the outer surface, their bract acute, coated with thick pale tomentum; stamens 30—40, with nearly sessile yellow anthers, and slightly divided connectives; pistillate not seen. Fruit subglobose to slightly ovoid or oblong, abruptly contracted at apex into a short point (J. elæopyren Dode), densely tomentose when half grown, 1′—1½′ in diameter, with a thin husk covered with close rufous pubescence; nut dark brown or black, slightly compressed, usually rather broader than high, or ovoid, rounded or bluntly acute at apex, rounded and sometimes depressed at base, longitudinally grooved with broad deep grooves, thick shelled; seed small and sweet.

A tree sometimes 50° high, with a straight trunk occasionally 3°—4° in diameter, or divided at the ground into several large stems, stout branches forming a narrow head, and slender branchlets thickly coated when they first appear with rufous pubescence, becoming red-brown, pubescent or puberulous and marked by many small pale lenticels at the end of their first season and ashy gray the following year.

Distribution. Banks of streams in the cañons of central and southern New Mexico and Arizona, and on Oak Creek near Flagstaff, Arizona on the Colorado plateau (P. Lowell).

4. Juglans rupestris Engelm. Walnut.

Leaves 9′—12′ long, with slender pubescent or puberulous petioles and rachis, and 13—23 narrow lanceolate long-pointed usually falcate finely serrate leaflets entire or nearly entire on their incurved margins, cuneate or rounded at base, thin, light green, glabrous or pubescent on the midrib below, 2′—3′ long and ⅓′—⅔′ wide. Flowers: staminate in slender aments, 3′—4′ long, pubescent when they first appear, becoming glabrous; calyx short-stalked, nearly orbicular, light yellow-green, puberulous on the outer surface, 3—5-lobed with rounded lobes, their bracts ovate-lanceolate, coated with hoary tomentum; stamens about 20, with nearly sessile yellow anthers and slightly lobed connectives; pistillate flowers oblong, narrowed at the ends, thickly coated with rufous pubescence; bract and bractlets irregularly divided into a laciniate border rather shorter than the ovate acute calyx-lobes; stigmas green tinged with red, ⅓′ long. Fruit globose or subglobose, tipped with the persistent remnants of the calyx, pubescent or puberulous with rusty hairs, ½′—¾′ in diameter, with a thin husk; nut subglobose to slightly ovoid, sometimes obscurely 4-ridged from the apex nearly to the middle (J. subrupestris Dode), deeply grooved with longitudinal simple or forked grooves, 4-celled at base, 2-celled at apex, thick shelled; seed small and sweet.

A shrubby round-headed tree occasionally 20°—30° high, with a short generally leaning [174] trunk 18′—30′ in diameter, usually branching from near the ground, and slender branchlets coated with pale scurfy pubescence often persistent for two or three years, orange-red and marked by pale lenticels in their first winter and ultimately ashy gray; often a shrub with clustered stems only a few feet high. Winter-buds: terminal, ¼′—½′ long, compressed, narrowed and often oblique at apex, covered with pale tomentum; axillary ⅛′ long, compressed, coated with pale pubescence. Wood heavy, hard, not strong, rich dark brown with thick white sapwood. The beauty of the veneers obtained from the stumps of the large trees is fast causing their destruction.

Distribution. Limestone banks of the streams of southern, central and western Texas from the Rio Grande to the mountains in the western part of the state; western Oklahoma (Kiowa, Greer, Beckham, Rogel, Mills and Ellis Counties); southeastern New Mexico.

Occasionally cultivated in the eastern United States and in Europe, and hardy as far north as eastern Massachusetts; interesting as producing the smallest nuts of any of the known Walnut-trees.

5. Juglans californica S. Wats.

Leaves 6′—9′ long, with glandular pubescent petioles and rachis, and 11—15, rarely 19, oblong-lanceolate acute or acuminate glabrous finely serrate leaflets cuneate or rounded at base, 1′—2½′ long and ⅓′—¾′ wide, the lower often rounded at apex. Flowers: staminate in slender glabrous or puberulous aments 2′—3′ long; calyx puberulous on the outer surface with acute or rarely rounded lobes, its bract, puberulous; stamens 30—40, with yellow anthers and short connectives bifid at apex; the pistillate subglobose, puberulous; stigmas yellow, ½′ long. Fruit globose, ⅓′—¾′ in diameter, with a thin dark-colored puberulous husk; nut nearly globose, deeply grooved with longitudinal grooves, thick shelled, 4-celled at base, imperfectly 2-celled at apex; seed small and sweet.

A shrubby round-headed tree or shrub generally 12°—20°, rarely 40°—50° high, usually branching from the ground or with a short trunk 1° or rarely 2°—3° in diameter, and slender branchlets coated with scurfy rufous pubescence when they first appear, glabrous, reddish brown and marked by pale lenticels at the end of their first season and gray the following year. Winter-buds coated with rufous tomentum.

Distribution. Banks of streams and bottom-lands in the southern California coast region from Santa Barbara and the Ojai valley to San Fernando and the Sierra Santa Monica, and along the foothills of the Sierra Madre to the San Bernardino Mountains and southward to the Sierra Santa Anna.

A curious seminal variety (var. quercina Babcock) with compound leaves composed of 3 oval leaflets, the terminal long-stalked and 2 or 3 times larger than the lateral leaflets, is occasionally cultivated in California.

6. Juglans Hindsii Rehd.

Juglans californica S. Wats., in part.
Juglans californica var. Hindsii Jep.

Leaves 9′—12′ long, with slender villose pubescent petioles and rachis, and 15—19, usually 19, ovate-lanceolate to lanceolate long-pointed often slightly falcate leaflets, serrate with remote teeth except toward the usually rounded cuneate or rarely cordate base, thin, puberulous above while young, becoming bright green, lustrous and glabrous on the upper surface, below furnished with conspicuous tufts of pale hairs, and villose-pubescent along the midrib and primary veins, 2½′—4′ long and ¾′—1′ wide. Flowers: staminate in slender glabrous or sparingly villose aments 3′—5′ long; calyx elongated, coated like its bract with scurfy pubescence, divided into 5 or 6 acute lobes; stamens 30—40, with short connectives bifid at apex; ovary of the pistillate flower oblong-ovoid, thickly covered with villose pubescence, ⅛′ long, the border of the thin bract and bractlets much shorter than the calyx-lobes; stigma yellow. Fruit globose, 1¼′—2′ in diameter, with a thin dark-colored husk covered with short soft pubescence; nut nearly globose, somewhat flattened at the ends, faintly grooved with remote longitudinal depressions, thick shelled; seed small and sweet.

A tree usually 30°—40°, occasionally 75° high, with a tall trunk 1°—2° in diameter, stout pendulous branches forming a narrow round-topped head, and comparatively slender branchlets thickly coated when they first appear with villose pubescence, reddish brown and puberulous, and marked by pale lenticels and small elevated obscurely 3-lobed leaf scars during their first winter, becoming darker and nearly glabrous in their second year. Winter-buds coated with hoary tomentum; terminal acute, compressed, more or less enlarged at apex, ¼′—⅓′ long; axillary usually solitary, nearly globose, about ¹/₁₆′ in diameter. Bark gray-brown, smoothish, longitudinally fissured into narrow plates. Wood heavy, hard, rather coarse-grained, dark brown often mottled, with thick pale sapwood of from 8 to 10 layers of annual growth.

Distribution. Coast region of central California; banks of the lower Sacramento River; along streams near the western base of Mt. Diablo, and on eastern slope of the Napa Range near Atlas Peak east of Napa Valley; near Loyalton in the Sierra Valley.

Often cultivated in California as a shade tree and as stock on which to graft varieties of Juglans regia L., and rarely in the eastern states and in Europe. In California, a hybrid known as “Paradox” between J. Hindsii and J. regia has been artificially produced.

2. CARYA NUTT. Hickory.

Trees, with smooth gray bark becoming on old trunks rough or scaly, strong hard tough brown heartwood, pale sapwood and tough terete flexible branchlets, solid pith, buds covered with few valvate or with numerous imbricated scales, the axillary buds much smaller than the terminal. Leaves often glandular-dotted, their petioles sometimes persistent on the branches during the winter, and in falling leaving large elevated oblong or semiorbicular more or less 3-lobed emarginate leaf-scars displaying small marginal clusters and central radiating lines of dark fibro-vascular bundle-scars; leaflets involute in the bud, ovate or obovate, usually acuminate, thick and firm, serrate, mostly unequal at base, with veins forked and running to the margins; turning clear bright yellow in the autumn. Aments of the staminate flowers ternate, slender, solitary or fascicled in the axils of leaves of the previous or rarely of the current year, or at the base of branches of the year from the inner scales of the terminal bud, the lateral branches in the axils of lanceolate acute persistent bracts; calyx usually 2 rarely 3-lobed, its bract free nearly to the base and usually much longer than the ovate rounded or acuminate calyx-lobes; stamens 3—10, in 2 or 3 series, their anthers ovate-oblong, emarginate or divided at apex, yellow or red, pilose or hirsute, as long or longer than their slender connectives; pistillate flowers sessile, in 2—10-flowered spikes, with a perianth-like involucre, slightly 4-ridged, unequally 4-lobed at apex, villose and covered on the outer surface with yellow scales more or less persistent on the fruit, the bract much longer than the bractlets and the single calyx-lobe; stigmas short, papillose-stigmatic. Fruit ovoid, globose or pyriform, with a thin or thick husk becoming hard and woody at maturity, 4-valved, the sutures alternate with those of the nut, sometimes more or less broadly winged, splitting to the base or to the middle; nut oblong, obovoid or subglobose, acute, acuminate, or rounded at apex, tipped by the hardened remnants of the style, narrowed and usually rounded at base, cylindric, or compressed contrary to the valves, the shell thin and brittle or thick, hard, and bony, smooth or variously rugose or ridged on the outer surface, 4-celled at base, 2-celled at apex. Seed compressed, variously grooved on the back of the flat or concave lobes, sweet or bitter.

Carya is confined to the temperate region of eastern North America from the valley of the St. Lawrence River to the highlands of Mexico, and to southern China where one species occurs. Of the seventeen species, fifteen inhabit the territory of the United States.

CONSPECTUS OF THE SPECIES OF THE UNITED STATES.

1. Carya pecan Asch. & Gr. Pecan.

Leaves 12′—20′ long, with slender glabrous or pubescent petioles, and 9—17 lanceolate to oblong-lanceolate more or less falcate long-pointed coarsely often doubly serrate leaflets [178] rounded or cuneate at the unequal base; sessile, except the terminal leaflet, or short-stalked, dark yellow-green and glabrous or pilose above, and pale and glabrous or pubescent below, 4′—8′ long, 1′—3′ wide, with a narrow yellow midrib and conspicuous veins. Flowers: staminate in slender puberulous clustered aments 3′—5′ long, from buds formed in the axils of leaves of the previous year or occasionally on shoots of the year, sessile or short-stalked, light yellow-green and hirsute on the outer surface, with broadly ovate acute lobes rather shorter than the oblong or obovate bract; stamens 5′ or 6′; anthers yellow, slightly villose; pistillate in few or many flowered spikes, oblong, narrowed at the ends, slightly 4-angled and coated with yellow scurfy pubescence. Fruit in clusters of 3—11, pointed at apex, rounded at the narrowed base, 4-winged and angled, 1′—2½′ long, ½′—1′ broad, dark brown and more or less thickly covered with yellow scales, with a thin, brittle husk splitting at maturity nearly to the base and often persistent on the branch during the winter after the discharge of the nut; nut ovoid to ellipsoidal, nearly cylindric or slightly 4-angled toward the pointed apex, rounded and usually apiculate at base, bright reddish brown, with irregular black markings with a thin shell and papery partitions; seed sweet, red-brown, its nearly flat lobes grooved from near the base to the apex by 2 deep longitudinal grooves.

A tree, 100°—180° high, with a tall massive trunk occasionally 6° or 7° in diameter above its enlarged and buttressed base, stout slightly spreading branches forming in the forest a narrow symmetrical and inversely pyramidal head, or with abundant room a broad round-topped crown, and branchlets at first slightly tinged with red and coated with loose pale tomentum, becoming glabrous or puberulous in their first winter, and marked by numerous oblong orange-colored lenticels and by large oblong concave leaf-scars with a broad thin membranaceous border surrounding the lower axillary bud. Winter-buds acute, compressed, covered with clusters of bright yellow articulate hairs and pale tomentum; terminal ½′ long; axillary ovoid, often stalked, especially the large upper bud. Bark 1′—1½′ thick, light brown tinged with red, and deeply and irregularly divided into narrow forked ridges broken on the surface into thick appressed scales. Wood heavy, hard, not strong, brittle, coarse-grained, light brown tinged with red, with thin light brown sapwood; less valuable than that of most Hickories, and used chiefly for fuel, and occasionally in the manufacture of wagons and agricultural implements. The nuts, which vary in size and shape and in the thickness of their shells and in the quality of the kernels, are an important article of commerce.

Distribution. Low rich ground in the neighborhood of streams; in the valley of the Mississippi River, Iowa (Clinton and Muscatine Counties), southern Illinois, southwestern [179] Indiana (Sullivan and Spencer Counties), western Kentucky and Tennessee, western Mississippi and Louisiana, extreme western and southwestern Missouri (Jackson County southward, common only on the Marias de Cygne River), eastern Kansas to Kickapoo Island in the Missouri River near Fort Leavenworth, Oklahoma to the valley of the Salt Fork of the Arkansas River (near Alva, Woods County) and to creek valleys near Cache, Comanche County (G. W. Stevens), through Arkansas; and in Texas to the valley of the Devil’s River and to that of Warder’s Creek, Hardiman County; reappearing on the mountains of Mexico; most abundant and of its largest size in southern Arkansas and eastern Texas.

Largely cultivated in the Southern States, in many selected varieties, for its valuable nuts.

2. Carya texana Schn. Bitter Pecan.

Leaves 10′—12′ long, with slender petioles, and 7—13 lanceolate acuminate finely serrate leaflets, hoary-tomentose when they unfold, and more or less villose in the autumn, thin and firm, dark yellow-green and nearly glabrous above, pale yellow-green and puberulous below, 3′—5′ long, about 1½′ wide, the terminal leaflet gradually narrowed to the acute base and short-stalked, the lateral often falcate, unsymmetrical at the base, subsessile or short-stalked. Flowers: staminate in villose aments 2′—3′ long, light yellow-green and villose on the outer surface, with oblong-ovate rounded lobes; pistillate in few fruited spikes, oblong, slightly 4-angled, villose. Fruit oblong or oblong-obovoid, apiculate at apex, slightly 4-winged at base, dark brown, more or less covered with yellow scales, 1½′—2′ long, with a thin husk; nut oblong-ovoid or oblong-obovoid, compressed, acute at the ends, short-pointed at apex, apiculate at base, obscurely 4-angled, bright red-brown, rough and pitted, with a thin brittle shell, thin papery walls, and a low basal ventral partition; seed very bitter, bright red-brown, flattened, its lobes rounded and slightly divided at apex, longitudinally grooved and deeply penetrated on the outer face by the prominent reticulated folds of the inner surface of the shell of the nut.

A tree, sometimes 100° high on the bottoms of the Brazos River, with a tall straight trunk 3° in diameter, and ascending branches, or on the borders of prairies in low wet woods usually 15°—25° tall, with a short trunk 8′—10′ in diameter, small spreading branches forming a narrow round-topped head, and slender branchlets coated at first with thick hoary tomentum sometimes persistent until the autumn, bright red-brown and marked by occasional large pale lenticels during their first winter and by the large concave obcordate leaf-scars nearly surrounding the lowest axillary bud, becoming darker in their second season and dark or light gray-brown in their third year. Winter-buds covered with light [180] yellow articulate hairs; the terminal oblong, acute, or acuminate, somewhat compressed, about ¼′ long, and rather longer than the upper lateral bud. Bark ½′—¾′ thick, light reddish brown, and roughened by closely appressed variously shaped plate-like scales. Wood close-grained, tough and strong, light red-brown, with pale brown sapwood.

Distribution. Bottom-lands and low wet woods; valley of the lower Brazos River, Texas; near Lake Charles, Calcasieu Parish, and Laurel Hill, West Feliciana Parish, Louisiana; near Natchez, Adams County, Mississippi; valley of the Arkansas River (Arkansas Post, Arkansas County, and Van Buren, Crawford County), Arkansas.

3. Carya cordiformis K. Koch. Pignut. Bitternut.

Leaves 6′—10′ long, with slender pubescent or hirsute petioles, and 7—9 lanceolate to ovate-lanceolate or obovate long-pointed sessile leaflets coarsely serrate except at the equally or unequally cuneate or subcordate base, thin and firm, dark yellow-green and glabrous above, lighter and pubescent below, especially along the midrib, 4′—6′ long, ¾′—1¼′ wide, or occasionally 2′—4′ wide (var. latifolia Sarg.). Flowers: staminate in slightly pubescent aments, 3′—4′ long, coated with rufous hairs like its ovate acute bract; stamens 4, with yellow anthers deeply emarginate and villose at apex; pistillate in 1 or 2-flowered spikes, slightly 4-angled, covered with yellow scurfy tomentum. Fruit cylindric or slightly compressed, ¾′—1½′ long, obovoid to subglobose, or oblong and acute at apex (var. elongata Ashe), 4-winged from the apex to about the middle, with a thin puberulous husk, more or less thickly coated with small yellow scales; nut ovoid or oblong, often broader than long, compressed and marked at base with dark lines along the sutures and alternate with them, depressed or obcordate, and abruptly contracted into a long or short point at apex, gray tinged with red or light reddish brown, with a thin brittle shell; seed bright reddish brown, very bitter, much compressed, deeply rugose, with irregular cross-folds.

A tree, often 100° high, with a tall straight trunk 2°—3° in diameter, stout spreading branches forming a broad handsome head, and slender branchlets marked by oblong pale lenticels, bright green and covered more or less thickly with rusty hairs when they first appear, reddish brown and glabrous or puberulous during their first summer, reddish brown and lustrous during the winter and ultimately light gray, with small elevated obscurely 3-lobed obcordate leaf-scars. Winter-buds compressed, scurfy pubescent, bright yellow; terminal ⅓′—¾′ long, oblique at apex, with 2 pairs of scales; lateral 2-angled, often stalked, ⅛′—¼′ long, with ovate pointed slightly accrescent scales keeled on the back. Bark ⅓′—¾′ thick, light brown tinged with red, and broken into thin plate-like scales separating [181] on the surface into small thin flakes. Wood heavy, very hard, strong, tough, close-grained, dark brown, with thick light brown or often nearly white sapwood; largely used for hoops and ox-yokes, and for fuel.

Distribution. Low wet woods near the borders of streams and swamps or on high rolling uplands often remote from streams, southern Maine to Quebec and Ontario, the northern shores of the Lower Peninsula of Michigan, northern Minnesota, southeastern Nebraska, eastern Kansas, eastern Oklahoma, and southward to northwestern Florida, Dallas County, Alabama, and eastern Texas; generally distributed, but not very abundant in all the central states east and west of the Appalachian Mountains; ranging farther north than the other species, and growing to its largest size on the bottom-lands of the lower Ohio basin; the common Hickory of Iowa, Nebraska, and Kansas.

A natural hybrid, × C. Brownii Sarg. of C. cordiformis with C. pecan, with characters intermediate between those of its supposed parents, occurs on bottom-land of the Arkansas River near Van Buren, Crawford County, Arkansas. Probably of the same parentage is the so-called Galloway Nut found in Hamilton County, Ohio. Another hybrid, × C. Brownii var. varians Sarg., probably of the same parentage also, occurs near Van Buren. × C. Laneyi Sarg., a natural hybrid evidently of C. cordiformis with C. ovata, has been found in Rochester, New York, and trees considered varieties of the same hybrid, var. chateaugayensis Sarg., occur near the mouth of the Chateaugay River, Province of Quebec, and at Summertown, Ontario.

4. Carya aquatica Nutt. Water Hickory.

Leaves 9′—15′ long, with slender dark red puberulous or tomentose petioles, and 7—13 ovate-lanceolate long-pointed falcate leaflets symmetrical and rounded or cuneate and unsymmetrical and oblique at base, finely or coarsely serrate, sessile or stalked, 3′—5′ long, ½′—1½′ wide, covered with yellow glandular dots, thin, dark green above, brown and lustrous or tomentose on the lower surface, especially on the slender midrib and primary veins, the terminal leaflet more or less decurrent by its wedge-shaped base on a slender stalk or rarely nearly sessile. Flowers: staminate in solitary or fascicled hirsute aments 2½′—3′ long, covered like their bract with yellow glandular pubescence; stamens 6, with yellow puberulous anthers; pistillate in several flowered spikes, oblong, slightly flattened, 4-angled, glandular-pubescent. Fruit often in 3 or 4-fruited clusters, much compressed, usually broadest above the middle, rounded at the slightly narrowed base, rounded or abruptly narrowed at apex, conspicuously 4-winged, dark brown or nearly black, covered [182] more or less thickly with bright yellow-scales, 1½′ long, 1′—1¼′ wide, with a thin brittle husk splitting tardily and usually only to the middle; nut flattened, slightly obovoid, nearly as broad as long, rounded and abruptly short-pointed at apex, rounded at the narrow base, 4-angled and ridged, dark reddish brown, and longitudinally and very irregularly wrinkled, with a thin shell; seed oblong, compressed, dark brown, irregularly and usually longitudinally furrowed, very bitter.

A tree, occasionally 80°—100° high, with a trunk rarely exceeding 2° in diameter, slender upright branches forming a narrow head, and slender dark reddish brown or ashy gray lustrous branchlets marked by numerous pale lenticels, at first slightly glandular and coated with loose pale tomentum, glabrous or puberulous during the summer, and marked during the winter by small nearly oval or obscurely 3-lobed slightly elevated leaf-scars, growing dark red-brown and ultimately gray. Winter-buds slightly flattened, acute, dark reddish brown, covered with caducous yellow scales; terminal ⅛′—¼′ long, often villose; axillary much smaller, frequently nearly sessile, often solitary. Bark ½′—⅔′ thick, separating freely into long loose plate-like light brown scales tinged with red. Wood heavy, strong, close-grained, rather brittle, dark brown, with thick light-colored or often nearly white sapwood; occasionally used for fencing and fuel.

Distribution. River swamps often inundated during a considerable part of the year from southeastern Virginia southward through the coast regions to the shores of Indian River and the valley of the Suwanee River, Florida, through the maritime portions of the Gulf states to the valley of the Brazos River, Texas, and northward through western Louisiana to southeastern Missouri, and to northeastern Louisiana, western Mississippi, and the valley of the lower Wabash River, Illinois; passing into the var. australis Sarg. with narrower leaflets, smaller ellipsoidal fruit, pale red-brown nuts without longitudinal wrinkles, and with close not scaly bark of the trunk. A large tree in dry sandy soil; high banks of the St. John’s River, near San Mateo, Putnam County, near Jupiter, Palm Beach County, banks of the Caloosahatchie River at Alma, Lee County, and Old Town, Lafayette County, Florida; near Marshall, Harrison County, Texas.

5. Carya myristicæformis Nutt. Nutmeg Hickory.

Leaves 7′—14′ long, with slender terete scurfy-pubescent petioles, and 7—9, occasionally 5, ovate-lanceolate to broadly obovate acute leaflets usually equally or sometimes unequally cuneate or rounded at the narrow base, coarsely serrate, short-stalked or nearly sessile, thin and firm, dark green above, more or less pubescent or nearly glabrous and silvery white and very lustrous below, 4′—5′ long, 1′—1½′ wide, with a pale scurfy pubescent [183] midrib; changing late in the season to bright golden-bronze color and then very conspicuous. Flowers: staminate in aments 3′—4′ long and coated like the ovate-oblong acute bract and calyx of the flower with dark brown scurfy pubescence; stamens 6, with yellow anthers; pistillate oblong, narrowed at the ends, slightly 4-angled, covered with thick brown scurfy pubescence. Fruit usually solitary, ellipsoidal or slightly obovoid, 4-ridged to the base, with broad thick ridges, 1½′ long, coated with yellow-brown scurfy pubescence, the husk not more than ¹/₃₂′ thick, splitting nearly to the base; nut ellipsoidal or sometimes slightly obovoid, 1′ long, ¾′ broad, rounded and apiculate at the ends, smooth, dark reddish brown, and marked by longitudinal broken bands of small gray spots covering the entire surface at the ends with a thick hard and bony shell, a thick partition, and a low thin dorsal division; seed sweet, small, dark brown; the lobes deeply 2-lobed at apex.

A tree, 80°—100° high, with a tall straight trunk often 2° in diameter, stout slightly spreading branches forming a comparatively narrow rather open head, and slender branchlets coated with lustrous golden or brown scales often persistent until the second year, light brown or ashy gray during their first winter, ultimately dark reddish brown, and marked by small scattered pale lenticels and small oval emarginate elevated leaf-scars. Winter-buds covered with thick brown scurvy pubescence; terminal ⅛′—¼′ long, ovoid, rather obtuse; axillary much smaller, acute, slightly flattened, sessile or short-stalked, often solitary. Bark ½′—¾′ thick, dark brown tinged with red, and broken irregularly into small thin appressed scales. Wood hard, very strong, tough, close-grained, light brown, with thick lighter colored sapwood of 80—90 layers of annual growth.

Distribution. Banks of rivers and swamps in rich moist soil or rarely on higher ground; eastern South Carolina, central Alabama, eastern, and northwestern (bluffs of the Yazoo River at Yazoo City) Mississippi, southern Arkansas, western Louisiana, southeastern Oklahoma to Clear Boggy Creek, western Choctaw County, and in Beaumont County, Texas; on the mountains of northeastern Mexico; rare and local; abundant only in southern Arkansas.

6. Carya ovata K. Koch. Shellbark Hickory. Shagbark Hickory.

Leaves 8′—14′ long, with stout glabrous or pubescent petioles, and 5 or rarely 7 ovate to ovate-lanceolate or obovate leaflets, acuminate or rarely rounded at apex, more or less thickly ciliate on the margins, finely serrate except toward the usually cuneate base, dark yellow-green and glabrous above, paler, glabrous and lustrous or puberulous below, the terminal leaflet decurrent on a slender stalk, 5′—7′ long, 2′—3′ wide, rather larger than the sessile or short-stalked upper leaflets, and two or three times as large as those of the lowest [184] pair. Flowers: staminate opening after the leaves have grown nearly to their full size, in slender light green glandular-hirsute aments 4′—5′ long, glandular-hirsute, their elongated ovate-lanceolate acute bract two or three times as long as the ovate concave rounded or acute calyx-lobes; stamens 4, with yellow or red anthers hirsute above the middle; pistillate in 2—5-flowered spikes, ⅓′ long, clothed with rusty tomentum. Fruit solitary or in pairs, subglobose, rather longer than broad or slightly obovoid, depressed at apex, dark reddish brown or nearly black at maturity, roughened by small pale lenticels, glabrous or pilose, 1′—2½′ long, the husk, ⅛′—½′ thick, splitting freely to the base; nut oblong, nearly twice as long as broad, or obovoid and broader than long, compressed, prominently or obscurely 4-ridged and angled, acute and gradually or abruptly narrowed or rounded or nearly truncate at apex, gradually narrowed and rounded at base, pale or nearly white, with a usually thin shell; seed light brown, lustrous, sweet, with an aromatic flavor.

A tree, 70°—90° and occasionally 120° high, with a tall straight trunk 3°—4° in diameter, in the forest often free of branches for 50°—60° above the ground and then divided into a few small limbs forming a narrow head, or with more space sometimes dividing near the ground or at half the height of the tree into stout slightly spreading limbs, forming a narrow inversely conic round-topped head of more or less pendulous branches, and stout branchlets marked with oblong pale lenticels, covered at first with caducous brown scurf and coated with pale glandular pubescence, soon bright reddish brown, and lustrous, glabrous or pubescent, growing dark gray in their second year and ultimately light gray, and marked by pale and slightly elevated ovate semiorbicular or obscurely 3-lobed leaf-scars. Winter-buds: terminal broadly ovoid, rather obtuse, ½′—¾′ long, ⅓′—½′ broad, the 3 or 4 outer scales nearly triangular, acute, dark brown, pubescent and hirsute on the outer surface, the exterior scales often abruptly narrowed into long rigid points and deciduous before the unfolding of the leaves, the inner scales lustrous, covered with resinous glands, yellow-green often tinged with red, oblong-obovate, pointed, becoming 2½′—3′ long and ½′ broad, usually persistent until after the fall of the staminate aments; axillary buds coated at first with thick white tomentum, becoming ⅓′—½′ long when fully grown. Bark light gray, ¾′—1′ thick, separating in thick plates often a foot or more long and 6′—8′ wide, and more or less closely attached to the trunk by the middle, giving it the shaggy appearance to which this tree owes its common name. Wood heavy, very hard and strong, tough, close-grained, flexible, light brown, with thin nearly white sapwood; largely used in the manufacture of agricultural implements, carriages, wagons, and for axe-handles, baskets, and fuel. The nut is the common Hickory nut of commerce.

Distribution. Low hills and the neighborhood of streams and swamps in rich deep moderately moist soil; southern Maine to the valley of the St. Lawrence River near Montreal, along the northern shores of Lakes Erie and Ontario to central Michigan, central Wisconsin, southeastern Minnesota, eastern Iowa and southeastern Nebraska, and southward to western Florida, northern Alabama and Mississippi, and to eastern Kansas, eastern Oklahoma, and eastern Texas; ranging further north than other Hickories with the exception of C. cordiformis; and in the Carolinas ascending to 3000° above the sea in valleys on the western slope of the Blue Ridge. Variable in the size and shape of the nut and in the character and amount of pubescence on the leaves and branchlets. These varieties are distinguished: var. Nuttallii Sarg., with nuts rounded, obcordate or rarely pointed at apex, rounded or abruptly pointed at base, much compressed, and only about ⅗′ long and ⅖′—½′ broad; not rare and widely distributed northward. Var. complanata Sarg., with oblong-obovoid fruit and broadly obovoid much compressed slightly angled nuts cuneate at base and rounded, truncate or slightly obcordate at apex; a single tree on the Drushel Farm near Mt. Hope, Holmes County, Ohio. Var. ellipsoidalis Sarg., with ellipsoidal much compressed nuts abruptly long-pointed at apex, and slender reddish branchlets; near Hannibal, Marion County, and Oakwood, Rolles County, northeastern Missouri, and Indian River, Lewis County, and near Rochester, Munroe County, New York. Var. pubescens Sarg., differing in the dense pubescence of pale fascicled hairs on the young branchlets, and on the petioles, rachis and under surface of the leaflets; bottoms of the [185] Savannah River, Calhoun Falls, Abbeville County, South Carolina, bottom of Little River, Walker County, Georgia, Chattanooga Creek, Hamilton County, Tennessee, Valley Head, DeKalb County, Alabama, and Columbus, Lowndes County, Starkville, Oktibbeha County, and Brookville, Noxubee County, Mississippi. More distinct is

Carya ovata var. fraxinifolia Sarg.

Leaves 7′—9′ long, with slender glabrous or puberulous petioles and 5 lanceolate to slightly oblanceolate acuminate finely serrate leaflets glabrous except on the under side of the midrib, the terminal leaflet 4′—7′ long and 1′—1½′ wide, the lateral sessile, unsymmetrical at base, those of the upper pair often larger than the terminal leaflet, those of the lower pair 2′—2½′ long and 1′—1¼′ wide. Flowers as in the species. Fruit obovoid, usually rounded at apex, compressed, about 1¼′ long, the husk splitting freely to the base, ⅙′—⅕′ in thickness; nut much compressed, rounded at the ends, prominently angled.

A large tree with bark separating in long loose plates, and slender reddish glabrous or puberulous branchlets.

Distribution. Near Rochester, Munroe County, New York; common; near Kingston, Ontario, and westward through Ohio and Indiana; at Keosauqua, Van Buren County, Iowa, and near Myers, Osage County, Oklahoma.

7. Carya carolinæ-septentrionalis Schn. Shagbark Hickory.

Leaves 4′—8′ long, with slender glabrous petioles, and usually 5 but occasionally 3 lanceolate long-pointed leaflets gradually narrowed at the acuminate symmetrical or unsymmetrical base, coarsely serrate, ciliate with long white hairs as the leaves unfold, thin, dark green above, pale yellow-green and lustrous below, the upper leaflets 3′—4′ long, 1′—1½′ wide, and about twice as large as those of the lower pair, turning dull brown or yellow-brown some time before falling. Flowers: staminate in slightly villose aments, glandular-hirsute on the outer surface, with linear elongated acuminate villose bracts; stamens 4; anthers puberulous; pistillate usually in 2-flowered spikes, oblong and covered with clustered golden hairs, their bract linear and ciliate on the margins. Fruit broader than high, or short-oblong, slightly depressed at apex, ¾′—1½′ wide, dark red-brown, roughened by small pale lenticels, the husk ⅛′—⅜′ thick, splitting freely almost to the base; nut ovoid, compressed, prominently 4-angled, acute at ends, nearly white or pale brown, with a thin shell; seed light brown, sweet.

A tree, on moist bottom-lands sometimes 80° tall, with a trunk 2°—3° in diameter, and [186] short small branches forming a narrow oblong head, or on dry hillsides usually not more than 20°—30° tall, with a trunk generally not exceeding a foot in diameter, and slender red-brown branchlets marked by numerous small pale lenticels and by the small low truncate or slightly obcordate leaf-scars, becoming ultimately dull gray-brown. Winter-buds: terminal ovoid, gradually narrowed to the obtuse apex, about ¼′ long, with glabrous bright red-brown and lustrous acute and apiculate strongly keeled spreading outer scales, the inner scales becoming when fully grown bright yellow, long-pointed, and sometimes 2′ long; axillary buds oblong, obtuse, not more than ¹/₁₆′ long. Bark light gray, ¼′—¾′ thick, separating freely into thick plates often a foot or more long, 3′ or 4′ wide, and long-persistent, giving to the trunk the shaggy appearance of the northern Shagbark Hickory. Wood hard, strong, very tough, light reddish brown, with thin nearly white sapwood.

Distribution. Dry limestone hills, river-bottoms and low flat often inundated woods, frequently in clay soil; central North Carolina to northern Georgia, and through western North Carolina to eastern Tennessee, eastern Mississippi, and in Cullman and Dallas Counties, Alabama.

8. Carya laciniosa Schn. Big Shellbark. King Nut.

Leaves 15′—22′ long, with stout glabrous or pubescent petioles often persistent on the branches during the winter, and 5—9, usually 7, ovate to oblong-lanceolate or broadly obovate leaflets, the upper 5′—9′ long and 3′—5′ wide and generally two or three times as large as those of the lowest pair, usually equilateral and acuminate at apex, equally or unequally cuneate or rounded at the often oblique base, finely serrate, sessile or short-stalked, dark green and lustrous above, pale yellow-green or bronzy brown and covered with soft pubescence below. Flowers: staminate in aments 5′—8′ long, glabrous or covered with rufous scurfy tomentum, with linear-lanceolate acute bracts two or three times as long as the broad rounded calyx-lobes; anthers hirsute, yellow, more or less deeply emarginate; pistillate in 2—5-flowered spikes, oblong-ovoid, about twice as long as broad, slightly angled, clothed with pale tomentum, their linear bracts acute much longer than the nearly triangular bractlets and calyx-lobe. Fruit solitary or in pairs, ellipsoidal, ovoid or subglobose, depressed at apex, roughened with minute orange-colored lenticels, downy or glabrous, light orange-colored or dark chestnut-brown at maturity, 1¾′—2½′ long and 1¼′—2′ broad, with a hard woody husk pale and marked on the inside with dark delicate veins, and ¼′—⅓′ thick; nut ellipsoidal or slightly obovoid, longer than broad or sometimes broader than long, flattened and rounded at the ends, or gradually narrowed and rounded at base [187] and occasionally acuminate at apex, more or less compressed, prominently 4-ridged and angled or often 6-ridged, furnished at base with a stout long point, light yellow to reddish brown, 1¼′—2½′ long and 1½′—1¾′ wide, with a hard bony shell sometimes ¼′ thick; seed light chestnut-brown, very sweet.

A tree, occasionally 120° high, with a straight slender trunk often free of branches for more than half its height and rarely exceeding 3° in diameter, comparatively small spreading branches forming a narrow oblong head, and stout dark or light orange-colored branchlets at first pilose or covered with pale or rufous pubescence or tomentum, roughened by scattered elevated long pale lenticels, orange-brown and glabrous or puberulous during their first winter, and marked by oblong 3-lobed emarginate leaf-scars. Winter-buds: terminal ovoid, rather obtuse, sometimes 1′ long and ⅔′ wide, and three or four times as large as the axillary buds, usually covered by 11 or 12 scales, the outer dark brown, puberulous, generally keeled, with a long point at apex, the inner scales obovate, pointed or rounded at apex, light green tinged with red, or bright red or yellow, covered with silky pubescence on the outer face, slightly resinous, becoming 2′—3′ long and 1′ wide. Bark 1′—2′ thick, light gray, separating into broad thick plates frequently 3°—4° long, sometimes remaining for many years hanging on the trunk. Wood heavy, very hard, strong and tough, close-grained, very flexible, dark brown, with comparatively thin nearly white sapwood. The large nuts are often sold in the markets of western cities and commercially are not often distinguished from those of the Shellbark Hickory.

Distribution. Rich bottom-lands usually inundated during several weeks of every year; central and western New York and southeastern Ontario, and westward through southern Ohio, southern Michigan, Indiana and Illinois to southeastern Iowa and southeastern Nebraska, through Missouri and Arkansas to southeastern Kansas and northeastern Oklahoma, and southward through eastern Pennsylvania to western West Virginia; in southeastern Tennessee; banks of the Alabama River, Dallas County, Alabama, and in West Feliciana Parish, Louisiana.

× Carya Nussbaumerii Sarg. with leaves like those of C. laciniosa, slender branchlets, and large fruit of the shape of that of the Pecan but without sutural wings and white or nearly white nuts, believed to be a hybrid of these species, has been found near Fayetteville, St. Clair County, Illinois, at Mt. Vernon, Posey County, Indiana, near Burlington, Des Moines County, Iowa, and from the neighborhood of Rockville, Bates County, Missouri.

Trees intermediate in character between C. laciniosa and C. ovata growing on the bottoms of the Genessee River at Golah, Munroe County, New York, and believed to be hybrids of these species, are × C. Dunbarii Sarg.

9. Carya alba K. Koch. Hickory.

Leaves glandular, resinous, fragrant, 8′—12′ long, with petioles covered like the rachis and the under surface of the leaflets with fascicled hairs, and 5 or 7 oblong-lanceolate to obovate-lanceolate leaflets gradually or abruptly acuminate, mostly equilateral, equally or unequally rounded or cuneate at base, minutely or coarsely serrate, sessile or short-stalked, dark yellow-green and rather lustrous above, lustrous, paler or light orange-colored or brown on the lower surface, the upper leaflets 5′—8′ long and 3′—5′ wide, and two or three times as large as those of the lowest pair. Flowers: staminate in aments 4′—5′ long, with slender light green stems coated with fascicled hairs, pale yellow-green, scurfy-pubescent, with elongated ovate-lanceolate bracts ending in tufts of long pale hairs, and three or four times as long as the calyx-lobes; stamens 4, with oblong bright red hirsute anthers; pistillate in crowded 2—5-flowered spikes, slightly contracted above the middle, coated with pale tomentum, the bract ovate, acute, sometimes ¼′ long, about twice as long as the broadly ovate nearly triangular bractlets and calyx-lobes; stigmas dark red. Fruit ellipsoidal or obovoid, gradually narrowed at the ends, acute at apex, abruptly contracted toward the base, rarely obovoid with a stipe-like base (var. ficoides Sarg.), or ovoid with a long acuminate apex (var. ovoidea Sarg.), pilose or nearly glabrous, dark red-brown, 1½′—2′ long, with a husk about ⅛′ thick splitting to the middle or nearly to the base; nut nearly globose, ellipsoidal, obovoid-oblong or ovoid, narrowed at ends, rounded at base, acute, and sometimes attenuated and long-pointed at apex, much or only slightly compressed, obscurely or prominently 4-ridged, light reddish brown, becoming darker and sometimes red with age, with a very thick hard shell and partitions; in drying often cracking transversely; seed small, sweet, dark brown, and lustrous.

A tree, rarely 100° high, usually much smaller, with a tall trunk occasionally 3° in diameter, comparatively small spreading branches forming a narrow or often a broad round-topped head of upright rigid or of gracefully pendulous branches, and stout branchlets clothed at first with pale fascicled hairs, rather bright brown, nearly glabrous or more or less pubescent, and marked by conspicuous pale lenticels during their first season, becoming light or dark gray, with pale emarginate leaf-scars almost equally lobed, or elongated with the lowest lobe two or three times as long as the others. Winter-buds: terminal broadly ovoid, acute or obtuse, ½′—¾′ long, two or three times as large as the axillary buds, the three or four outer bud-scales ovate, acute, often keeled and apiculate, thick and firm, dark reddish brown and pilose, usually deciduous late in the autumn, the inner scales [189] ovate, rounded or acute and short-pointed at apex, light green covered with soft silky pubescence on the outer, and often bright red and pilose on the inner surface, becoming 1′—1½′ long and ½′ broad. Bark ½′—¾′ thick, close, slightly ridged by shallow irregular interrupted fissures and covered by dark gray closely appressed scales. Wood very heavy, hard, tough, strong, close-grained, flexible, rich dark brown, with thick nearly white sapwood; used for the same purposes as that of the Shellbark Hickory.

Distribution. Eastern Massachusetts southward to Lake County, Florida, and eastern Texas, and through Ohio, southwestern Ontario, southern Michigan, Illinois and Indiana to southeastern Iowa, and through Missouri to eastern Oklahoma; comparatively rare at the north, growing on dry slopes and ridges and less commonly on alluvial bottom-lands; absent from eastern Canada, northern and western New England, and New York except in the neighborhood of the coast; the most abundant and generally distributed Hickory-tree of the southern states, growing to its largest size in the basin of the lower Ohio River and in Missouri and Arkansas; commonly in southern Arkansas and eastern Texas, and occasionally in other southern states represented by var. subcoriacea Sarg., differing in its larger, thicker, more pubescent leaflets, more prominently angled fruit with a thicker husk, larger nuts, and in its longer winter-buds often ⅘′ long and ¾′ in diameter.

× Carya Schneckii Sarg., believed to be a hybrid of C. alba and C. pecan, has been found at Lawrenceville, Lawrence County, Illinois, and near Muscatine, Muscatine County, Iowa.

10. Carya leiodermis Sarg.

Leaves 12′—14′ long, with slender petioles and rachis slightly or densely pubescent with fascicled hairs, becoming glabrous or nearly glabrous, and 7 or rarely 5 thin finely serrate leaflets, long-pointed at apex, and gradually narrowed, cuneate and unsymmetrical at base, at first hoary tomentose below and pubescent above, becoming dark green and lustrous on the upper surface and pale and slightly pubescent on the lower surface, especially on the stout midrib, the terminal oblong-obovate with a stalk ⅕′—⅗′ in length, or nearly sessile, of the same shape and often smaller than the nearly sessile upper leaflets, 4′—5′ long and 2′—2½′ wide, and much larger than the lanceolate lower leaflets. Flowers: staminate opening after the leaves have grown nearly to their full size, in slender puberulous aments [190] 4′—4½′ long; bract of the flower ovate, lanceolate, ciliate on the margins with long white hairs mixed with stipitate glands, a third longer than the ciliate calyx-lobes; stamens 4, anthers red, covered with long rigid white hairs; pistillate in short spikes, their involucre and bracts densely clothed with white hairs. Fruit broadly obovoid, smooth, glabrous or puberulous, covered with scattered white scales, 1½′—1¾′ long, about 1¼′ in diameter, the husk ⅕′ to nearly ¼′ thick, opening freely to the base usually only by two sutures; nut ellipsoidal or slightly obovoid, little compressed, rounded at the ends, tinged with red, with a shell ⅙′—⅕′ thick; seed small and sweet.

A tree 60°—75° tall with a trunk occasionally 3° in diameter, stout often pendulous branches forming a narrow round-topped head, and slender reddish brown lustrous branchlets puberulous or pubescent when they first appear, becoming glabrous or nearly glabrous by the end of their first season. Winter-buds: terminal acute, about ½′ long, the outer scales pubescent, the inner covered with appressed pale hairs and ciliate on the margins; axillary buds ovoid and rounded at apex or subglobose. Bark close, pale, only slightly ridged.

Distribution. Low wet woods; Louisiana to southern Arkansas, and in northwestern Mississippi (bluffs, Yazoo County); most abundant in western Louisiana from the neighborhood of the coast to the valley of Red River, and in Tangipahoa Parish east of the Mississippi River.

Passing into var. callicoma Sarg., differing in the thinner husk of the fruit and in the bright red color of the unfolding leaves.

Distribution. Low wet woods; valley of the Calcasieu River (near Lake Charles), western Louisiana to that of the Neches River (near Beaumont), Texas; in western and southern Mississippi (Warren, Adams, Hinds, Lafayette, Copiah, Lowndes and Oktibbeha Counties).

11. Carya pallida Ashe.

Leaves 7′—15′ long, with slender petioles and rachis covered, like the under side of the midrib, with prominent persistent clusters of fascicled hairs mixed with silvery scales, and usually 7, rarely 9, lanceolate or oblanceolate leaflets, the terminal rarely obovate, finely serrate, resinous, fragrant, acuminate and long-pointed at apex, cuneate or rounded and often unsymmetrical at base, covered in spring with small silvery peltate scales, and at maturity light green and lustrous above, pale and pubescent or puberulous below, the terminal short-stalked or nearly sessile, 4′—6′ long and 1′—2′ wide, and as large or slightly larger than the upper lateral leaflets, those of the lower pairs usually not more than 2′ long and [191] ½′ wide. Flowers: staminate in aments covered with fascicled hairs and silvery scales, 2½′—5′ long, puberulous and glandular on the outer surface, with linear acuminate bracts; stamens 4, anthers hirsute; pistillate usually solitary, oblong, covered with yellow scales, their bract ovate-lanceolate, ciliate on the margin. Fruit pubescent and covered with yellow scales, ellipsoidal to obovoid, broad-obovoid, subglobose to depressed-globose, and from ½′—1½′ in length, with a husk from ⅛′—⅙′ in thickness, splitting tardily to the base by 2 or 3 of the sutures, or occasionally remaining unopened until midwinter; nut white, rounded at the ends, or obcordate or obtusely pointed at apex, compressed, more or less prominently ridged nearly to the base, with a shell ⅛′—¹/₁₂′ thick; seed small and sweet.

A tree occasionally 90°—110° high, with a tall trunk 2½°—3° in diameter, usually not more than 30°—40° tall, with a trunk 12′—18′ in diameter, stout branches, the upper erect, the lower often pendulous, and slender red-brown glabrous or pubescent branchlets. Winter-buds acute or obtuse, reddish brown, puberulous and covered with silvery scales, the terminal ¼′ long with 6—9 scales and rather larger than the lateral buds usually covered with fewer scales. Bark of large trees grown in good soil pale and slightly ridged, that of trees on dry ridges, rough, deeply furrowed, dark gray and southward often nearly black. Wood brown with nearly white sapwood; probably little used except as fuel.

Distribution. Sandy soil in the neighborhood of Cape May, New Jersey, in southern Delaware, and in the southern part of the Maryland peninsula; common in rich soil in Gloucester and James City Counties, Virginia, growing here to its largest size, and southward from southeast Virginia through the Piedmont region of North and South Carolina, ascending to altitudes of 2200° in the mountain valleys of these states; common in northern and central Georgia and southeastern Tennessee, occasionally reaching the Georgia coast and the southwestern part of that state; in western Florida, through northern and central Alabama to Dallas County, and through southern Mississippi to northeastern Louisiana (near Kentwood, Tangipahoa Parish); in Mississippi extending northward to the valley of the Yazoo River in Yazoo County; in northern Tennessee (Lexington, Henderson County); in Alabama the common Hickory on the dry gravelly and poor soils of the upland table-lands and ridges of the central part of the state.

12. Carya glabra Sweet. Pignut.

Leaves 8′—12′ long, with slender glabrous petioles and rachis, and 5 or rarely 7 lanceolate or oblanceolate finely serrate leaflets acuminate at the ends, yellow-green and glabrous above, glabrous, or pubescent on the midrib below, the terminal leaflet sometimes obovate, 4′—4½′ long and 5′ or 6′ wide, and raised on a glabrous or sparingly pubescent stalk, ¼′—½′ in length, the lateral leaflets sessile, those of the upper pair about the size of the terminal leaflet, and two or three times larger than those of the lower pair. Flowers: staminate in short-stalked pubescent aments 2′—2½′ long, yellow-green, the bract villose, much longer than the calyx-lobes; stamens 4, anthers yellow, villose toward the apex; pistillate in few-flowered spikes, oblong, coated with hoary tomentum like the lanceolate acuminate bract. Fruit obovoid, compressed, rounded at apex, gradually narrowed below and often abruptly contracted into a stipe-like base, about 1′ long and ¾′ wide, with a husk from ¹/₁₂′—⅛′ in thickness, opening late by one or two sutures or often remaining closed; nut obovoid, compressed, without ridges, rounded or slightly obcordate at apex, gradually narrowed and rounded below, with a hard thick shell; seed small and sweet.

A tree 60°—90° high, with a trunk 2°—2½° in diameter, with small spreading often drooping branches forming a tall narrow head, and slender glabrous reddish branchlets marked by pale lenticels. Winter-buds ovoid, acute, light brown, glabrous, ⅓′—½′ long and ⅕′—¼′ in diameter, the inner scales covered with close pubescence. Bark close, ridged, light gray. Wood heavy, hard, strong and tough, flexible, light or dark brown, with thick lighter-colored sapwood; used for the handles of tools and in the manufacture of wagons and agricultural implements, and largely for fuel.

Distribution. Hillsides and dry ridges; southwestern Vermont to western New York, southeastern Ontario, southern Indiana and southwestern Illinois, and southward to Delaware, the District of Columbia and eastern Virginia, and along the Appalachian Mountains to North Carolina; in northern, central and eastern Georgia, northern Alabama and eastern Mississippi.

The name “Pignut” usually applied to this tree and to the forms of C. ovalis Sarg., especially in the north, properly belongs to C. cordiformis Schn.

Carya glabra var. megacarpa Sarg.

Leaves 12′—14′ long, with slender glabrous petioles and 5—7 lanceolate to oblanceolate leaflets long-pointed and acuminate at apex, gradually narrowed and unsymmetrical at base, finely serrate, glabrous or very rarely pubescent, often furnished below with small clusters of axillary hairs, the three upper 8′—10′ long and 1½′—2½′ wide and about twice as large as those of the lowest pair. Flowers: staminate in slightly villose aments 2½′—3′ in length, villose, their bract long-pointed, acuminate, villose, twice longer than the calyx-lobes, stamens 4—6, anthers yellow, villose above the middle; pistillate in short-stalked spikes, their involucre only slightly angled, covered with pale yellow hairs, the bract acuminate, twice longer than the bractlets and calyx-lobes. Fruit oblong-obovoid with a stipe-like base to short-obovate and rounded or abruptly cuneate at base, rarely depressed at apex, slightly flattened, often covered with bright yellow scales, 1′—2′ long, 1′—1½′ in diameter, with a husk ⅛′—⅕′ in thickness, opening tardily to the middle usually by one or by two sutures, or often remaining closed; nut broadest toward the rounded apex or oblong and occasionally acute at apex, gradually narrowed and acute at base, often compressed, slightly or rarely prominently angled (f. angulata Sarg.), with a shell ⅛′—⅙′ in thickness; seed small and sweet.

A tree 50°—70° high, with a trunk up to 2° in diameter, stout spreading and drooping branches, and stout or rarely slender glabrous branchlets, reddish brown at the end of their [193] first season, becoming dark gray-brown. Winter-buds ovoid, acute, glabrous, up to ½′ in length, the inner scales puberulous. Bark close, only slightly ridged, light or dark gray.

Distribution. Rochester, Munroe County, New York, through southern Ohio and Indiana to southern Illinois (Tunnel Hill, Johnson County); coast of New Jersey; District of Columbia and southward to the shores of Indian River and the valley of the Callusahatchie River, Florida, and through southern Alabama to western Louisiana; one of the commonest Hickories in the coast region of the south Atlantic and east Gulf states, occasionally ranging inland to central and northern Georgia and western Mississippi.

13. Carya ovalis Sarg.

Leaves 6′—10′ long, with slender petioles often scurfy-pubescent early in the season, soon glabrous, and 7 or rarely 5 lanceolate to oblanceolate, or occasionally obovate finely serrate leaflets, long-pointed and acuminate or rarely rounded at apex, cuneate and unsymmetrical at base, early in the season often scurfy-pubescent and furnished below with small axillary tufts of pale hairs, soon glabrous, the upper 6′ or 7′ long and 1½′—2′ wide, and raised on a stalk ¼′—½′ in length, the lateral sessile, those of the upper pairs as large or slightly smaller than the terminal leaflet. Flowers: staminate in puberulous aments 6′—7′ long, pubescent, their bracts twice longer than the ovate acute calyx-lobes; stamens 4, anthers yellow, thickly covered with pale hairs; pistillate in 1 or 2-flowered spikes, obovoid, more or less thickly covered with yellow scales. Fruit ellipsoidal, acute or rounded at apex, rounded at base, puberulous, 1′—1¼′ long, about ¾′ in diameter, with a husk ¹/₁₂′—¹/₁₀′ in thickness, splitting freely to the base; nut pale, oblong, slightly flattened, rounded at base, acute or acuminate and 4-angled at apex, the ridges extending for one-third or rarely for one-half of its length, with a shell rarely more than ⅕′ in thickness; seed small and sweet.

A tree sometimes 100° high, with a tall trunk occasionally 3° in diameter, small spreading branches forming a narrow often pyramidal head, and slender lustrous red-brown branchlets marked by pale lenticels, often slightly pubescent when they first appear, soon glabrous. Winter-buds ovoid, obtuse, acute or acuminate; the terminal often ½′ long and twice as large as the lateral, the outer scales red-brown, lustrous and glabrous, the inner covered with close pale tomentum. Bark slightly ridged, pale gray, usually separating freely into small plate-like scales, or occasionally close. Wood heavy, hard and tough, flexible, light or dark brown, with thick lighter-colored sapwood; used for the handles of tools, in the manufacture of wagons and agricultural implements, and largely for fuel.

Distribution. Hillsides and rich woods; western New York, eastern Pennsylvania and the District of Columbia to southern Illinois and central Iowa (Ames, Story County), and southward to the mountains of North Carolina and Tennessee, and to central Georgia and Alabama; usually rare and local; most abundant and generally distributed in Indiana. With its varieties usually but incorrectly called “Pignut.”

The following varieties differing in the shape of their fruit are distinguished:

Carya ovalis var. obcordata Sarg.

Fruit subglobose to short-oblong or slightly obovoid, 1′—1¼′ in diameter, with a husk ¹/₁₂′—⅛′ in thickness, splitting freely to the base or nearly to the base by often narrow-winged sutures; nut much compressed, slightly angled and often broadest above the middle, rounded and usually more or less obcordate at apex, narrowed and rounded at base.

Distribution. Southern New England to southern Wisconsin, southwestern Missouri, western North Carolina, central and eastern Georgia, eastern Mississippi and central Alabama; the common and most widely distributed northern variety of Carya ovalis; common in the mountain districts of central Alabama; varying to the f. vestita Sarg. with stouter branchlets covered during their first year with rusty tomentum and more or less pubescent in their second and third seasons, leaflets slightly pubescent below, and with more compressed nuts and puberulous winter-buds. A single tree near Davis Pond, Knox County, Indiana.

Carya ovalis var. odorata Sarg.

Carya microcarpa Darling. in part.
Hicoria microcarpa Britt. in part.
Hicoria glabra var. odorata Sarg. in part.

Fruit subglobose or slightly longer than broad, much flattened, ½′—⅗′ in diameter, with a husk not more than ¹/₂₄′ in thickness, splitting freely to the base by sutures sometimes furnished with narrow wings; nut compressed, rounded at apex, rounded or acute at base, slightly or not at all ridged, pale or nearly white, with a shell ¹/₁₂′ or less in thickness.

Distribution. Southern New England, eastern Pennsylvania and the District of Columbia to western New York, and southeastern Ontario, and through Ohio and Indiana to southern Illinois; near Atlanta, Georgia, and Starkville, Oktibbaha County, Mississippi; less variable in the size and shape of the fruit than the other varieties of C. ovalis.

Carya ovalis var. obovalis Sarg.

Fruit more or less obovoid, about 1′ long and ⅘′ in diameter, with a husk ¹/₁₂′—⅛′ thick, splitting freely to the base.

Distribution. Southern New England to Missouri and northern Arkansas; on the mountains of North Carolina, on the coast of Georgia and in north central Alabama. The common “Pignut” in the middle western states, varying to f. acuta Sarg. with nuts pointed at the ends and closer bark; only near Rochester, Munroe County, New York.

Other forms of C. ovalis are var. hirsuta Sarg. (Hicoria glabra hirsuta Ashe) with obovoid compressed fruit narrowed into a stipitate base, with a husk ¹/₁₂′—⅛′ in thickness, scaly bark, pubescent winter-buds, leaves with pubescent petioles and leaflets pubescent on the lower surface; a common tree on the mountains of North Carolina up to altitudes of 2000° above the sea; and var. borealis Sarg. (Hicoria borealis Ashe) with pubescent branchlets [196] and winter-buds, leaves pubescent early in the season, ellipsoidal or ovoid flattened fruit with a husk ⅛′—⅕′ in thickness, an ovoid nut ridged to the base, and scaly bark; only in southeastern Michigan.

14. Carya floridana Sarg.

Leaves 6′—8′ long, with slender petioles rusty pubescent when they first appear, soon glabrous, with 5 or rarely 7 lanceolate to oblanceolate leaflets long-pointed and acuminate at apex, unsymmetrical and rounded or cuneate at base, serrate with remote cartilaginous teeth, sessile or the terminal leaflet short-stalked, covered when they unfold with rufous pubescence, soon glabrous, at maturity thin, conspicuously reticulate-venulose, yellow-green above, often brownish below, the upper three 3½′—4′ long, 1′—2′ wide, and about twice larger than those of the lowest pair. Flowers: staminate in long-stalked scurfy pubescent aments 1′—1½′ in length, produced at the base of branchlets of the year from the axils of [197] bud-scales, and often of leaves, scurfy pubescent, their bract ovate, acuminate, a third longer than the calyx-lobes; stamens 4 or 5, anthers yellow, slightly villose near the apex; pistillate in 1 or 2-flowered spikes, obovoid, thickly covered, like their bracts, with yellow scales. Fruit obovoid, gradually narrowed, rounded and sometimes slightly depressed at apex, narrowed below into a short stipe-like base, occasionally slightly winged at the sutures, often roughened by prominent reticulate ridges, puberulous and covered with small yellow scales, ⅘′—1½′ long, ¾′—1′ in diameter with a husk ¹/₁₂′—⅛′ thick, splitting freely to the base by 2 or 3 sutures; nut pale or reddish, subglobose, not more than ⅗′ in diameter, or ovoid or rarely oblong, acute at base, narrowed and rounded at apex, slightly compressed, with a shell ¹/₁₂′—⅛′ in thickness.

A tree 50°—70° high with a trunk up to 20′ in diameter, slender spreading branches forming a broad head, and slender branchlets at first coated with rufous pubescence, soon puberulous or glabrous, bright red-brown and marked by pale lenticels during their first winter; or in dry sand often a shrub producing abundant fruit on stems 3° or 4° high. Winter-buds ovoid, acute or obtuse, the outer scales covered with thick rusty pubescence and more or less thickly with yellow or rarely silvery scales, the inner coated with pale pubescence; the terminal ⅕′—⅓′ in length and twice as large as the axillary buds. Bark slightly ridged, close dark gray-brown. Wood dark brown, with pale sapwood; probably used only for fuel.

Distribution. Dry sandy ridges and low hills, Florida; east coast, Volusia County to Jupiter Island, Palm Beach County; in the interior of the peninsula as a shrub, from Orange to De Soto Counties, and on the shores of Pensacola Bay.

15. Carya Buckleyi Durand.

Leaves 8′—12′ long, with slender petioles rusty pubescent and sparingly villose early in the season, and 5—7, usually 7, lanceolate to oblanceolate acuminate bluntly serrate sessile leaflets, the terminal occasionally broadly obovate and abruptly pointed, and sometimes raised on a winged stalk ¼—½′ in length, when they unfold thickly covered with rusty pubescence mixed with small white scales and villose on the lower side of the midrib and veins, [198] and at maturity dark green, lustrous, glabrous or puberulous along the midrib above, paler, glabrous or sparingly villose and furnished with small tufts of axillary hairs below, the upper three leaflets 4′—6′ long and 2′—2¼′ wide, and twice the size of those of the lowest pair. Flowers: staminate in rusty pubescent aments 2′—3′ long, their bract slender, long acuminate, 3 or 4 times longer than the acuminate calyx-lobes; stamens 4 or 5, anthers yellow, slightly villose toward the apex; pistillate in 1 or 2-flowered short-stalked spikes, slightly angled, thickly coated with rufous hairs like the bract and bractlets. Fruit subglobose, puberulous, 1¼′—1¾′ in diameter, with a husk ¹/₁₂′—⅛′ thick, splitting freely to the base by slightly winged sutures; nut slightly compressed, rounded at base, abruptly narrowed and acute at apex, 4-angled above the middle or nearly to the base, dark reddish brown, conspicuously reticulate-venulose with pale veins, with a shell about ⅛′ thick; in drying often cracking longitudinally between the angles; seed small and sweet.

A tree, usually 30°—45° or rarely 60° high, with a trunk 12′—24′ in diameter, large spreading often drooping more or less contorted branches forming a narrow head, and slender light red-brown branchlets marked by pale lenticels, more or less densely rusty pubescent during their first season and dark gray-brown and glabrous or nearly glabrous the following year. Winter-buds ovoid, covered with rusty pubescence mixed with silvery scales, furnished at apex with long pale hairs; the terminal bud abruptly contracted and long-pointed at apex, ⅖′—½′ in length and ¼′—⅓′ in diameter, and 2 or 3 times larger than the flattened acute lateral buds. Bark thick, deeply furrowed, rough, dark often nearly black. Wood hard, brittle, little used except for fuel.

Distribution. Dry sandy uplands with Post and Black Jack Oaks; northern and eastern Texas (Grayson, Cherokee, San Augustine and Atascosa Counties), and in central Oklahoma (dry sand hills, Muskogee County).

Carya Buckleyi var. arkansana Sarg.

Differing from Carya Buckleyi in the shape of the fruit and sometimes in the bark of the trunk. Fruit obovoid, rounded at apex, rounded or gradually narrowed or abruptly contracted into a more or less developed stipe at base, or ellipsoidal, or ovoid and rounded at the ends, ⅘′—1½′ in length and in diameter, with a husk ¹/₁₂′—⅙′ thick, splitting to the middle or nearly to the base by slightly winged sutures; nut oblong to slightly obovoid, rounded at [199] the ends, compressed, slightly 4-angled occasionally to the middle, pale brown, with a shell ⅙′—⅕′ in thickness; seed small and sweet.

A tree from 60°—75° high, with a trunk 2° in diameter; southward usually much smaller. Bark on some trees dark gray, irregularly fissured, separating into thin scales, and on others close, nearly black and deeply divided into rough ridges.

Distribution. Dry hillsides, rocky ridges, or southward on sandy upland; southwestern Indiana (Knox County), southern Illinois, northeastern Missouri and southward through Missouri and Arkansas to eastern Oklahoma, western Louisiana and northern and eastern Texas to the valley of the Atascosa River, Atascosa County; the common Hickory of the Ozark Mountain region, Arkansas, and here abundant on dry rocky ridges at altitudes of 1200°—1800°; in Texas the common Hickory from the coast to the base of the Edwards Plateau; trees with the smallest fruit northward; those with the largest fruit with thickest husks in Louisiana, and in southern Arkansas (f. pachylemma Sarg.), a tree with slender nearly glabrous branchlets, deeply fissured pale gray bark, rusty pubescent winter-buds and fruit 2½′ long and 2′ in diameter, with a husk ½′ in thickness.

Carya Buckleyi var. villosa Sarg.

Hicoria glabra var. villosa Sarg.
Hicoria villosa Ashe.
Carya villosa Schn.
Carya glabra var. villosa Robins.

Leaves 6′—10′ long, with slender petioles and rachis pubescent with fascicled hairs early in the season, generally becoming glabrous, and 5—7, usually 7, lanceolate to oblanceolate finely serrate leaflets long-pointed and acuminate at apex, cuneate or rounded and often unsymmetrical at base, sessile or the terminal leaflet sometimes short-stalked, dark green and glabrous above, pale and pubescent below, the lower side of the midrib often covered with fascicled hairs, the upper leaflets 3′—4′ long and 1′—1½′ wide, and twice as long as those of the lowest pair. Flowers: staminate in aments pubescent with fascicled hairs, 4′—8′ long, pubescent, their bract acuminate, not much longer than the rounded calyx-lobes; pistillate in 1 or 2-flowered spikes, rusty pubescent, slightly angled. Fruit obovoid to ellipsoidal, rounded at apex, cuneate and often abruptly narrowed into a stipitate base, rusty pubescent and covered with scattered yellow scales, about 1′ long and ¾′ in diameter, with a husk ¹/₁₂′ in thickness, splitting tardily to the base by 1 or 2 sutures or indehiscent; nut ovoid, rounded [200] at base, pointed at apex, only slightly angled, faintly tinged with red, with a shell rarely more than ¹/₁₂′ in thickness; seed small and sweet.

A tree 30°—40° high, with a trunk 12′—18′ in diameter, stout often contorted branches and slender branchlets covered at first with rusty pubescence mixed with fascicled hairs and pubescent or glabrous during their first winter. Winter-buds ovoid, acute, covered with rusty pubescence mixed with yellow scales, often furnished near the apex with tufts of white hairs, the terminal ¼′ long and about twice as large as the compressed axillary buds.

Distribution. Dry rocky hills, Allenton, Saint Louis County, Missouri. Distinct from other forms of Carya Buckleyi in the often indehiscent fruit and more numerous and longer fascicled hairs, and possibly better considered a species.

IX. BETULACEÆ.

Trees, with sweet watery juice, without terminal buds, their slender terete branchlets marked by numerous pale lenticels and lengthening by one of the upper axillary buds formed in early summer, and alternate simple penniveined usually doubly serrate deciduous stalked leaves, obliquely plicately folded along the primary veins, their petioles in falling leaving small semioval slightly oblique scars showing three equidistant fibro-vascular bundle-scars; stipules inclosing the leaf in the bud, fugacious. Flowers vernal, appearing with or before the unfolding of the leaves, or rarely autumnal, monœcious, the staminate 1—3 together in the axils of the scales of an elongated pendulous lateral ament and composed of a 2—4-parted membranaceous calyx and 2—20 stamens inserted on a receptacle, with distinct filaments and 2-celled erect extrorse anthers opening longitudinally, or without a calyx, the pistillate in short lateral or capitate aments, with or without a calyx, a 2-celled ovary, narrowed into a short style divided into two elongated branches longer than the scales of the ament and stigmatic on the inner face or at the apex, and a single anatropous pendulous ovule in each cell of the ovary. Fruit a small mostly 1-celled 1-seeded nut, the outer layer of the shell light brown, thin and membranaceous, the inner thick, hard, and bony. Seed solitary by abortion, filling the cavity of the nut, suspended, without albumen, its coat membranaceous, light chestnut-brown; cotyledons thick and fleshy, much longer than the short superior radicle turned toward the minute hilum.

Of the six genera, all confined to the northern hemisphere, five are found in North America; of these only Corylus is shrubby.

CONSPECTUS OF THE NORTH AMERICAN ARBORESCENT GENERA.

1. CARPINUS L. Hornbeam.

Trees, with smooth close bark, hard strong close-grained wood, elongated conic buds covered by numerous imbricated scales, the inner lengthening after the opening of the buds. Leaves open and concave in the bud, ovate, acute, often cordate; stipules strap-shaped to oblong-obovate. Flowers: staminate in aments emerging in very early spring from buds produced the previous season near the ends of short lateral branchlets of the year and inclosed during the winter, composed of 3—20 stamens crowded on a pilose receptacle adnate to the base of a nearly sessile ovate acute coriaceous scale longer than the stamens; filaments short, slender, 2-branched, each branch bearing a 1-celled oblong yellow half-anther hairy at the apex; pistillate in lax semi-erect aments terminal on leafy branches of the year, in pairs at the base of an ovate acute leafy deciduous scale, each flower subtended by a small acute bract with two minute bractlets at its base; calyx adnate to the ovary and dentate on the free narrow border. Nut ovoid, acute, compressed, conspicuously longitudinally ribbed, bearing at the apex the remnants of the calyx, marked on the broad base by a large pale scar and separating at maturity in the autumn from the leaf-like 3-lobed conspicuously serrate green involucre formed by the enlargement of the bract and bractlets of the flower and inclosing only the base of the nut, fully grown at mid-summer and loosely imbricated into a long-stalked open cluster. (Eucarpinus.)

Carpinus is confined to the northern hemisphere, and is distributed from the Province of Quebec through the eastern United States to the highlands of Central America in the New World, and from Sweden to southern Europe, Asia Minor, the temperate Himalayas, Korea, southern China, Japan and Formosa in the Old World. Fifteen or sixteen species are recognized. Of the exotic species, the European and west Asian Carpinus Betulus L. is frequently planted as an ornamental tree in the northeastern United States, where some of the species of eastern Asia promise to become valuable.

1. Carpinus caroliniana Walt. Hornbeam. Blue Beech.

Leaves often somewhat falcate, long-pointed, sharply doubly serrate with stout spreading glandular teeth, except at the rounded or wedge-shaped often unequal base, pale bronze-green, and covered with long white hairs when they unfold, at maturity thin and firm, pale dull blue-green above, light yellow-green and glabrous or puberulous below, with small tufts of white hairs in the axils of the veins, 2′—4′ long, 1′—1¾′ wide, with a slender yellow midrib, numerous slender veins deeply impressed and conspicuous above, and prominent cross veinlets; turning deep scarlet and orange color late in the autumn; [202] petioles slender, terete, hairy, about ⅓′ long, bright red while young; stipules ovate-lanceolate, acute, pubescent, hairy on the margins, bright red below, light yellow-green at the apex, ⅓′ long. Flowers: staminate aments 1½′ long when fully grown, with broadly ovate acute boat-shaped scales green below the middle, bright red above; pistillate aments ½′—¾′ long, with ovate acute hairy green scales; styles scarlet. Fruit: nut ⅓′ long, its involucre short-stalked, with one of the lateral lobes often wanting, coarsely serrate, but usually on one margin only of the middle lobe, 1′—1½′ long, nearly 1′ wide, crowded on slender terete pubescent red-brown stems 5′—6′ in length.

A bushy tree, rarely 40° high, with a short fluted trunk occasionally 2° in diameter, long slightly zigzag slender tough spreading branches pendulous toward the ends, and furnished with numerous short thin lateral branches growing at acute angles, and branchlets at first pale green coated with long white silky hairs, orange-brown and sometimes slightly pilose during the summer, becoming dark red and lustrous during their first winter and ultimately dull gray tinged with red. Winter-buds ovoid, acute, about ⅛′ long, with ovate acute chestnut-brown scales white and scarious on the margins. Bark light gray-brown, sometimes marked with broad dark brown horizontal bands, ¹/₁₆′—⅛′ thick. Wood light brown, with thick nearly white sapwood; sometimes used for levers, the handles of tools, and other small articles.

Distribution. Borders of streams and swamps, generally in deep rich moist soil; Nova Scotia and southern and western Quebec to the northern shores of Georgian Bay, southward to the shores of Indian River and those of Tampa Bay, Florida, and westward to central Minnesota, eastern Iowa (Sharpy County), eastern Nebraska (reported), eastern Kansas, eastern Oklahoma, and eastern Texas; reappearing on the mountains of southern Mexico and Central America; common in the eastern and central states; most abundant and of its largest size on the western slopes of the southern Alleghany Mountains and in southern Arkansas and eastern Texas.

2. OSTRYA Scop. Hop Hornbeam.

Trees, with scaly bark, heavy hard strong close-grained wood, and acute elongated winter-buds formed in early summer and covered by numerous imbricated scales, the inner lengthening after the opening of the bud. Leaves open and concave in the bud; petioles slender, nearly terete, hairy; stipules strap-shaped to oblong-obovate. Flowers: staminate in long clustered sessile or short-stalked aments developed in early summer from lateral buds near the ends of short lateral branchlets of the year and coated while young with hoary tomentum, naked and conspicuous during the winter, and composed of 3—14 stamens crowded on a pilose receptacle adnate to the base of an ovate concave scale rounded and abruptly short-pointed at the apex, ciliate on the margins, longer than the stamens; filaments short, 2-branched, each branch bearing a 1-celled half-anther hairy at the apex; pistillate in erect lax aments terminal on short leafy branches of the year, in pairs at the base of an elongated ovate acute leaf-like ciliate scale persistent until midsummer, each flower inclosed in a hairy sack-like involucre formed by the union of a bract and 2 bractlets; calyx adnate to the ovary, denticulate on the free narrow border. Nut ovoid, acute, flattened, obscurely longitudinally ribbed, crowned with the remnants of the calyx, marked at the narrow base by a small circular pale scar, inclosed in the much enlarged pale membranaceous conspicuously longitudinally veined reticulate-venulose involucres of the flower, short, pointed and hairy at the apex, hirsute at the base, with sharp rigid stinging hairs, imbricated into a short strobile fully grown at midsummer, and suspended on a slender hairy stem.

Ostrya is widely distributed in the northern hemisphere from Nova Scotia to Texas, northern Arizona, and to the highlands of southern Mexico and Guatemala in the New World, and through southern Europe and southwestern Asia, and in northern Japan and on the Island of Quelpart in the Old World. Of the four species now recognized two are North American.

CONSPECTUS OF THE NORTH AMERICAN SPECIES.

1. Ostrya virginiana K. Koch. Hop Hornbeam. Ironwood.

Leaves oblong-lanceolate, gradually narrowed into a long slender point or acute at apex, narrowed and rounded, cordate, or wedge-shaped at the often unequal base, sharply serrate, with slender incurved callous teeth terminating at first in tufts of caducous hairs, when they unfold light bronze-green, glabrous above and coated below on the midrib and primary veins with long pale hairs, at maturity thin and extremely tough, dark dull yellow-green above, light yellow-green and furnished with conspicuous tufts of pale hairs in the axils of the veins below, 3′—5′ long, 1½′—2′ wide, with a slender midrib impressed and puberulous above, light yellow and pubescent below, and numerous slender veins forked near the margins; turning clear yellow before falling in the autumn; petioles hairy about ¼′ long; stipules rounded and often short-pointed at apex, ciliate on the margins with long pale hairs, hairy on the back, about ½′ long and ⅛′ wide. Flowers: staminate aments about ½′ long during their first season, with light red-brown rather loosely imbricated scales narrowed into a long slender point, becoming when the flowers open 2′ long, with broadly obovate scales rounded and abruptly contracted at apex into a short point, ciliate on the margins, green tinged with red above the middle, light brown toward the base; pistillate aments slender, about ¼′ long, on thin hairy stems, their scales lanceolate, acute, light green, often flushed with red above the middle, hirsute at the apex, decreasing in size from the lowest. Fruit: nuts ⅓′ long, about ⅛′ wide, rather abruptly narrowed below the apex, their involucres in clusters 1½′—2′ long and ⅔′—1′ wide, on slender hairy stems about 1′ in length.

A tree, occasionally 50°—60° high, with a short trunk 2° in diameter, usually not more than 20°—30° tall, with a trunk 18′—20′ thick, long slender branches drooping at the ends and forming a round-topped or open head frequently 50° across, and slender, very tough branchlets, light green, coated with pale appressed hairs when they first appear, becoming light orange color and very lustrous by midsummer, glabrous, dark red-brown and lustrous during their first winter, and then growing gradually darker brown and losing their lustre; or covered like the petioles and peduncles with short erect glandular hairs (var. glandulosa Sarg.). [204] Winter-buds ovoid, light chestnut-brown, slightly puberulous, ¼′ long. Bark about ¼′ thick, broken into thick narrow oblong closely appressed plate-like light brown scales slightly tinged with red on the surface. Wood strong, hard, tough, durable, light brown tinged with red or often nearly white, with thick pale sapwood of 40—50 layers of annual growth; used for fence-posts, handles of tools, mallets, and other small articles.

Distribution. Dry gravelly slopes and ridges often in the shade of oaks and other large trees; Island of Cape Breton and the shores of the Bay of Chaleur, through the valley of the St. Lawrence River, and along the northern shores of Lake Huron to western Ontario, Manitoba, Minnesota, eastern North Dakota, the foothills of the Black Hills of South Dakota, eastern, northern and northwestern Nebraska, eastern Kansas and Oklahoma, and southward to northern Florida and eastern Texas; most abundant and of its largest size in southern Arkansas and in Texas. From Quebec and Ontario to western New England, western New York, Ohio and in Central Michigan, the glandular form prevails: the two forms occur in New Jersey, Pennsylvania, Indiana, northern Illinois, southwestern Missouri, Oklahoma, and southward on the high Appalachian Mountains.

2. Ostrya Knowltonii Cov. Ironwood.

Leaves elliptic to obovate, acute or round at apex, gradually narrowed and often unequal at the rounded cuneate rarely cordate base, sharply serrate with small triangular callous teeth, covered with loose pale tomentum when they unfold, at maturity dark yellow-green and pilose above, pale and soft-pubescent below, 1′—2′ long, 1′—1½′ wide, with a slender yellow midrib slightly raised on the upper side, and slender primary veins connected by obscure reticulate veinlets; turning dull yellow in the autumn before falling; petioles ⅛′—¼′ long; stipules pale yellow-green, often tinged with red toward the apex, ½′ long, about ½′ wide. Flowers: staminate aments on stout stalks covered with rufous tomentum and sometimes ½′ long, rarely sessile, about ½′ long during their first season, with dark brown puberulous scales gradually contracted into a long slender subulate point, becoming when the flowers open 1′—1¼′ long, with broadly ovate concave scales abruptly narrowed into a nearly triangular point, yellow-green near the base, bright red above the middle; pistillate aments about ¼′ long, with ovate-lanceolate light yellow-green puberulous scales ciliate on the margins. Fruit: nuts ¼′ long, gradually narrowed at the apex, their involucres 1′ long, nearly glabrous at the apex, sometimes slightly stained with red toward the base, in clusters 1′—1½′ long and about ¾′ broad, on stems ½′ in length.

A tree 20°—30° high, with a trunk 12′—18′ in diameter, usually divided 1° or 2° above the ground into 3 or 4 stout upright stems 4′—5′ thick, slender pendulous often much contorted branches forming a narrow round-topped symmetrical head, and slender branchlets dark green and coated with hoary tomentum when they first appear, dark red-brown and pubescent during their first summer, becoming light cinnamon-brown, glabrous, and lustrous in the winter, and ultimately ashy gray. Winter-buds ovoid, dark brownish red, about ⅛′ long. Bark internally bright orange color, ⅛′ thick, separating into loose hanging plate-like scales light gray slightly tinged with red, and 1′—2′ long and wide. Wood light reddish brown, with thin sapwood.

Distribution. On the southern slope of the cañon of the Colorado River in Coconino County, Arizona, at altitudes of 6000°—7000° above the sea (Hance trail, seventy miles north of Flagstaff); in the cañon of Oak Creek, south of Flagstaff (P. Lowell); and on Grand River, Utah (Moab, Grant County, M. E. Jones).

3. BETULA L. Birch.

Trees, with smooth resinous bark marked by long longitudinal lenticels, often separating freely into thin papery plates, becoming thick, deeply furrowed, and scaly at the base of old trunks, short slender branches more or less erect and forming on young trees a narrow symmetrical pyramidal head, becoming horizontal and often pendulous on older trees, tough branchlets, short stout spur-like 2-leaved lateral branchlets much roughened by the crowded leaf-scars of many years, and elongated winter-buds covered by numerous ovate acute scales, and fully grown and bright green at midsummer. Leaves open and convex in the bud, often incisely lobed; stipules ovate and acute or oblong-obovate, scarious. Flowers in 3-flowered cymes, the lateral flowers of the cyme subtended by bractlets adnate to the base of the scale of the ament; staminate aments long, pendulous, solitary or clustered, appearing in summer or autumn in the axils of the last leaves of a branchlet or near the ends of short lateral branchlets, erect and naked during the winter, their scales in the spring broadly ovate, rounded, short-stalked, yellow or orange-color below the middle and dark chestnut-brown and lustrous above it; staminate flowers composed of a membranaceous 4-lobed calyx often 2-lobed by suppression, the anterior lobe obovate, rounded at apex, as long as the stamens, much longer than the minute posterior lobe, and of 2 stamens inserted on the base of the calyx, with short 2-branched filaments, each branch bearing an erect half-anther; pistillate aments oblong or cylindric, terminal on the short spur-like lateral branchlets, their scales closely imbricated, oblong-ovate, 3-lobed, light yellow, often tinged with red above the middle, accrescent, becoming brown and woody at maturity, and forming sessile or stalked erect or pendulous short or elongated strobiles usually ripening in the autumn, deciduous with the nuts from the slender rachis; calyx of the pistillate flower 0; ovary sessile, compressed, with styles stigmatic at apex. Nut minute, oval or obovoid, compressed, bearing at the apex the persistent stigmas, marked at the base by a small pale scar, the outer coat of the shell produced into a marginal wing interrupted at the apex.

Betula is widely distributed from the Arctic circle to Texas in the New World, and to southern Europe, the Himalayas, China, and Japan in the Old World, some species forming great forests at the north, or covering high mountain slopes. Of the twenty-eight or thirty species now recognized twelve are found in North America; of these nine are trees. Of exotic species the European and Asiatic Betula pendula Roth. in a number of forms is a common ornamental tree in the northern states, where several of the Birch-trees of eastern Asia also flourish. Many of the species produce wood valued by the cabinet-maker, or used in the manufacture of spools, shoe-lasts, and other small articles. The thin layers of the bark are impervious to water and are used to cover buildings, and for shoes, canoes, and boxes. The sweet sap provides an agreeable beverage.

CONSPECTUS OF THE NORTH AMERICAN ARBORESCENT SPECIES.

1. Betula lenta L. Cherry Birch. Black Birch.

Leaves ovate to oblong-ovate, acute or acuminate, gradually narrowed and often unequal at the cordate or rounded base, sharply serrate with slender incurved teeth, or very rarely laciniately lobed (f. laciniata Rehdr.), when they unfold light green, coated on the lower surface with long white silky hairs, and slightly hairy on the upper surface, at maturity thin and membranaceous, dark dull green above, light yellow-green below, with small tufts of white hairs in the axils of the veins, 2½′—6′ long, 1½′—3′ wide, with a yellow midrib and primary veins prominent and hairy on the lower surface, and obscure reticulate cross veinlets; turning bright clear yellow late in the autumn; petioles stout, hairy, deeply grooved on the upper side, ¾′—1′ long; stipules ovate, acute, light green or nearly white, scarious and ciliate above the middle. Flowers: staminate aments during the winter about ¾′ long, nearly ¼′ thick, with ovate acute apiculate scales bright red-brown above the middle and light brown below it, becoming 3′—4′ long; pistillate aments ½′—¾′ long, about ⅛′ thick, with ovate pale green scales rounded at the apex; styles light pink. Fruit: strobiles oblong-ovoid, sessile, erect, glabrous, 1′—1½′ long, about ½′ thick; nut obovoid, pointed at base, rounded at apex, about as broad as its wing.

A tree, with aromatic bark and leaves, 70°—80° high, with a trunk 2°—5° in diameter, slender branches spreading almost at right angles, becoming pendulous toward the ends and gradually forming a narrow round-topped open graceful head, and branchlets light green, slightly viscid and pilose when they first appear, soon turning dark orange-brown, lustrous during the summer, bright red-brown in their first winter, becoming darker and finally dark dull brown slightly tinged with red. Winter-buds ovoid, acute, about ¼′ long, with ovate acute light chestnut-brown loosely imbricated scales, those of the inner ranks becoming ½′—¾′ long. Bark on young stems and branches close, smooth, lustrous, dark brown tinged with red, and marked by elongated horizontal pale lenticels, becoming on old trunks ½′—¾′ thick, dull, deeply furrowed and broken into large thick irregular plates covered with closely appressed scales. Wood heavy, very strong and hard, close-grained, dark brown tinged with red, with thin light brown or yellow sapwood of 70—80 layers of annual growth; largely used for floors, in the manufacture of furniture and for fuel, and occasionally in ship and boat-building. Sweet birch-oil distilled from the wood and bark is used for medicinal purposes and for flavoring as a substitute for oil of wintergreen, and beer is obtained by fermenting the sugary sap.

Distribution. Rich uplands from southern Maine to northwestern Vermont, and eastern Ohio and southward to northern Delaware and along the Appalachian Mountains up to altitudes of 4000° to northern Georgia; in Alabama, and in eastern Kentucky and Tennessee; a common forest tree at the north, and of its largest size on the western slopes of the southern Alleghany Mountains.

× Betula Jackii Schn., a natural hybrid of B. lenta with B. pumila Michx., has appeared in the Arnold Arboretum.

2. Betula lutea Michx. Yellow Birch. Gray Birch.

Leaves ovate to oblong-ovate, acuminate or acute at apex, gradually narrowed to the rounded cuneate or rarely heart-shaped usually oblique base, sharply doubly serrate, when they unfold bronze-green or red, and pilose with long pale hairs above and on the under side of the midrib and veins, at maturity dull dark green above, yellow-green below, 3′—4½′ long, 1½′—2′ wide, with a stout midrib and primary veins covered below near the base of the leaf with short pale or rufous hairs; turning clear bright yellow in the autumn; petioles slender, pale yellow, hairy, ¾′—1′ long; stipules ovate, acute, light green tinged with pink above the middle, about ½′ long. Flowers: staminate aments during the winter ¾′—1′ long, about ⅛′ thick, with ovate rounded scales light chestnut-brown and lustrous above the middle, ciliate on the margins, becoming 3′—3½′ long and ⅓′ thick; pistillate aments [208] about ⅔′ long, with acute scales, pale green below, light red and tipped with clusters of long white hair at apex, and pilose on the back. Fruit: strobiles erect, sessile, short-stalked, pubescent, 1′—1½′ long, about ¾′ thick; nut ellipsoidal to obovoid, about ⅛′ long, rather broader than its wing.

A tree, with slightly aromatic bark and leaves, occasionally 100° high, with a trunk 3°—4° in diameter, spreading and more or less pendulous branches forming a broad round-topped head, and branchlets at first green and covered with long pale hairs, light orange-brown and pilose during their first summer, becoming glabrous and light brown slightly tinged with orange, and ultimately dull and darker. Winter-buds about ¼′ long, somewhat viscid and covered with loose pale hairs during the summer, becoming light chestnut-brown, acute, and slightly puberulous in winter. Bark of young stems and of the branches bright silvery gray or light orange color, very lustrous, separating into thin loose persistent scales more or less rolled on the margins, becoming on old trees ½′ thick, reddish brown, and divided by narrow irregular fissures into large thin plates covered with minute closely appressed scales, or sometimes dull yellowish brown (B. alleghaniensis Britt.). Wood heavy, very strong, hard, close-grained, light brown tinged with red, with thin nearly white sapwood; largely used for floors, in the manufacture of furniture, button and tassel moulds, boxes, the hubs of wheels, and for fuel.

Distribution. Moist uplands, and southward often in swamps; one of the largest deciduous-leaved trees of northeastern America; Newfoundland and along the northern shores of the Gulf of St. Lawrence to the valley of Rainy River, and southward to Long Island (Cold Spring Harbor) and western New York, Pennsylvania, northern Delaware, southeastern Ohio, northern Indiana, southwestern Wisconsin, northern, northeastern and central Iowa, and from the mountains of Virginia and West Virginia to the highest peaks of North Carolina and Tennessee at altitudes between 3000° and 5000°; very abundant and of its largest size in the eastern provinces of Canada and in northern New York and New England; small and rare in southern New England and southward.

× Betula Purpusii Schn. believed to be a natural hybrid of B. lutea with B. pumila var. glandulifera Regel has been found in Michigan and in Tamarack Swamps in Hennepin, Pine and Anoka Counties, Minnesota.

3. Betula nigra L. Red Birch. River Birch.

Leaves rhombic-ovate, acute, abruptly or gradually narrowed and cuneate at base, doubly serrate, and on vigorous young branches often more or less laciniately cut into acute [209] doubly serrate lobes, when they unfold light yellow-green and pilose above and coated below, especially on the midrib and petioles, with thick white tomentum, at maturity thin and tough, 1½′—3′ long, 1′—2′ wide, deep green and lustrous above, glabrescent, pubescent or ultimately glabrous below, except on the stout midrib and remote primary veins; turning dull yellow in the autumn; petioles slender, slightly flattened, tomentose, about ½′ long; stipules ovate, rounded or acute at apex, pale green, covered below with white hairs. Flowers: staminate aments clustered, during the winter about ⅞ long and ¹/₁₆′ thick, with ovate rounded dull chestnut-brown lustrous scales, becoming 2′—3′ long and ⅛′ thick; pistillate aments about ⅓′ long, with bright green ovate scales pubescent on the back, rounded or acute at apex, and ciliate with long white hairs. Fruit ripening in May and June; strobiles cylindric, pubescent, 1′—1½′ long, ½′ thick, erect on stout tomentose peduncles ½′ long; nut ovoid to ellipsoidal, ⅛′ in length, pubescent or puberulous at apex, about as broad as its thin puberulous wing, ciliate on the margin.

A tree, 80°—90° high, with a trunk often divided 15°—20° above the ground into 2 or 3 slightly diverging limbs, and sometimes 5° in diameter, slender branches forming in old age a narrow irregular picturesque crown, and branchlets coated at first with thick pale or slightly rufous tomentum gradually disappearing before winter, becoming dark red and lustrous, dull red-brown in their second year, and then gradually growing slightly darker until the bark separates into the thin flakes of the older branches; or often sending up from the ground a clump of several small spreading stems forming a low bushy tree. Winter-buds ovoid, acute, about ¼′ long, covered in summer with thick pale tomentum, glabrous or slightly puberulous, lustrous and bright chestnut-brown in winter, the inner scales strap-shaped, light brown tinged with red, and coated with pale hairs. Bark on young stems and large branches thin, lustrous, light reddish brown or silvery gray, marked by narrow slightly darker longitudinal lenticels, separating freely into large thin papery scales persistent for several years, and turning back and showing the light pink-brown tints of the freshly exposed inner layers, becoming at the base of old trunks from ¾′—1′ thick, dark red-brown, deeply furrowed and broken on the surface into thick closely appressed scales. Wood light, rather hard, strong, close-grained, light brown, with pale sapwood of 40—50 layers of annual growth; used in the manufacture of furniture, wooden ware, wooden shoes, and in turnery.

Distribution. Banks of streams, ponds, and swamps, in deep rich soil often inundated for several weeks at a time; near Manchester, Hillsboro County, New Hampshire, northeastern Massachusetts, Long Island, New York, southward to northern Florida through the region east of the Alleghany Mountains except in the immediate neighborhood of the [210] coast, through the Gulf states to the valley of the Navasota River, Brazos County, Texas, and through Arkansas, eastern Oklahoma, southeastern Kansas, and Missouri to Tennessee and Kentucky, southern and eastern Iowa, southern Minnesota, the valley of the Eau Claire River, Eau Claire County, Wisconsin, southern Illinois, the valley of the Kankakee River, Indiana, and southern Ohio; the only semiaquatic species and the only species ripening its seeds in the spring or early summer; attaining its largest size in the damp semitropical lowlands of Florida, Louisiana, and Texas; the only Birch-tree of such warm regions.

Often cultivated in the northeastern states as an ornamental tree, growing rapidly in cultivation.

4. Betula populifolia Marsh. Gray Birch. White Birch.

Leaves nearly triangular to rhombic, long-pointed, coarsely doubly serrate with stout spreading glandular teeth except at the broad truncate or slightly cordate or cuneate base, thin and firm, dark green and lustrous and somewhat roughened on the upper surface early in the season by small pale glands in the axils of the conspicuous reticulate veinlets, 2½′—3′ long, 1½′—2½′ wide, with a stout yellow midrib covered with minute glands, and raised and rounded on the upper side, and obscure yellow primary veins; turning pale yellow in the autumn; petioles slender, terete, covered with black glands, often stained with red on the upper side, ¾′—1′ long; stipules broadly ovate, acute, membranaceous, light green slightly tinged with red. Flowers: staminate aments usually solitary or rarely in pairs, 1¼′—1½′ long, about ⅛′ thick during the winter, becoming 2½′—4′ long, with ovate acute apiculate scales; pistillate aments slender, as long as their glandular peduncles about ½′ in length, with ovate acute pale green glandular scales. Fruit: strobiles cylindric, pubescent, obtuse at apex, about ¾′ long and ⅓′ thick, pendant or spreading on slender stems; nut ellipsoidal to obovoid, acute or rounded at base, a little narrower than its obovate wing.

A short-lived tree, 20°—30° or exceptionally 40° high, with a trunk rarely 18′ in diameter, short slender often pendulous more or less contorted branches usually clothing the stem to the ground and forming a narrow pyramidal head, and branchlets roughened by small raised lenticels, resinous-glandular when they first appear, gradually growing darker, bright yellow and lustrous before autumn like the young stems, bright reddish brown during their first winter, and ultimately white near the trunk; often growing in clusters of spreading stems springing from the stumps of old trees. Winter-buds ovoid, acute, pale chestnut-brown, glabrous, about ¼′ long. Bark about ⅓′ thick, dull chalky white on the outer surface, bright orange on the inner, close and firm, with dark triangular markings at the [211] insertion of the branches, becoming at the base of old trees thicker, nearly black, and irregularly broken by shallow fissures. Wood light, soft, not strong, close-grained, not durable, light brown, with thick nearly white sapwood; used in the manufacture of spools, shoe-pegs and wood pulp, for the hoops of barrels, and largely for fuel.

Distribution. Dry gravelly barren soil or on the margins of swamps and ponds; Prince Edward Island, Nova Scotia, New Brunswick, and the valley of the lower St. Lawrence River southward to northeastern, central and on South Mountain, Franklin County, Pennsylvania, and northern Delaware, and westward through northern New England and New York, ascending sometimes to altitudes of 1800°, to the southern shores of Lake Ontario, and at the foot of Lake Michigan, Indiana; rare and local in the interior, very abundant in the coast region of New England and the middle states; springing up in great numbers on abandoned farm-lands or on lands stripped by fire of their original forest covering; most valuable in its ability to grow rapidly in sterile soil and to afford protection to the seedlings of more valuable and less rapid-growing trees.

A form with deeply divided leaves (var. laciniata Loud.) and one with purple leaves (var. purpurea E & B) are occasionally cultivated.

A shrub believed to be a natural hybrid of B. populifolia with B. pumila Michx. has been found near Mt. Mansfield, Vermont.

5. Betula cœrulea Blanch. Blue Birch.

Leaves ovate, long-pointed, broadly or narrowly concave-cuneate at the entire often unequal base, sharply mostly doubly serrate above with straight or incurved glandular often apiculate teeth, covered above when they unfold with pale deciduous glands, at maturity dull bluish green above, pale yellow-green below, and sparingly villose along the under side of the slender yellow midrib and primary veins, 2′—2½′ long, 1′—1½′ wide; petioles slender, ¾′—1¼′ long, yellow more or less deeply tinged with red. Flowers: staminate aments usually in pairs, or singly or in 3’s, 1¼′—2′ long, about ³/₁₆′ thick, with ovate rounded short-pointed scales; pistillate aments slender, about ⅓′ long, with acuminate pale green much reflexed scales. Fruit: strobiles cylindric, pubescent, slightly narrowed at the obtuse apex, about 1′ long and ¼′ thick, pendant on slender peduncles ¼′—½′ in length; nut ellipsoidal, much narrower than its broad wing.

A tree, rarely more than 30° high, with a trunk 8′—10′ in diameter, small ascending finally spreading branches, and slender branchlets marked by numerous small raised pale lenticels, purplish and sparingly villose when they first appear, soon glabrous, becoming [212] bright red-brown; often forming clumps of several stems. Bark thin, white tinged with rose, lustrous, not readily separable into layers, the inner bark light orange color.

Distribution. Moist slopes, Stratton and Windham, Windham County, Vermont, at altitudes of about 1800° (W. H. Blanchard), Haystack Mountain, Aroostook County, Maine (M. L. Fernald); the American representative of the European Betula pendula Roth., and probably widely distributed over the hills of northern New England and eastern Canada. Perhaps with its variety best considered a natural hybrid between B. papyrifera and B. populifolia.

Apparently passing into a form with larger leaves often rounded and truncate at the broad base, 3′—3½′ long and 2′ wide, stouter staminate aments, and strobiles frequently 1½′ long and ½′ thick (var. Blanchardii Sarg. fig. 198 A). This under favorable conditions is a tree 60°—70° high, with a trunk 18′ in diameter; common with Betula cœrulea at Windham and Stratton, Vermont (W. H. Blanchard), and on a hill near the coast in Washington County, Maine (M. L. Fernald).

6. Betula papyrifera Marsh. Canoe Birch. Paper Birch.

Leaves ovate, acute or acuminate with a short broad point, coarsely usually doubly and often very irregularly serrate except at the rounded abruptly cuneate or gradually narrowed base, bright green, glandular-resinous, pubescent and clothed below on the midrib and primary veins and on the petioles with long white hairs when they unfold, at maturity thick and firm, dull dark green and glandless or rarely glandular on the upper surface, light yellow-green and glabrous or puberulous, with small tufts of pale hairs in the axils of the primary veins and covered with many black glands on the lower surface, 2′—3′ long, 1½′—2′ wide, with a slender yellow midrib marked, like the remote primary veins, with minute black glands, turning light clear yellow in the autumn; petioles stout, yellow, glandular, glabrous or pubescent, ½′—¾′ long; stipules ovate, acute, ciliate on the margins with pale hairs, light green. Flowers: staminate aments clustered during the winter, ¾′—1¼′ long, about ⅛′ thick, with ovate, acute scales light brown below the middle, dark red-brown above it, becoming 3½′—4′ long, and about ⅓′ thick; pistillate aments 1′—1¼′ long, about ¹/₁₆′ thick, with light green lanceolate scales long-pointed and acute or rounded at apex; styles bright red. Fruit: strobiles cylindric, glabrous, about 1½′ long and ⅓′ thick, hanging on slender stalks, their scales very rarely entire (var. elobata Sarg.); nut ellipsoidal, about ¹/₁₆′ long, much narrower than its thin wing.

A tree, usually 60°—70° tall, with a trunk 2°—3° in diameter, becoming in old age, or when crowded by other trees, branchless below and supporting a narrow open head of [213] short pendulous branches, and branchlets at first light green, slightly viscid, marked by scattered orange-colored oblong lenticels and covered with long pale hairs, dark orange color and glabrous or pubescent during the summer, becoming dull red in their first winter, gradually growing dark orange-brown, lustrous for four or five years and ultimately covered with the white papery bark of older branches. Winter-buds obovoid, acute, about ¼′ long, pubescent below the middle and coated with resinous gum at midsummer, dark chestnut-brown, glabrous and slightly resinous during the winter, their inner scales becoming strap-shaped, rounded at apex, about ½′ long and ⅛′ wide. Bark on young trunks and large limbs thin, creamy white or rarely bronze color or orange-brown and lustrous on the outer surface, bright orange color on the inner, marked by long narrow slightly darker colored raised lenticels, separating into thin papery layers, pale orange color when first exposed to the light, becoming on old trunks for a few feet above the ground sometimes ½′ thick, dull brown or nearly black, sharply and irregularly furrowed and broken on the surface into thick closely appressed scales. Wood light, strong, hard, tough, very close-grained, light brown tinged with red, with thick nearly white sapwood; largely used for spools, shoe-lasts, pegs, and in turnery, the manufacture of wood-pulp, and for fuel. The tough resinous durable bark impervious to water is used by all the northern Indians to cover their canoes and for baskets, bags, drinking-cups, and other small articles, and often to cover their wigwams in winter.

Distribution. Rich wooded slopes and the borders of streams, lakes, and swamps scattered through forests of other trees; Labrador to the southern shores of Hudson’s Bay, and southward to Long Island, New York, northern Pennsylvania, central Michigan, northern Wisconsin, northern-central Iowa, eastern Nebraska, North and South Dakota and Wyoming; common in the maritime provinces of Canada and North of the Great Lakes, and in northern New England and New York; small and comparatively rare in the coast region of southern New England and southward; on the highest mountains of New England and northward the var. minor S. Wats and Cov. is common as a small shrub.

× Betula Sandbergii Britt. and its f. maxima Rosend. generally believed to be natural hybrids of B. papyrifera and B. pumila var. glandulifera Regl. occur in Tamarack swamps in Hennepin County, Minnesota.

Betula papyrifera var. cordifolia Fern.

Leaves ovate, abruptly pointed and acuminate or acute at apex, cordate at base, coarsely doubly serrate, glabrous or pilose on the under side of the midrib and veins, often furnished [214] below with axillary tufts of pale hairs, 1½′—3′ long, 1′—2½′ wide; petioles glabrous or rarely villose, ½′—¾′ in length. Fruit: strobiles ¾′—2′ long and ¼′—½′ thick, on villose peduncles up to ¾′ in length; scales glabrous or pubescent.

A tree rarely more than 30° tall, with slender glabrous or pubescent branchlets, and at high altitudes on the New England mountains reduced to a low shrub. Bark separating in thin layers, white or dark reddish brown.

Distribution. Labrador and Newfoundland to northern New England, and westward to the shores of Green Bay, Wisconsin, and those of Lake Superior, Minnesota (Grand Marais, Cook County); on Mt. Mitchell, North Carolina, at an altitude of 5550° (W. W. Ashe).

Betula papyrifera var. subcordata Sarg.

Leaves ovate, acute or acuminate at apex, slightly cordate or rounded at base, rarely slightly lobed above the middle, finely often doubly serrate with teeth pointing forward or spreading, glabrous, 2′—2½′ long, 1′—1½′ wide; petioles sparingly villose or glabrous, ½′—¾′ in length. Fruit: strobiles drooping on slender peduncles 1′—1½′ long, about ⅓′ thick, their scales puberulous, ciliate on the margins, the middle lobe acute, rather longer than the broad truncate lateral lobes; nut obovoid, cuneate at base, ¹/₁₂′ long, narrower than its wings.

A tree 25°—40° or occasionally 60° high, with a trunk 12′—18′ in diameter, and slightly glandular glabrous red-brown branchlets. Bark separating freely into thin layers, white or occasionally dark reddish brown or orange color.

Distribution. Alberta (Crow Nest Pass, neighborhood of Jasper and Cypress Hills), through northern Montana and Idaho to western Washington, northeastern Oregon (Minum River Valley) and British Columbia.

Betula papyrifera var. montanensis Sarg.

Leaves broadly ovate, acute at apex, truncate or rounded at base to oblong-ovate or lanceolate and long-pointed and acuminate at apex, narrowed and rounded at base, coarsely doubly serrate, thick, dark green above, paler, sparingly pubescent and furnished with [215] conspicuous tufts of axillary hairs below, 3′—5′ long, 2′—2¼′ wide; petioles puberulous, ¾′—1′ in length. Flowers unknown. Fruit: strobiles cylindric, 1¾′—2′ long, ½′ thick, pendent on puberulous peduncles ½′—¾′ in length, their scales puberulous, finely ciliate on the margins, the slender base of those below the middle of the ament rather more than twice as long as the expanded upper portion of the scale.

A tree 40°—50° high, with a trunk 12′—18′ in diameter, and slender branchlets red-brown, lustrous, marked by small pale lenticels and puberulous during their first season. Winter-buds narrow-obovoid, acuminate, dark red-brown, resinous, ⅓′ long. Bark white, or dark gray or brown.

Distribution. Shore of Yellow Bay, Flathead Lake, Flathead County, Montana, and at Sandpoint, Bonner County, Idaho.

Betula papyrifera var. occidentalis Sarg.

Leaves ovate, acute, or abruptly acuminate at apex, rounded or occasionally cordate or rarely cuneate at the broad base, coarsely and generally doubly serrate with straight or incurved glandular teeth, thin and firm in texture, dull dark green above, pale yellow-green below, and puberulous on both sides of the stout yellow midrib and slender primary veins, 3′—4′ long, 1½′—2′ wide; petioles stout, glandular, at first tomentose, ultimately pubescent or puberulous, about ¾′ long; stipules oblong-obovate, rounded and acute or apisculate at apex, ciliate on the margin, puberulous, glandular-viscid. Flowers: staminate aments during the winter about ¾′ long and ⅛′ thick, with ovate scales rounded or abruptly narrowed and acute at apex; puberulous on the outer surface, ciliate on the margins, becoming 3′—4′ long and about ¼′ thick; pistillate aments about 1′ long and ¹/₁₆′ thick, with acuminate bright green scales. Fruit: strobiles cylindric, puberulous, spreading, 1¼′—1½′ long, ¼′—½′ thick, on stout peduncles ¾′ in length, their scales ciliate on the margins; nut oval, about ¹/₁₆′ in length, and nearly as wide as its wings.

A tree, 100°—120° high, with a trunk 3°—4° in diameter, comparatively small branches often pendulous on old trees, and pale orange-brown branchlets more or less glandular and coated with long pale hairs when they first appear, becoming bright orange-brown and nearly destitute of glands during their first winter, and in their second year orange-brown, glabrous, and very lustrous. Winter-buds acute, bright orange-brown, ⅛′—¼′ long, their light brown inner scales sometimes becoming ¾′ in length. Bark thin, marked by long oblong horizontal raised lenticels, dark orange-brown or white, very lustrous, separating [216] freely into thin papery layers displaying in falling the bright orange-yellow inner bark.

Distribution. Banks of streams and lakes; southwestern British Columbia and northwestern Washington and eastward through eastern Washington and northern Idaho to northern Montana west of the continental divide; nowhere common and probably of its largest size on the alluvial banks of the lower Fraser River, and on the islands of Puget Sound.

Betula papyrifera var. kenaica A. Henry. Red Birch. Black Birch.

Leaves ovate, acute or acuminate, broadly cuneate or somewhat rounded at the entire base, irregularly coarsely often doubly serrate, glabrous, dark dull green above, pale yellow-green below, 1½′—2′ long, 1′—1¾′ wide, with a slender yellow midrib and 5 pairs of thin primary veins; petioles slender, ¾′—1′ long. Flowers: staminate aments clustered, 1′ long, [217] with ovate acute scales apiculate at apex, puberulous on the outer surface; pistillate aments, ⅓′—½′ long, about ¹/₁₆′ thick, on slender glandular pubescent peduncles ½′—¾′ in length; scales acuminate light green strongly reflexed; styles bright red. Fruit: strobiles cylindric, glabrous, 1′ long, their scales ciliate on the margins; nut oval, somewhat narrower than its thin wing.

A tree, 30°—40° high, with a trunk 12′—20′ in diameter, wide-spreading branches, stout branchlets marked by numerous small pale lenticels, bright red-brown during 2 or 3 years, gradually becoming darker. Bark thin, more or less furrowed, very dark brown or nearly black near the base of the trunk, grayish white or light reddish brown and separating into thin layers higher on the stem and on the branches.

Distribution. Coast of Alaska from Cook Inlet southward to the head of the Lynn Canal.

7. Betula alaskana Sarg. White Birch.

Leaves rhombic to deltoid-ovate, long-pointed, truncate, rounded or broadly cuneate, or on leading shoots occasionally cordate at the entire base, coarsely and often doubly glandular-serrate, thin, dark green above, pale and yellow-green below, 1½′—3′ long, 1′—1½′ wide, with a slender midrib and primary veins pubescent or ultimately glabrous below; petioles often bright red, somewhat hairy at first, finally glabrous, about 1′ long; Flowers: staminate aments clustered, sessile, 1′ long, ⅛′ thick, with ovate acuminate scales puberulous on the outer surface, and bright red, with yellow margins; pistillate aments slender, cylindric, glandular, 1′ long, ⅛′ thick, on stout peduncles nearly ½′ in length. Fruit: strobiles glabrous, pendulous or spreading, 1′—1¼′ long, ⅓′—½′ thick, their scales ciliate on the margins; nut oval, narrower than its broad wing.

A tree, usually 30°—40°, occasionally 80°, high, with a trunk 6′—12′ in diameter, slender erect and spreading or pendulous branches, and glabrous bright red-brown branchlets more or less thickly covered during their first year with resinous glands sometimes persistent until the second or third season. Winter-buds ovoid, obtuse at the gradually narrowed apex, about ¼′ long, with light red-brown shining outer scales sometimes ciliate on the margins, and oblong rounded scarious inner scales hardly more than ½′ long when fully grown. Bark thin, marked by numerous elongated dark slightly raised lenticels, dull reddish brown or sometimes nearly white on the outer surface, light red on the inner surface, close and firm, finally separable into thin plate-like scales.

Distribution. Valley of the Saskatchewan northwestward to the valley of the Yukon, [218] growing sparingly near the banks of streams in forests of coniferous trees and in large numbers on sunny slopes and hillsides; the common Birch-tree of the Yukon basin.

× Betula commixta Sarg., a shrub, growing on the tundra near Dawson, Yukon Territory, is believed to be a hybrid between B. alaskana and B. glandulosa Michx.

8. Betula fontinalis Sarg. Black Birch.

Leaves ovate, acute or acuminate, sharply and often doubly serrate, except at the rounded or abruptly cuneate often unequal base, and sometimes slightly laciniately lobed, pale green, pilose above, and covered by conspicuous resinous glands when they unfold, at maturity thin and firm, dark dull green above, pale yellow-green, rather lustrous and covered by minute glandular dots below, 1′—2′ long, ¾′—1′ wide, with a slender pale midrib, remote glandular veins, and rather conspicuous reticulate veinlets; turning dull yellow in the autumn; petioles stout, puberulous, light yellow, glandular-dotted, flattened on the upper side, often flushed with red, ⅓′—½′ long; stipules broadly ovate, acute or rounded at apex, slightly ciliate, bright green, soon becoming pale and scarious. Flowers: staminate aments clustered, ½′—¾′ long and ¹/₁₆′ thick during the winter, with ovate acute light chestnut-brown scales pale and slightly ciliate on the margins, becoming 2′—2½′ long, and about ⅛′ thick, with apiculate scales; pistillate aments short-stalked, about ¾′ long, with ovate acute green scales; styles bright red. Fruit: strobiles cylindric, rather obtuse, puberulous or nearly glabrous, 1′—1¼′ long, ½′ thick, erect or pendulous on slender glandular peduncles, ¼′ to nearly ¾′ in length; their scales ciliate, puberulous, the lateral lobes ascending, shorter than the middle lobe; nut ovoid or obovoid, puberulous at apex, nearly as wide as its wing.

A tree 20°—25° high with a short trunk, rarely more than 12′ or 14′ in diameter, ascending spreading and somewhat pendulous branches forming a broad open head, and slender branchlets, when they first appear light green glabrous or puberulous and covered with lustrous resinous glands persistent during their second season, and dark red-brown in their first winter; more commonly shrubby, with many thin spreading stems forming open clusters, 15°—20° high; often much lower, and frequently crowded in almost impenetrable thickets. Winter-buds ovoid, acute, very resinous, chestnut-brown, ¼′ long. Bark about ¼′ thick, dark bronze color, very lustrous, marked by pale brown longitudinal lenticels becoming on old trunks often 6′—8′ long and ¼′ wide. Wood soft and strong, light brown, with thick lighter-colored sapwood; sometimes used for fuel and fencing.

Distribution. Moist soil near the banks of streams usually in mountain cañons; generally [219] distributed, although nowhere very common: valley of the Saskatchewan (Saskatoon), Saskatchewan, westward to the basin of the upper Fraser and Pease Rivers, British Columbia, southward along the Rocky Mountains to eastern Utah, northern New Mexico and Arizona, the valleys of the Shasta region and the eastern slope of the Sierra Nevada, northern California, and eastward in the United States to the eastern foothills of the Rocky Mountains of Colorado, the Black Hills of South Dakota, and northwestern Nebraska. Passing into

Betula fontinalis var. Piperi Sarg.

A tree occasionally 50°—60° high with a tall trunk 12′—18′ in diameter, short spreading branches, and usually longer and often narrower strobiles.

Distribution. Spokane, Spokane County, Almota and Pullman, Whitman County, eastern Washington.

9. Betula Eastwoodæ Sarg.

Leaves broad-ovate to elliptic, acute, rounded or abruptly short-pointed at apex, coarsely serrate except at the cuneate base, thick, glabrous, dark green above, pale below, reticulate-venulose, [220] the veinlets more conspicuous on the lower surface, 1′—1½′ long, ¾′—1½′ wide; petioles slender, glabrous ⅕′—⅓′ in length; stipules scarious, ovate-oblong, rounded at apex. Flowers: staminate aments usually solitary or in pairs, sessile, 1′—1¼′ long, ⅕′ thick, with broadly ovate pubescent dark red scales acute and apiculate at apex; pistillate aments ½′ long, about ¹/₁₂′ thick, with acute light green scales. Fruit: strobiles pendulous on peduncles ⅓′—½′ long, cylindric, ¾′ in length, about ⅙′ thick, their scales glabrous longer than broad, the lobes narrowed at the rounded apex, ciliate, the lateral slightly spreading, one third shorter than the terminal lobe.

A tree 18°—20° high, with a trunk rarely more than 6′ in diameter, and slender red glabrous branchlets thickly covered with circular white glands. Bark close, chestnut-brown, marked by conspicuous horizontal white lenticels, about ⅕′ thick.

Distribution. Swamps near Dawson, Yukon Territory, forming jungles with Betula glandulosa Michx., B. alaskana Sarg., and various Willows; as a large shrub in Jasper Park near Jasper, Alberta.

4. ALNUS L. Alder.

Trees and shrubs, with astringent scaly bark, soft straight-grained wood, naked stipitate winter-buds formed in summer and nearly inclosed by the united stipules of the first leaf, becoming thick, resinous, and dark red. Leaves open and convex in the bud, falling without change of color; stipules of all but the first leaf ovate, acute, and scarious. Flowers vernal, or rarely opening in the autumn from aments of the year, in 1—3-flowered cymes in the axils of the peltate short-stalked scales of stalked aments formed in summer or autumn in the axils of the last leaves of the year or of those of minute leafy bracts; staminate aments elongated, pendulous, paniculate, naked and erect during the winter, each staminate flower subtended by 3—5 minute bractlets adnate to the scales of the ament, and composed of a 4-parted calyx, and 1—3 or usually 4 stamens inserted on the base of the calyx opposite its lobes, with short simple filaments; pistillate aments ovoid or oblong, erect, stalked, produced in summer in the axils of the leaves of a branch developed from the axils of an upper leaf of the year, and below the staminate inflorescence, inclosed at first in the stipules of the first leaf, emerging in the autumn and naked during the winter, or remaining covered until early spring; pistillate flowers in pairs, each flower subtended by 2—4 minute bractlets adnate to the fleshy scale of the ament becoming at maturity thick and woody, obovate, 3—5-lobed or truncate at the thickened apex, forming an ovoid or subglobose strobile persistent after the opening of its closely imbricated scales; calyx 0; ovary compressed; nut minute, bright chestnut-brown, ovoid to oblong, flat, bearing at the apex the remnants of the style, marked at the base by a pale scar, the outer coat of the shell produced into lateral wings often reduced to a narrow membranaceous border.

Alnus inhabits swamps, river bottom-lands, and high mountains, and is widely and generally distributed through the northern hemisphere, often forming the most conspicuous feature of vegetation on mountain slopes, ranging at high altitudes southward in the New World through Central America to Colombia, Peru, and Bolivia, and to upper Assam and Japan in the Old World. Of the eighteen or twenty species now recognized nine are North American; of these, six attain the size and habit of trees. Of the exotic species, Alnus vulgaris Hill., a common European, north African, and Asiatic timber-tree, was introduced many years ago into the northeastern states, where it has become locally naturalized. The wood of Alnus is very durable in water, and the astringent bark and strobiles are used in tanning leather and in medicine.

CONSPECTUS OF THE NORTH AMERICAN ARBORESCENT SPECIES.

1. Alnus sinuata Rydb. Alder.

Leaves ovate, acute, full and rounded and often unsymmetrical and somewhat oblique or abruptly narrowed and cuneate at base, divided into numerous short acute lateral lobes, sharply and doubly serrate with straight glandular teeth, glandular-viscid as they unfold, at maturity membranaceous, yellow-green on the upper surface, pale and very lustrous on the lower surface, glabrous, or villose along the under side of the stout midrib with short brown hairs also forming tufts in the axils of the numerous slender primary veins, 3′—6′ long, 1½′—4′ wide; petioles stout, grooved, abruptly enlarged at the base, ½′—¾′ in length; stipules oblong to spatulate, rounded and apiculate at apex, puberulous, about ¼′ long. Flowers: staminate aments sessile, in pairs in the axils of the upper leaves sometimes reduced to small bracts, and single in the axil of the leaf next below, during the winter about ½′ long and ⅛′ thick, with dark red-brown shining puberulous apiculate scales, becoming when the flowers open from spring to midsummer 4′ or 5′ long, with a puberulous light red rachis and ovate acute apiculate 3-flowered scales; calyx-lobes rounded, shorter than the 4 stamens; pistillate aments in elongated panicles, inclosed during winter in buds formed the previous summer in the axils of the leaves of short lateral branchlets, long-pedunculate, ⅓′ long, ⅙′ thick. Fruit: strobiles on slender peduncles in elongated sometimes leafy panicles 4′—6′ in length, oblong, ½′—⅝′ long, about ⅓′ thick, their truncate scales thickened at the apex; nut oval, about as wide as its wings.

A tree, sometimes 40° high, with a trunk 7′—8′ in diameter, short small nearly horizontal branches forming a narrow crown, and slender slightly zigzag branchlets puberulous and very glandular when they first appear, bright orange-brown and lustrous and marked by numerous large pale lenticels during their first season, much roughened during their second year by the elevated crowded leaf-scars, becoming light gray. Winter-buds acuminate, dark purple, covered especially toward the apex with close fine pubescence, about ½′ long. Bark thin bluish gray, with bright red inner bark; often a shrub only a few feet tall spreading into broad thickets.

Distribution. Northwest coast from the borders of the Arctic Circle to the high mountains of northern California; common in the valley of the Yukon and eastward through British Columbia to Alberta, and through Washington and Oregon to the western slopes of the Rocky Mountains in Montana; at the north with dwarf Willows, forming great thickets; in southeastern Alaska often a tall tree on rich moist bottom-lands near the mouths of mountain streams, and at the upper limits of tree growth a low shrub; very abundant in the valley of the Yukon on the wet banks of streams and often arborescent in habit; in British Columbia and the United States generally smaller and a shrub, growing usually only at altitudes of more than 3000° above the sea, and often forming thickets on the banks of streams and lakes.

2. Alnus rubra Bong. Alder.

Leaves ovate to elliptic, acute, abruptly or gradually narrowed and cuneate at base, crenately lobed, dentate with minute gland-tipped teeth, and slightly revolute on the margins, covered when they unfold with pale tomentum, at maturity thick dark green and glabrous or pilose with scattered white hairs above, clothed below with short rusty pubescence, 3′—5′ long, 1¾′—3′ wide, or on vigorous branchlets sometimes 8′—10′ long, with a broad midrib and primary veins green on the upper side and orange-colored on the lower, the primary veins running obliquely to the points of the lobes and connected by conspicuous slightly reticulate cross veinlets; petioles orange-colored, nearly terete, slightly grooved, ¼′—¾′ in length; stipules ovate, acute, pale green flushed with red, tomentose, ⅛′—¼′ long. Flowers: staminate aments in red-stemmed clusters, during the winter 1¼′ long, ⅛′ thick, with dark red-brown lustrous closely appressed scales, becoming 4′—6′ long and ¼′ thick, with ovate acute orange-colored glabrous scales; calyx yellow, with ovate rounded [223] lobes rather shorter than the 4 stamens; pistillate aments in short racemes usually inclosed during the winter in buds formed during the early summer and opening in the early spring, ⅓′—½′ long, about ¹/₁₆′ thick, with dark red acute scales; styles bright red. Fruit: strobiles raised on stout orange-colored peduncles sometimes ½′ in length, ovoid or oblong, ½′—1′ long, ⅓′—½′ thick, with truncate scales much thickened toward the apex; nut orbicular to obovoid, surrounded by a membranaceous wing.

A tree, usually 40°—50°, occasionally 90° high, with a trunk sometimes 3° in diameter, slender somewhat pendulous branches forming a narrow pyramidal head, and slender branchlets marked by minute scattered pale lenticels, light green and coated at first with hoary tomentum sometimes persistent until their second year, becoming during the first winter bright red and lustrous and ultimately ashy gray. Winter-buds about ⅓′ long, dark red, covered with pale scurfy pubescence. Bark rarely more than ¾′ thick, close, roughened by minute wart-like excrescences, pale gray or nearly white, with a thin outer layer, and bright red-brown inner bark. Wood light, soft, brittle, not strong, close-grained, light brown tinged with red, with thick nearly white sapwood; in Washington and Oregon largely used in the manufacture of furniture and for smoking salmon; by the Indians of Alaska the trunks are hollowed into canoes.

Distribution. Shores of Yakutat Bay, southeastern Alaska, southward near the coast to the cañons of the Santa Inez Mountains, Santa Barbara County, California; common along the banks of streams, and of its largest size near the shores of Puget Sound; in California most abundant in Mendocino, Humboldt and Marin Counties, forming groves on bottom-lands near the coast; often ranging inland for 20 or 30 miles, and occasionally ascending to altitudes of 2000° above the sea.

3. Alnus tenuifolia Nutt. Alder.

Leaves ovate-oblong, acute or acuminate, broad and rounded or cordate or occasionally abruptly narrowed and cuneate at base, usually acutely laciniately lobed and doubly serrate, when they unfold light green often tinged with red, pilose on the upper surface and coated on the lower with pale tomentum, at maturity thin and firm, dark green and glabrous above, pale yellow-green and glabrous or puberulous below, 2′—4′ long, 1½′—2½′ wide, with a stout orange-colored midrib impressed on the upper side, and slender primary veins running to the points of the lobes; petioles stout, slightly grooved, orange-colored, ½′—1′ in length; stipules ovate, acute, thin, and scarious, ½′ long, about ⅛′ wide, covered with pale pubescence. Flowers: staminate aments 3 or 4 in number in slender-stemmed racemes, nearly sessile or raised on stout peduncles often ½′ long, during the winter light purple, ¾′—1′ long [224] and ¼′ thick, becoming 1½′—2′ in length; calyx-lobes rounded, shorter than the 4 stamens; pistillate aments naked during the winter, dark red-brown, nearly ¼′ long, with acute apiculate loosely imbricated scales, only slightly enlarged when the flowers open. Fruit: strobiles obovoid-oblong, ⅓′—½′ long, their scales much thickened, truncate and 3-lobed at apex; nut nearly circular to slightly obovoid, surrounded by a thin membranaceous border.

A tree, occasionally 30° tall, with a trunk 6′—8′ in diameter, small spreading slightly pendulous branches forming a narrow round-topped head, and slender branchlets marked at first by a few large orange-colored lenticels and coated with fine pale or rusty caducous pubescence, becoming light brown or ashy gray more or less deeply flushed with red in their first winter and ultimately paler; more often shrubby, with several spreading stems, and at the north and at high altitudes frequently only 4°—5° tall. Winter-buds ¼′—⅓′ long, bright red, and puberulous. Bark rarely more than ¼′ thick, bright red-brown and broken on the surface into small closely appressed scales.

Distribution. Banks of streams and mountain cañons from Francis Lake in latitude 61° north to the valley of the lower Fraser River, British Columbia, eastward along the Saskatchewan to Prince Albert, and southward through the Rocky Mountains to northern New Mexico; on the Sierra Nevada of southern California, and in Lower California; the common Alder of mountain streams in the northern interior region of the continent; very abundant on the eastern slopes of the Cascade Mountains, and on the southern California Sierras; forming great thickets at 6000°—7000° above the sea along the headwaters of the rivers of southern California flowing to the Pacific Ocean; the common Alder of eastern Washington and Oregon, and of Idaho and Montana; very abundant and of its largest size in Colorado and northern New Mexico.

4. Alnus rhombifolia Nutt. White Alder. Alder.

Leaves ovate or oval or sometimes nearly orbicular, rounded or acute at apex, especially on vigorous shoots, gradually or abruptly narrowed and cuneate at base, finely or sometimes coarsely and occasionally doubly serrate, slightly thickened and reflexed on the somewhat undulate margins, when they unfold pale green and covered with deciduous matted white hairs, at maturity dark green and lustrous on the upper surface, frequently marked, especially on the midrib, with minute glandular dots, light yellow-green and slightly puberulous below, 2′—3′ long, 1½′—2′ wide, with a stout yellow midrib and primary veins; petioles slender, yellow, hairy, flattened and grooved on the upper side, ½′—¾′ long; stipules ovate, acute, scarious, puberulous, about ¼′ in length. Flowers: staminate aments in slender-stemmed pubescent clusters, usually short-stalked, during the summer dark olive-brown [225] and lustrous, ¾′—1′ long and about ¹/₁₆′ thick, beginning to lengthen late in the autumn before the leaves fall, fully grown and 4′—6′ long and ¼′ thick in January, with dark orange-brown scales, and deciduous in February before the appearance of the new leaves; calyx yellow, 4-lobed, rather shorter than the 2 or occasionally 3 or rarely single stamen; pistillate aments in short pubescent racemes emerging from the bud in December, their scales broadly ovate and rounded. Fruit: strobiles oblong, ⅓′—½′ long, with thin scales slightly thickened and lobed at apex, fully grown at midsummer, remaining closed until the trees flower the following year; nut broadly ovoid, with a thin margin.

A tree, frequently 70°—80° high, with a tall straight trunk 2°—3° in diameter, long slender branches pendulous at the ends, forming a wide round-topped open head, and slender branchlets marked by small scattered lenticels, at first light green and coated with pale caducous pubescence, soon becoming dark orange-red and glabrous, and darker during the winter and following summer. Winter-buds nearly ½′ long, very slender, dark red, and covered with pale scurfy pubescence. Bark on old trunks 1′ thick, dark brown, irregularly divided into flat often connected ridges broken into oblong plates covered with small closely appressed scales. Wood light, soft, not strong, brittle, close-grained, light brown, with thick lighter colored often nearly white sapwood.

Distribution. Banks of streams from northern Idaho to the eastern slope of the Cascade Mountains of Washington and southeastern Oregon, and southward from the valley of the Willamette River, Oregon (near Salem, Marion County, J. C. Nelson) over the coast ranges and along the western slopes of the Sierra Nevada to the mountains of southern California (San Bernardino, San Jacinto, and Cuyamaca Ranges); the common Alder of the valleys of central California, occasionally ascending on the southern Sierra Nevada to altitudes of 8000°, and the only species at low altitudes in the southern part of the state.

5. Alnus oblongifolia Torr. Alder.

Leaves oblong-lanceolate, acute; or rarely obovate and rounded at apex, gradually narrowed and cuneate at base, sharply and usually doubly serrate, more or less thickly covered, especially early in the season, with black glands, dark yellow-green and glabrous or slightly puberulous above, pale and glabrous or puberulous below, especially along the slender yellow midrib and veins, with small tufts of rusty hairs in the axils of the primary veins, 2′—3′ long, about 1½′ wide; petioles slender, grooved, pubescent, ¾′ long; stipules ovate-lanceolate, [226] brown and scarious, about ¼′ in length. Flowers: staminate aments in short stout-stemmed racemes, during the winter light yellow, ½′—¾′ long and about ¹/₁₆′ thick, becoming when the flowers open at the end of February before the appearance of the leaves 2′—2½′ in length, with ovate pointed dark orange-brown scales; calyx 4-lobed; stamens 3 or occasionally 2, with pale red anthers soon becoming light yellow; pistillate aments naked during the winter, ⅛′ to nearly ¼′ long, with light brown ovate rounded scales; stigmas bright red. Fruit: strobiles ½′—1′ long, with thin scales slightly thickened and nearly truncate at apex; nut broadly ovoid, with a narrow membranaceous border.

A tree, in the United States rarely more than 20°—30° high, with a trunk sometimes 8′ in diameter, long slender spreading branches forming an open round-topped head, and slender branchlets slightly puberulous when they first appear, light orange-red and lustrous during their first winter, and marked by small conspicuous pale lenticels, becoming in their second year dark red-brown or gray tinged with red and much roughened by the elevated leaf-scars. Winter-buds acute, red, lustrous, glabrous, ½′ long. Bark thin, smooth, light brown tinged with red.

Distribution. Banks of streams in cañons of the mountains of southern New Mexico and Arizona at altitudes of 4000°—6000° above the sea; in Oak Creek Cañon near Flagstaff, northern Arizona (tree 100° × 3°, P. Lowell); and on the mountains of northern Mexico.

6. Alnus maritima Nutt. Alder.

Leaves oblong-ovate, or obovate, acute, acuminate or rounded at apex, gradually narrowed and cuneate at base, remotely serrate with minute incurved glandular teeth, and somewhat thickened on the slightly undulate margins, when they unfold, light green tinged with red, hairy on the midrib, veins, and petioles, and coated above with pale scurfy pubescence, at maturity dark green, very lustrous, and covered below by minute pale glandular dots, 3′—4′ long, 1½′—2′ wide, with a stout yellow midrib and primary veins prominent and glandular on the upper side and slightly puberulous below; petioles stout, yellow, glandular, flattened and grooved on the upper side, ½′—¾′ in length; stipules oblong, acute, about ⅛′ long, dark reddish brown, caducous. Flowers opening in the autumn: aments appearing in July on branches of the year and fully grown in August or early in September; staminate in short scurfy-pubescent glandular-pitted racemes on slender peduncles sometimes 1′ in length from the axils of upper leaves, covered at first with ovate acute dark green very lustrous scales slightly ciliate on the margins and furnished at apex with minute red points, at maturity 1½′—2½′ long, ¼′ to nearly ½′ thick, with dark orange-brown scales raised on slender stalks, and 4 bright orange-colored stamens; pistillate usually solitary [227] from the axils of the lower leaves on stout pubescent peduncles, bright red at apex and light green below before opening, with ovate acute scales slightly ciliate on the margins, about ⅛′ long when the styles protrude from between the scales, beginning to enlarge the following spring. Fruit attaining full size at midsummer and then raised on a stout peduncle, broadly ovoid, rounded and depressed at base, gradually narrowed to the rather obtuse apex, about ⅝′ long and ½′ broad, with thin lustrous scales slightly thickened and crenately lobed at apex, turning dark reddish brown or nearly black and opening late in the autumn and remaining on the branches until after the flowers open the following year; nut oblong-obovoid, gradually narrowed and apiculate at apex, with a thin membranaceous border.

A tree, occasionally 30° high, with a tall straight trunk 4′—5′ in diameter, small spreading branches forming a narrow round-topped head, slender slightly zigzag branchlets, light green and hairy at first, pale yellow-green, very lustrous, slightly puberulous, marked with occasional small orange-colored lenticels, and glandular with minute dark glandular dots during their first summer, becoming dull light orange or reddish brown in the winter, and ashy gray often slightly tinged with red the following season; more often shrubby, with numerous slender spreading stems 15°—20° tall. Winter-buds acute, dark red, coated with pale lustrous scurfy pubescence, about ¼′ long. Bark ⅛′ thick, smooth, light brown or brown tinged with gray. Wood light, soft, close-grained, light brown, with thick hardly distinguishable sapwood.

Distribution. Banks of streams and ponds in southern Delaware and Maryland, and in south central Oklahoma (Johnson and Bryan Counties).

Occasionally cultivated as an ornamental tree in the eastern states and hardy as far north as Massachusetts.

X. FAGACEÆ.

Trees, with watery juice, slender terete branchlets marked by numerous usually pale lenticels, alternate stalked penniveined leaves, and narrow mostly deciduous stipules. Flowers monœcious, the staminate in unisexual heads or aments, composed of a 4—8-lobed calyx, and 4 or 8 stamens, with free simple filaments and introrse 2-celled anthers, the cells parallel and contiguous, opening longitudinally; the pistillate solitary or clustered, in terminal unisexual or bisexual spikes or heads, subtended by an involucre of imbricated bracts becoming woody and partly or entirely inclosing the fruit, and composed of a 4—8-lobed calyx adnate to the 3—7-celled ovary with as many styles as its cells and 1 or 2 pendulous anatropous or semianatropous ovules in each cell. Fruit a nut 1-seeded by abortion, the outer coat cartilaginous, the inner membranaceous or bony. Seed filling the cavity of the nut, without albumen; seed-coat membranaceous; cotyledons fleshy, including the minute superior radicle; hilum, basal, minute.

The six genera of this widely distributed family occur in North America with the exception of Nothofagus, separated from Fagus to receive the Beech-trees of the southern hemisphere.

CONSPECTUS OF THE NORTH AMERICAN GENERA.

1. FAGUS L. Beech.

Trees, with smooth pale bark, hard close-grained wood, and elongated acute bright chestnut-brown buds, their inner scales accrescent and marking the base of the branchlets with persistent ring-like scars. Leaves convex and plicate along the veins in the bud, thick and firm, deciduous; petioles short, nearly terete, in falling leaving small elevated semioval leaf-scars, with marginal rows of minute fibro-vascular bundle-scars; stipules linear-lanceolate, infolding the leaf in the bud. Flowers vernal after the unfolding of the leaves; staminate short-pedicellate, in globose many-flowered heads on long drooping bibracteolate stems at base of shoots of the year or from the axils of their lowest leaves, and composed of a subcampanulate 4—8-lobed calyx, the lobes imbricated in æstivation, ovate and rounded, and 8—16 stamens inserted on the base of and longer than the calyx, with slender filaments and oblong green anthers; pistillate in 2—4-flowered stalked clusters in the axils of upper leaves of the year, surrounded by numerous awl-shaped hairy bracts, the outer bright red, longer than the flowers, deciduous, the inner shorter and united below into a 4-lobed involucre becoming at maturity woody, ovoid, thick-walled, and covered by stout recurved prickles, inclosing or partly inclosing the usually 3 nuts, and ultimately separating into 4 valves; calyx urn-shaped, villose, divided into 4 or 5 linear-lanceolate acute lobes, its 3-angled tube adnate to the 3-celled ovary surmounted by 3 slender recurved pilose styles green and stigmatic toward the apex and longer than the involucre; ovules 2 in each cell. Nut ovoid, unequally 3-angled, acute or winged at the angles, concave and longitudinally ridged on the sides, chestnut-brown and lustrous, tipped with the remnants of the styles, marked at the base by a small triangular scar, with a thin shell covered on the inner surface with rufous tomentum. Seed dark chestnut-brown, suspended with the abortive ovules from the tip of the hairy dissepiment of the ovary pushed by the growth of the seed into one of the angles of the nut; cotyledons sweet, oily, plano-convex.

Fagus as here limited is confined to the northern hemisphere, with a single American species and seven Old World species; of these one is widely distributed through Europe, another is found in the Caucasus, and the others are confined to eastern temperate Asia. Of exotic species, the European Fagus sylvatica L., an important timber-tree, is frequently planted for ornament in the eastern states in several of its forms, especially those with purple leaves, and with pendulous branches. The wood of Fagus is hard and close-grained. The sweet seeds are a favorite food of swine, and yield a valuable oil.

1. Fagus grandifolia Ehrh. Beech.

Leaves remote at the ends of the branches and clustered on short lateral branchlets, oblong-ovate, acuminate with a long slender point, coarsely serrate with spreading or incurved triangular teeth except at the gradually narrowed generally cuneate base, when they unfold pale green and clothed on the lower surface and margins with long pale lustrous silky hairs, at maturity dull dark bluish green above, light yellow-green, very lustrous, and glabrous or rarely pilose below (f. pubescens Fern. & Rehd.) with tufts of long pale hairs in the axils of the veins, 2½′—5′ long, 1′—3′ wide, with a slender yellow midrib covered above with short pale hairs, and slender primary veins running obliquely to the points of the teeth; turning bright clear yellow in the autumn; very rarely deeply laciniate; petioles hairy, ¼′—½′ length; stipules ovate-lanceolate on the lower leaves, strap-shaped to linear-lanceolate on the upper, brown or often red below the middle, membranaceous, [229] lustrous, 1′—1½′ long. Flowers opening when the leaves are about one third grown; staminate in globose heads 1′ in diameter, on slender hairy peduncles about 2′ long; pistillate in usually 2-flowered clusters, on short clavate hoary peduncles ½′—¾′ long. Fruit: involucres ½′—¾′ in length often shorter than the nuts, on stout hairy club-shaped peduncles ¼′—¾′ long, fully grown at midsummer, and then puberulous, dark orange-green, and covered by long slender recurved prickles red above the middle, becoming at maturity in the autumn light brown and tomentose, with crowded much recurved pubescent prickles, persistent on the branch after opening late into the winter; nut about ¾′ long.

A tree, usually 70°—80° but exceptionally 120° high, sending up from the roots numerous small stems sometimes extending into broad thickets round the parent tree, in the forest with a long comparatively slender stem free of branches for more than half its length, and short branches forming a narrow head, in open situations short-stemmed, with a trunk often 3°—4° in diameter, and numerous limbs spreading gradually and forming a broad compact round-topped head of slender slightly drooping branches clothed with short leafy laterals, and branchlets pale green and coated with long soft caducous hairs when they first appear, olive-green or orange-colored during their first summer, and conspicuously marked by oblong bright orange lenticels, gradually growing red, bright reddish brown during their first winter, darker brown in their second season and ultimately ashy gray. Winter-buds puberulous, especially toward the apex, ¾′ to nearly 1′ long, about ⅛′ broad, the inner scales hirsute on the inner surface and along the margins and when fully grown often 1′ long, lustrous, brown above the middle, and reddish below. Bark ¼′—½′ thick, with a smooth light steel-gray surface. Wood hard, strong, tough, very close-grained, not durable, difficult to season, dark or often light red, with thin nearly white sapwood of 20—30 layers of annual growth; largely used in the manufacture of chairs, shoe-lasts, plane-stocks, the handles of tools, and for fuel. The sweet nuts are gathered and sold in the markets of Canada and of some of the western and middle states.

Distribution. Rich uplands and mountain slopes, often forming nearly pure forests, and southward on the bottom-lands of streams and the margins of swamps; valley of the Restigouche River, New Brunswick, to the northern shores of Lake Huron and the southern shores of Lake Superior, and southward to Virginia, Ohio, Michigan, the ravines of Rock River near Oregon, Ogle County, Illinois, Minnesota and northern Missouri; southward passing into the var. caroliniana Fern. & Rehd., differing in its ovate to short-ovate thicker leaves, usually rounded or subcordate at base, and often less coarsely serrate or [230] undulate on the margins, glabrous or rarely densely soft pubescent below (f. mollis Fern. & Rehd.), in the often shorter involucre of the fruit with shorter and less crowded prickles; usually on the bottom-lands of streams and the borders of swamps, New Jersey, and southern Ohio and Missouri to western Florida, Alabama, Mississippi, Louisiana, eastern Texas, and northeastern Oklahoma; ascending on the southern Appalachian Mountains to altitudes of 3000°; probably growing to its largest size in eastern Louisiana.

The northern form is occasionally planted in the northern states as a shade and park tree.

2. CASTANEA Adans. Chestnut.

Trees or shrubs, with furrowed bark, porous brittle wood, durable in the ground, terete branchlets without terminal buds, axillary buds covered by 2 pairs of slightly imbricated scales, the outer lateral, the others accrescent, becoming oblong-ovate and acute and marking the base of the branch with narrow ring-like scars, and stout perpendicular tap-roots; producing when cut numerous stout shoots from the stump. Leaves convolute in the bud, ovate, acute, coarsely serrate, except at the base, with thin veins running to the points of the slender glandular teeth, deciduous; petioles leaving in falling small elevated semioval leaf-scars marked by an irregular marginal row of minute fibro-vascular bundle-scars; stipules ovate to linear-lanceolate, acute, scarious, infolding the leaf in the bud, caducous. Flowers opening in early summer, unisexual, strong-smelling; the staminate, in 3—7-flowered cymes, in the axils of minute ovate bracts, in elongated simple deciduous aments first appearing with the unfolding of the leaves from the inner scales of the terminal bud and from the axils of the lower leaves of the year, composed of a pale straw-colored slightly puberulous calyx deeply divided into 6 ovate rounded segments imbricated in the bud, and 10—20 stamens inserted on the slightly thickened torus, with filiform filaments incurved in the bud, becoming elongated and exserted, and ovoid or globose pale yellow anthers; the pistillate scattered or spicate at the base of the shorter persistent androgynous aments from the axils of later leaves, sessile, 2 or 3 together or solitary within a short-stemmed or sessile involucre of closely imbricated oblong acute bright green bracts scurfy-pubescent or tomentose below the middle, subtended by a bract and 2 lateral bractlets, each flower composed of an urn-shaped calyx, with a short limb divided into 6 obtuse lobes, minute sterile stamens shorter than the calyx-lobes, an ovary 6-celled after fecundation, with 6 linear spreading white styles hairy below the middle and tipped by minute acute stigmas, and 2 ovules in each cell, attached on its inner angle, descending, semianatropous. Fruit maturing in one season, its involucre inclosing 1—3 nuts, globose or short-oblong, pubescent or tomentose and spiny on the outer surface, with elongated ridged bright green ultimately brown branched spines fascicled between the deciduous scales, coated on the inner surface with lustrous pubescence, splitting at maturity into 2—4 valves; nut ovoid, acute, crowned by the remnants of the style, bright chestnut-brown and lustrous, tomentose or pubescent at apex, cylindrical, or when more than 1 flattened, marked at the broad base by a large conspicuous pale circular or oval thickened scar, its shell lined with rufous or hoary tomentum. Seed usually solitary by abortion, dark chestnut-brown, marked at apex by the abortive ovules, with thick and fleshy more or less undulate ruminate sweet farinaceous cotyledons.

Castanea is confined to the northern hemisphere, and is widely distributed through eastern North America, southern Europe, northern Africa, southwestern Asia, and central and northern China, Korea, and Japan. Seven species are distinguished. In the countries of the Mediterranean Basin much attention has been given to improving the fruit of the native species Castanea sativa Mill., which is occasionally planted in the middle United States; in Japan the seeds of Castanea crenata S. & Zucc. in many varieties and in China those of Castanea mollissima Bl. are important articles of food. Castanea produces coarse-grained wood very durable in contact with the soil, and rich in tannin. Chestnut-trees suffer in the eastern United States from the attacks of a fungus, Endothia parasitica Anders. which has nearly exterminated them in many parts of the country.

CONSPECTUS OF THE NORTH AMERICAN SPECIES.

1. Castanea dentata Borkh. Chestnut.

Leaves oblong-lanceolate, acute and long-pointed at apex, gradually narrowed and cuneate at base, when they unfold puberulous on the upper surface and clothed on the lower with fine cobweb-like tomentum, at maturity thin, glabrous, dark dull yellow-green above, pale yellow-green below, 6′—8′ long, about 2′ wide, with a pale yellow midrib and primary veins; turning bright clear yellow late in the autumn; petioles stout, slightly angled, puberulous, ½′ long, often flushed with red; stipules ovate-lanceolate, acute, yellow-green, puberulous, about ½′ long. Flowers: staminate aments about ½′ long when they first appear, green below the middle and red above, becoming when fully grown 6′—8′ long, with stout green puberulous stems covered from base to apex with crowded flower-clusters; androgynous aments, slender, puberulous, 2½′—5′ long, with 2 or 3 irregularly scattered involucres of pistillate flowers near their base. Fruit: involucre attaining its full size by the middle of August, 2′—2½′ in diameter, sometimes a little longer than broad, somewhat flattened at apex, pubescent and covered on the outer surface with crowded fascicles of long slender glabrous much-branched spines, opening with the first frost and gradually shedding their nuts; nuts usually much compressed, ½′—1′ wide, usually rather broader than long, coated at apex or nearly to the middle with thick pale tomentum, the interior of the shell lined with thick rufous tomentum; seed very sweet.

A tree, occasionally 100° high, with a tall straight columnar trunk 3°—4° in diameter, or often when uncrowded by other trees with a short trunk occasionally 10°—12° in diameter, and usually divided not far above the ground into 3 or 4 stout horizontal limbs forming a broad low round-topped head of slightly pendulous branches frequently 100° across, and branchlets at first light yellow-green sometimes tinged with red, somewhat angled, lustrous, slightly puberulous, soon becoming glabrous and olive-green tinged with yellow or brown [232] tinged with green and ultimately dark brown. Winter-buds ovoid, acute, about ¼′ long, with thin dark chestnut-brown scales scarious on the margins. Bark from 1′—2′ thick, dark brown and divided by shallow irregular often interrupted fissures into broad flat ridges separating on the surface into small thin closely appressed scales. Wood light, soft, not strong, liable to check and warp in drying, easily split, reddish brown, with thin lighter colored sapwood of 3 or 4 layers of annual growth; largely used in the manufacture of cheap furniture and in the interior finish of houses, for railway-ties, fence-posts, and rails. The nuts, which are superior to those of the Old World chestnuts in sweetness were formerly gathered in great quantities in the forest and sold in the markets of the eastern cities.

Distribution. Southern Maine to Woodstock, Grafton County, New Hampshire (rare) and to the valley of the Winooski River, Vermont, southern Ontario, and southern Michigan, southward to Delaware and Ohio, southern Indiana, and southwestern Illinois (Pulaski County) along the Appalachian Mountains up to altitudes of 4000° to northern Georgia, and to western Florida (Crestview, Walton County) southeastern (Henry and Dale Counties) and south central (Dallas County) Alabama, Northern, central and southeastern Mississippi (Pearl River County), and to central Kentucky and Tennessee; very common on the glacial drift of the northern states and, except at the north, mostly confined to the Appalachian hills; attaining its greatest size in western North Carolina and eastern Tennessee.

Formerly sometimes planted in the eastern states as an ornamental and timber tree, and for its nuts, of which several varieties have been recognized.

× Castanea neglecta Dode with leaves intermediate between those of C. dentata and C. pumila and an involucre containing a single large nut occurs on the Blue Ridge near Highlands, Macon County, North Carolina.

2. Castanea pumila Mill. Chinquapin.

Leaves oblong-elliptic to oblong-obovate, acute, coarsely serrate, with slender rigid spreading or incurved teeth, gradually narrowed and usually unequal and rounded or cuneate at base, when they unfold tinged with red and coated above with pale caducous tomentum and below with thick snowy white tomentum, at maturity rather thick and firm in texture, bright yellow-green on the upper surface, hoary or silvery pubescent on the lower, 3′—5′ long, 1½′—2′ wide; turning dull yellow in the autumn; petioles stout, pubescent, flattened on the upper side, ¼′—½′ long; stipules light yellow-green, pubescent, those of the 2 lowest leaves broad, ovate, acute, covered at apex by rufous tomentum, on later leaves ovate-lanceolate, often oblique and acute, becoming linear at the end of the branch. Flowers: [233] staminate aments ½′ long when they first appear, pubescent, green below, bright red at apex, becoming when fully grown 4′—6′ long, with stout hoary tomentose stems and crowded or scattered flower-clusters; androgynous aments silvery tomentose, 3′—4′ long; involucres 1-flowered, scattered at the base of the ament or often spicate and covering its lower half, sessile or short-stalked. Fruit: involucre 1′—1½′ in diameter, with thin walls covered with crowded fascicles of slender spines tomentose toward the base; nut ovoid, terete, rounded at the slightly narrowed base, gradually narrowed and pointed at apex, more or less coated with silvery white pubescence, dark chestnut-brown, very lustrous, ¾′—1′ long, ⅓′ thick, with a thin shell lined with a coat of lustrous hoary tomentum, and a sweet seed.

A round-topped tree, rarely 50° high, with a short straight trunk 2°—3° in diameter, slender spreading branches, and branchlets coated at first with pale tomentum, becoming during their first winter pubescent or remaining tomentose at the apex, bright red-brown, glabrous, lustrous, olive-green or orange-brown during their second season and ultimately darker; east of the Mississippi River often a shrub spreading into broad thickets by prolific stolons, with numerous intricately branched stems often only 4° or 5° tall. Winter-buds ovoid, or oval, about ⅛′ long, clothed when they first appear in summer with thick hoary tomentum, becoming red during the winter and scurfy-pubescent. Bark ½′—1′ thick, light brown tinged with red, slightly furrowed and broken on the surface into loose plate-like scales. Wood light, hard, strong, coarse-grained, dark brown, with thin hardly distinguishable sapwood of 3 or 4 layers of annual growth; used for fence-posts, rails, and railway-ties. The sweet nuts are sold in the markets of the western and southern states.

Distribution. Dry sandy ridges, rich hillsides and the borders of swamps; southern New Jersey and Pennsylvania to central (Lake County) and western Florida and westward through the Gulf States to the valley of the Neches River, Texas, and through Arkansas to eastern Oklahoma and southwestern Missouri; on the Appalachian Mountains ascending to altitudes of 4500°; most abundant and of its largest size in southern Arkansas and eastern Texas.

3. Castanea alnifolia Nutt. Chinquapin.

A low shrub spreading into broad thickets by underground stems, with leaves pale pubescent on the lower surface; and distributed in the neighborhood of the coast from the valley of the Cape Fear River, North Carolina, to southern Georgia. Passing into

Castanea alnifolia var. floridana Sarg. Chinquapin.

Leaves oblong-obovate to elliptic, acute, acuminate or rounded at apex, gradually narrowed and cuneate or rounded at base, irregularly sinuate-toothed with apiculate teeth, [234] hoary tomentose below when they unfold, soon glabrous with the exception of the last leaves of vigorous summer shoots, and at maturity thin, glabrous, dark green above, light green and lustrous below, 3′—4′ long and 1′—1¾′ wide; petioles stout, glabrous, about ¹/₁₂′ in length. Flowers: staminate aments pale pubescent, 4′—5′ long; androgynous aments pubescent, as long or rather longer with ten or twelve involucres of pistillate flowers below the middle, often only the lowest being fertilized. Fruit: involucre 1-seeded, subglobose to short-oblong, pale tomentose, ¾′ to 1¼′ in diameter, covered with stout pubescent scattered spines divided at base into numerous branches; nut ovoid, terete, acute, dark chestnut-brown, lustrous, ½′ to nearly ¾′ in length.

A tree occasionally 40°—45° high, with a tall trunk sometimes a foot in diameter, small irregularly spreading branches forming a narrow head, and slender glabrous or rarely pilose red-brown branchlets; more often a shrub sometimes with broader obovoid leaves sometimes puberulous on the lower surface.

Dry sandy soil; coast of North Carolina, near Wrightsville, New Hanover County; Dover, near the Ogechee River, Screven County, Georgia; Jacksonville, Duval County, and Panama City on Saint Andrew’s Bay, Bay County, Florida; near Selma, Dallas County, Alabama; and Covington, St. Tammany Parish, Louisiana.

3. CASTANOPSIS Spach.

Trees, with scaly bark, astringent wood, and winter-buds covered by numerous imbricated scales. Leaves convolute in the bud, 5-ranked, coriaceous, entire or dentate, penniveined, persistent; stipules obovate or lanceolate, scarious, mostly caducous. Flowers in 3-flowered cymes, or the pistillate rarely solitary or in pairs, in the axils of minute bracts, on slender erect aments from the axils of leaves of the year; the staminate on usually elongated and panicled aments, and composed of a campanulate 5 or 6-lobed or parted calyx, the lobes imbricated in the bud, usually 10 or 12 stamens inserted on the slightly thickened torus, with elongated exserted filiform filaments and oblong anthers, and a minute hirsute rudimentary ovary; the pistillate on shorter simple or panicled aments or scattered at the base of the staminate inflorescence, the cymes surrounded by an involucre of imbricated scales; calyx urn-shaped, the short limb divided into 6 obtuse lobes; abortive stamens inserted on the limb of the calyx and opposite its lobes; ovary sessile on the thin disk, 3-celled after fecundation, with 3 spreading styles terminating in minute stigmas, and 2 ovules in each cell attached to its interior angle. Fruit maturing at the end of the second or rarely of the first season, its involucre inclosing 1—3 nuts, ovoid or globose, sometimes more or less depressed, rarely obscurely angled, dehiscent or indehiscent, covered by stout spines, tuberculate or marked by interrupted vertical ridges; nut more or less angled by mutual pressure when more than 1, often pilose, crowned with the remnants of the style, marked at the base by a large conspicuous circular depressed scar, the thick shell tomentose on the inner surface. Seed usually solitary by abortion, bearing at apex the abortive ovules; cotyledons plano-convex, fleshy, farinaceous.

Castanopsis inhabits California with two species, and southeastern Asia where it is distributed with about twenty-five species from southern China to the Malay Archipelago and the eastern Himalayas. Of the California species one is usually arborescent and the other Castanopsis sempervirens Dudley is a low alpine shrub of the coast ranges and the Sierra Nevada.

Castanopsis from κὰστανα and ὄψις, in allusion to its resemblance to the Chestnut-tree.

1. Castanopsis chrysophylla A. DC. Chinquapin. Golden-leaved Chestnut.

Leaves lanceolate or oblong-ovate, gradually narrowed at the ends or sometimes abruptly contracted at apex into a short broad point, entire with slightly thickened revolute margins, when they unfold thin, coated below with golden yellow persistent scales and above with scattered white scales, at maturity thick and coriaceous, dark green and [235] lustrous above, 2′—6′ long, ½′ to nearly 2′ wide, with a stout midrib raised and rounded on the upper side; turning yellow at maturity and falling gradually at the end of their second or in their third year; petioles ¼′—⅓′ in length; stipules ovate, rounded or acute at apex, brown and scarious, puberulous, ¼′—⅓′ long. Flowers appearing irregularly from June until February in the axils of broadly ovate apiculate pubescent bracts on staminate and androgynous scurfy stout-stemmed aments 2′—2½′ long and crowded at the ends of the branches; calyx of the staminate flower coated on the outer surface with hoary tomentum, divided into broadly ovate rounded lobes much shorter than the slender stamens; calyx of the pistillate flower oblong-campanulate, free from the ovary, clothed with hoary tomentum, divided at apex into short rounded lobes, rather shorter than the minute abortive stamens; anthers red; ovary conic, hirsute, with elongated slightly spreading thick pale stigmas. Fruit ripening at the end of the second season, involucre globose, dehiscent, irregularly 4-valved, often slightly shorter than the nuts, sessile, solitary, or clustered, tomentose and covered on the outer surface by long stout or slender rigid spines, 1′—1½′ in diameter, containing 1 or occasionally 2 nuts; nuts broadly ovoid, acute, obtusely 3-angled, light yellow-brown and lustrous; seeds dark purple-red, sweet and edible.

A tree, 50°—100° high, with a massive trunk 3°—6° in diameter, frequently free of branches for 50°, stout spreading branches forming a broad compact round-topped or conic head, and rigid branchlets coated when they first appear with bright golden-yellow scurfy scales, dark reddish brown and slightly scurfy during their first winter, and gradually growing darker in their second season; often much smaller and sometimes reduced to a shrub, 2°—12° high (var. minor A. De Candolle). Winter-buds fully grown at mid-summer, usually crowded near the end of the branch, ovoid or subglobose, with broadly ovate apiculate thin and papery light brown scales slightly puberulous on the back, ciliate on the scarious often reflexed margins, the terminal about ¼′ long and broad and rather larger than the often stipitate axillary buds. Bark 1′—2′ thick and deeply divided into rounded ridges 2′—3′ wide, broken into thick plate-like scales, dark red-brown on the surface and bright red internally. Wood light, soft, close-grained, not strong, light brown tinged with red, with thin lighter colored sapwood of 50—60 layers of annual growth; occasionally used in the manufacture of ploughs and other agricultural implements.

Distribution. Skamania County, Washington, valley of the lower Columbia River, Oregon, southward along the western slopes of the Cascade Mountains, and in California along the western slopes of the Sierra Nevada and through the coast ranges to the elevated valleys of the San Jacinto Mountains, sometimes ascending to altitudes of 4000° above the sea; of its largest size in the humid coast valleys of northern California.

4. LITHOCARPUS Bl.

Trees, with astringent properties, pubescence of fascicled hairs, deeply furrowed scaly bark, hard close-grained brittle wood, stout branchlets, and winter-buds covered by few erect or spreading foliaceous scales. Leaves convolute in the bud, petiolate, persistent, entire or dentate, with a stout midrib, primary veins running obliquely to the points of the teeth, or on entire leaves forked and united near the margins, and reticulate veinlets; stipules oblong-obovate to linear-lanceolate, those of the upper leaves persistent and surrounding the buds during the winter. Flowers in erect unisexual and in bisexual tomentose aments from the axils of leaves of the year, from the inner scales of the terminal bud or from separate buds in the axils of leaves of the previous year; staminate in 3-flowered clusters in the axils of ovate rounded bracts, the lateral flowers subtended by similar but smaller bracts, each flower composed of a 5-lobed tomentose calyx, with nearly triangular acute lobes, 10 stamens, with slender elongated filaments and small oblong or emarginate anthers, and an acute abortive hairy ovary; pistillate scattered at the base of the upper aments below the staminate flowers, solitary in the axils of acute bracts, furnished with minute lateral bractlets, and composed of a 6-lobed ovoid calyx, with rounded lobes, inclosed in the tomentose involucral scales, 6 stamens, with abortive anthers, an ovoid-oblong 3-celled ovary, 3 elongated spreading light green styles thickened and stigmatic at apex, and 2 anatropous ovules in each cell. Fruit an oval or ovoid nut maturing at the end of the second season, 1-seeded by abortion, surrounded at base by the accrescent woody cupular involucre of the flower, marked by a large pale circular basal scar, the thick shell tomentose on the inner surface. Seed red-brown, filling the cavity of the nut, bearing at apex the abortive ovules; cotyledons thick and fleshy, yellow and bitter.

Lithocarpus is intermediate between the Oaks and the Chestnuts, and, with the exception of one California species, is confined to southeastern Asia, where it is distributed with many species from southern Japan and southern China through the Malay Peninsula to the Indian Archipelago.

Lithocarpus from λίθος and καρπός, in allusion to the character of the fruit.

1. Lithocarpus densiflora Rehd. Tan Bark Oak. Chestnut Oak.

Leaves oblong or oblong-obovate, rounded or acute or rarely cordate at base, acute or occasionally rounded at apex, or rarely lanceolate and acuminate (f. lanceolata Rehdr.) repand-dentate, with acute callous teeth, or entire with thickened revolute margins, coated when they unfold with fulvous tomentum and glandular on the margins with dark caducous glands, at maturity pale green, lustrous and glabrous or covered with scattered pubescence on the upper surface, rusty-tomentose on the lower, ultimately becoming glabrous above and glabrate and bluish white below, 3′—5′ long, ¾′—3′ wide, with a midrib raised and rounded on the upper side, thin or thick primary veins and fine conspicuous reticulate veinlets; persistent until the end of their third or fourth year; petioles stout, rigid, tomentose, ½′—¾′ in length; stipules brown and scarious, hirsute on the outer surface. Flowers in early spring and frequently also irregularly during the autumn; aments stout-stemmed, 3′—4′ long; staminate flowers crowded, hoary-tomentose in the bud, their bracts tomentose. Fruit solitary or often in pairs, on a stout tomentose peduncle ½′—1′ in length; nut full and rounded at base, gradually narrowed and acute or rounded at apex, scurfy-pubescent when fully grown, becoming light yellow-brown, glabrous and lustrous at maturity, ¾′—1′ long, ½′—1′ thick, its cup shallow, tomentose with lustrous red-brown hairs on the inner surface, and covered by long linear rigid spreading or recurved light brown scales coated with fascicled hairs, frequently tipped, especially while young, with dark red glands and often tomentose near the base of the cup.

A tree, usually 70°—80° but sometimes 150° high, with a trunk 1°—4° in diameter, stout branches ascending in the forest and forming a narrow spire-like head, or in open positions spreading horizontally and forming a broad dense symmetrical round-topped crown, and branchlets coated at first with a thick fulvous tomentum of fascicled hairs often persistent until the second or third year, becoming dark reddish brown and frequently covered with a glaucous bloom; or sometimes reduced to a shrub, with slender stems only a few feet high (var. montana Rehdr.). Winter-buds ovoid, obtuse, ¼′—⅓′ long, often surrounded by the persistent stipules of the upper leaves, with tomentose loosely imbricated scales, those of the outer ranks linear-lanceolate, increasing in width toward the interior of the bud, those of the inner ranks ovate or obovate and rounded at apex. Bark ¾′—1½′ thick, deeply divided by narrow fissures into broad rounded ridges broken into nearly square plates covered by closely appressed light red-brown scales. Wood hard, strong, close-grained, brittle, reddish brown, with thick darker brown sapwood; largely used as fuel. The bark is exceedingly rich in tannin and is largely used for tanning leather.

Distribution. Valley of the Umpqua River, Oregon, southward through the coast ranges to the Santa Inez Mountains, California, and along the western slope of the Sierra Nevada up to elevations of 4000° above the sea to Mariposa County; very abundant in the humid coast region north of San Francisco Bay and on the Santa Cruz and Santa Lucia Mountains, and of its largest size in the Redwood forest of Napa and Mendocino Counties; southward and on the Sierras less abundant and of smaller size; the form lanceolata in southern Oregon and in Del Norte and Mendocino Counties, California; the var. montana at high altitudes on the Siskiyou Mountains, in the region of Mount Shasta and on the northern Sierra Nevada.

5. QUERCUS L. Oak.

Trees or shrubs, with astringent properties, pubescence of fascicled hairs, scaly or dark and furrowed bark, hard and close-grained or porous brittle wood, slender branchlets marked by pale lenticels and more or less prominently 5-angled. Winter-buds clustered at the ends of the branchlets, with numerous membranaceous chestnut-brown slightly accrescent caducous scales closely imbricated in 5 ranks, in falling marking the base of the branchlet with ring-like scars. Leaves 5-ranked, lobed, dentate or entire, often variable on the same branch, membranaceous or coriaceous, the primary veins prominent and extending to the margins or united within them and connected by more or less reticulate veinlets, deciduous in the autumn or persistent until spring or until their third or fourth year; [238] petioles in falling leaving slightly elevated semiorbicular more or less obcordate leaf-scars broader than high, marked by the ends of numerous scattered fibro-vascular bundles; stipules obovate to lanceolate, scarious, caducous, or those of upper leaves occasionally persistent through the season. Flowers vernal with or after the unfolding of the leaves; staminate solitary in the axils of lanceolate acute caducous bracts, or without bracts, in graceful pendulous clustered aments, from separate or leaf-buds in the axils of leaves of the previous year, or from the axils of the inner scales of the terminal bud or from those of the leaves of the year; calyx campanulate, lobed or divided to the base into 4—7, usually 6, membranaceous lobes; stamens 4—6, rarely 2, or 10—12, inserted on the slightly thickened torus, with free filiform exserted filaments and ovate-oblong or subglobose glabrous or rarely hairy 2-celled usually yellow anthers; pistillate solitary, subtended by a caducous bract and 2 bractlets, in short or elongated few-flowered spikes from the axils of leaves of the year; calyx urn-shaped, with a short campanulate 6-lobed limb, the tube adnate to the incompletely 3 or rarely 4 or 5-celled ovary inclosed more or less completely by an accrescent involucre of imbricated scales, becoming the cup of the fruit; styles as many as the cells of the ovary, short or elongated, erect or incurved, dilated above, stigmatic on the inner face or at apex only, generally persistent on the fruit; ovules anatropous or semianatropous, 2 in each cell. Fruit a nut (acorn) maturing in one or in two years, ovoid, subglobose, or turbinate, short-pointed at apex, 1-seeded by abortion, marked at base by a large conspicuous circular scar, with a thick shell, glabrous or coated on the inner surface with pale tomentum, more or less surrounded or inclosed in the accrescent cupular involucre of the flower (cup), its scales thin or thickened, loosely or closely imbricated. Seed marked at base or at apex or rarely on the side by the abortive ovules; cotyledons thick and fleshy, usually plano-convex and entire.

Quercus inhabits the temperate regions of the northern hemisphere and high altitudes within the tropics, ranging in the New World southward to the mountains of Colombia and in the Old World to the Indian Archipelago. Two hundred and seventy-five species have been described; of the North American species fifty-four are large or small trees. Of exotic species, the European Quercus Robur L., and Quercus sessiliflora Salisb., have been frequently cultivated as ornamental trees in the eastern United States, where, however, they are usually short-lived and unsatisfactory. Many of the species are important timber-trees; their bark is often rich in tannin and is used for tanning leather, and all produce wood valuable for fuel and in the manufacture of charcoal.

CONSPECTUS OF THE NORTH AMERICAN ARBORESCENT SPECIES.

1. Quercus borealis Michx. Red Oak.

Leaves obovate or oblong, acute or acuminate, abruptly or gradually cuneate or rounded at the broad or narrow base, usually divided about half way to the midrib by wide oblique sinuses rounded at the bottom into 11 or sometimes into 7 or 9 acute oblique ovate lobes tapering from broad bases and mostly sinuately 3-toothed at apex with elongated bristle-pointed teeth, or sometimes oblong-obovate, gradually narrowed and cuneate at base, and sinuately lobed with broad acute usually entire or slightly dentate lobes, when they unfold pink, covered with soft silky pale pubescence on the upper surface and [242] below with thick white tomentum, soon glabrous, at maturity thin and firm, dark green, dull and glabrous above, pale yellow-green, glabrous or rarely puberulous and sometimes furnished with small tufts of rusty hairs in the axils of the veins below, 5′—9′ long, 4′—6′ wide; falling early in the autumn after turning dull or sometimes bright orange color or brown; petioles stout, yellow or red, 1′—2′ in length. Flowers: staminate in pubescent aments 4′—5′ long; calyx divided into 4 or 5 narrow ovate rounded lobes shorter than the stamens; pistillate on short glabrous peduncles, their involucral scales broadly ovate, dark reddish brown, shorter than the conspicuous linear acute bract of the flower and as long as the lanceolate acute calyx-lobes; stigmas bright green. Fruit solitary or in pairs, sessile or short-stalked, ovoid, gradually narrowed and acute at apex or cylindric and rounded at apex, pale brown, lustrous, more or less tomentose toward the ends, ½′—1′ long; ½′—¾′ in diameter; cup cup-shaped, puberulous on the inner surface, covered with small closely appressed ovate acute red-brown pubescent scales slightly thickened on the back toward the base of the cup, with a thin dark-colored tip and margins.

A tree usually not more than 60°—70° high, with a trunk 2°—3° in diameter, often much smaller, stout branches forming a narrow head, and slender lustrous branchlets light green and covered with pale scurfy pubescence when they first appear, dark red during their first winter and ultimately dark brown. Winter-buds ovoid, gradually narrowed to the acute apex, about ¼′ long, with thin ovate acute light chestnut-brown scales. Bark on young stems and on the upper part of the limbs of old trees 1′—1½′ thick, dark brown tinged with red and divided into small thick appressed plates scaly on the surface. Wood heavy, hard, strong, close-grained, light reddish brown, with thin lighter-colored sapwood; used in construction, for the interior finish of houses, and in furniture.

Distribution. Nova Scotia and New Brunswick, through Quebec to southern Ontario, and southward to northern New England, western New York, northern Pennsylvania (Presque Isle, Erie County), northern Michigan, southeastern Wisconsin, central Minnesota, central Iowa (Winneshick County), and on the Appalachian Mountains of North Carolina at altitudes of about 4000°. Passing with many intermediate forms differing in the size of the nut and in the depth of the cup into

Quercus borealis var. maxima Ashe. Red Oak.

Fruit solitary or in pairs, sessile or short-stalked; nut ovoid to slightly obovoid, gradually narrowed and rounded at apex, slightly narrowed at base, usually 1′—1¼′ long and ½′—⅔′ thick, occasionally not more than ⅔′ long and thick, inclosed only at the base in a thick saucer-shaped cup.

A tree, usually 70°—80°, or occasionally 150° high, with a trunk 3°—4° in diameter, and stout spreading and ascending branches forming a broad head.

Distribution. Province of Quebec in the neighborhood of Montreal, and southern Ontario, westward through southern Michigan to southeastern Nebraska, and southward to northern Georgia, on the southern Appalachian Mountains up to altitudes of 3000°, southern Kentucky, eastern and central Tennessee, northeastern (Tishomingo County), northwestern (Yazoo County), and central and southern (Hinds and Union Counties) Mississippi, northern and southwestern Alabama (Dekalb, Cullman, Jefferson, and Dallas Counties), northwestern Arkansas, and eastern Kansas and Oklahoma; one of the largest and most generally distributed trees of the northern states; rare and local in the south; of its largest size in the region north of the Potomac and Ohio Rivers.

Often planted as a park and shade tree in the northeastern states and in the counties of western and northern Europe; generally more successful in Europe than other American Oaks.

× Quercus Lowellii Sarg., a possible hybrid of Quercus borealis and Q. ilicifolia, has been found in the neighborhood of Seabury, York County, Maine.

× Quercus Porterii Trel., probably a hybrid of Quercus borealis var. maxima and Q. velutina, has been found on Bowditch Hill, Jamaica Plain, Suffolk County, Massachusetts, on College Hill, Easton, Northampton County, Pennsylvania, and near Columbus, Franklin County, Ohio.

× Quercus runcinata Engelm., believed to be a hybrid of Quercus borealis var. maxima and Q. imbricaria first found near St. Louis, occurs also in the neighborhood of Independence, Jackson County, and at Williamsville, Wayne County, Missouri, and in Richland and Wayne Counties, Illinois.

2. Quercus Shumardii Buckl.

Leaves obovate, seven rarely five-lobed, the lobes two or three-lobed and sometimes dentate at apex, on leaves of lower branches short and broad, and separated by narrow sinuses pointed or rounded in the bottom, on upper branches deeply divided by broad rounded sinuses into narrow acuminate lobes, when they unfold often tinged with red and covered with pale loose tomentum deciduous before they are half grown, at maturity glabrous, dark green and lustrous above, paler and furnished below with large axillary tufts [244] of pale hairs, 6′—8′ long, 4′—5′ wide, with a thin midrib and slender primary veins running to the points of the lobes; petioles slender, glabrous, 2′—2½′ in length. Flowers: staminate in slender glabrous aments 6′—7′ long; calyx divided into 4 or 5 rounded slightly villose lobes shorter than the stamens; pistillate on pubescent peduncles, their involucral scales ovate, light brown, pubescent; stigmas red. Fruit: nut oblong-ovoid, narrowed and rounded at apex, ¾′—1¼′ long, ½′—1′ in diameter, inclosed at the base only in the thick saucer-shaped cup with a slightly incurved rim and covered with closely appressed ovate pale pubescent or nearly glabrous scales narrowed above the middle, abruptly long-pointed, thin or often conspicuously tuberculate.

A tree up to 120° high, with a tall trunk occasionally 5° in diameter, stout wide-spreading branches forming a broad rather open head, and gray or grayish brown glabrous branchlets. Winter-buds ovoid, acute or acuminate, about ¼′ long, with closely imbricated gray glabrous or rarely pubescent scales. Bark 1′—1½′ thick, ridged, broken into small appressed plates scaly on the surface. Wood heavy, hard, close-grained, light reddish brown, often manufactured into lumber in the Mississippi valley and considered more valuable than that of the northern Red Oak.

Distribution. Borders of streams and swamps in moist rich soil; coast region of Texas eastward from the Colorado River and ranging inland up the valley of that river to Burnet County, southeastern Oklahoma, through Arkansas, southeastern Kansas and Missouri to Fayette County, Iowa, southern Illinois and Indiana, the neighborhood of Columbus, Franklin County, Ohio, and southeastern Michigan (near Portage Lake, Jackson County); through the eastern Gulf States to western and central Florida and northward in the neighborhood of the coast to the valley of the Neuse River, North Carolina; Chesapeake Beach, Calvert County, Maryland (W. W. Ashe); ranging inland in the south Atlantic States to Rome, Floyd County, Georgia, Calhoun Falls, Abbeville County, and Columbia, Richland County, South Carolina, and Chapel Hill, Orange County, North Carolina. Passing into

Quercus Shumardii var. Schneckii Sarg.

Differing from the type in the deep cup-shaped cup of the fruit covered with thin scales, rarely much thickened and tuberculate at base (only on river banks near Vicksburg, [245] Warren County, Mississippi), and connected with it by forms with the cups of the fruit differing from saucer to deep cup-shaped.

Distribution. Growing with Quercus Shumardii; more common in Texas and in the Mississippi valley than the type, and ranging eastward through Louisiana and Mississippi to central and southern Alabama, central and southeastern Tennessee (neighborhood of Chattanooga), and central Kentucky; apparently not reaching the Atlantic States.

3. Quercus texana Buckl.

Leaves widest above the middle, broad-cuneate, concave-cuneate or nearly truncate at base, deeply or rarely only slightly divided by broad sinuses rounded in the bottom into 5 or 7 lobes, the terminal lobe 3-lobed and acute at apex, the upper lateral lobes broad and more or less divided at apex and much larger and more deeply lobed than those of the lowest pair, when they unfold densely covered with fascicled hairs and often bright red, soon glabrous, thin, dark green and lustrous above, pale and lustrous and rarely furnished below with small inconspicuous axillary tufts of pale hairs, 3′—3½′ long, 2½′—3′ wide, with a thin midrib and slender primary veins running to the points of the lobes; petioles slender, soon glabrous, ¾′—1½′ in length. Flowers: staminate in slender villose aments 3′—4′ long; calyx thin, villose on the outer surface, divided into 4 or 5 acute lobes shorter than the stamens; pistillate on short hoary tomentose peduncles, their involucral scales brown tinged with red; stigmas bright red. Fruit short-stalked, usually solitary; nut ovoid, narrowed and rounded at apex, light red-brown, often striate, ¼′—¾′ long and broad, sometimes acute, nearly 1′ in length and not more than ⅓′ in diameter; cup turbinate, covered with thin ovate acuminate slightly appressed glabrous scales, in the small fruit of trees on dry hills inclosing a third or more of the nut, in the larger fruit of trees on better soil comparatively less deep.

A tree on dry hills rarely more than 30° tall, with a trunk 8′—10′ in diameter, small spreading or erect branches and slender red or reddish brown glabrous or rarely pubescent branchlets; often a shrub; on better soil at the foot of hills occasionally 50° high with a trunk 12′—18′ in diameter. Winter-buds ovoid, acute, ⅙′—¼′ long and covered with closely imbricated acute slightly or densely pubescent red scales. Bark light brown tinged with red, ¾′—1′ thick, deeply ridged and broken into plate-like scales.

Distribution. Dry limestone hills and ridges, and in the more fertile soil at their base; central and western Texas (Dallas, Tarrant County to Travis and Bexar Counties), and to the Edwards Plateau (San Saba, Kerr, Brown, Coke and Uvalde Counties); westward [246] replaced by the var. chesosensis Sarg. differing in the acuminate lobes of the leaves and smaller cups of the fruit; known only on the dry rocky slopes of the Chesos Mountains, Brewster County, Texas; and by the var. stellapila Sarg., differing in the presence of fascicled hairs on both surfaces of the mature leaves and on the branchlets of the year; above Fort Davis, Jeff Davis County, Texas.

4. Quercus ellipsoidalis E. J. Hill. Black Oak.

Leaves elliptic to obovate-orbicular, acute or acuminate, truncate or broadly cuneate at base, deeply divided by wide sinuses rounded in the bottom into 5—7 oblong lobes repandly dentate at apex, or often, especially those of the upper pair, repandly lobulate, when they unfold slightly tinged with red and hoary-tomentose, soon becoming glabrous with the exception of small tufts of pale hairs in the axils of the principal veins, at maturity thin and firm, bright green and lustrous above, paler and sometimes entirely glabrous below, 3′—5′ long, 2½′—4′ wide, with a stout midrib and primary veins and prominent reticulate veinlets; late in the autumn turning yellow or pale brown more or less blotched with purple; petioles slender, glabrous or rarely puberulous, 1½′—2′ in length. Flowers: staminate in puberulous aments 1½′—2′ long; calyx campanulate, usually tinged with red, 2—5-lobed or parted into oblong-ovate or rounded segments, glabrous or slightly villose, fringed at apex with long twisted hairs, about as long as the 2—5 stamens, with short filaments and oblong anthers; pistillate on stout tomentose 1—3-flowered peduncles, red, their involucral scales broad, oblong, acute, hairy; calyx campanulate, 4—7-lobed, ciliate on the margins. Fruit short-stalked or nearly sessile, solitary or in pairs; nut ellipsoidal to subglobose, chestnut-brown, often striate and puberulous, inclosed for one third to one half its length in a turbinate or cup-shaped cup gradually narrowed at base, thin, light red-brown, and covered by narrow ovate obtuse or truncate brown pubescent closely appressed scales.

A tree, 60°—70° high, with a short trunk rarely 3° in diameter, much forked branches ascending above and often pendulous low on the stem, forming a narrow oblong head, and slender branchlets covered at first with matted pale hairs, bright reddish brown during their first winter, becoming dark gray-brown or reddish brown in their second season. Winter-buds ovoid, obtuse or acute, sometimes slightly angled, about ⅛′ long, with ovate or oval red-brown lustrous slightly puberulous outer scales ciliate on the margins. Bark thin, light yellow internally, close, rather smooth, divided by shallow connected fissures into thin plates, dark brown near the base of the tree, dull above, gray-brown and only slightly furrowed on the large branches.

Distribution. In the neighborhood of Chicago, Illinois, to southeastern Minnesota common; often covering large areas of sandy soil with a stunted growth and on the prairies sometimes a low shrub; eastern Iowa (Muscatine County), and the Lower Peninsular of Michigan (Montmorency, Arenac, and St. Clair Counties).

5. Quercus coccinea Muench. Scarlet Oak. Spanish Oak.

Leaves oblong-obovate or elliptic, truncate or cuneate at base, deeply divided by wide sinuses rounded in the bottom into 7 or rarely 9 lobes repand-dentate at apex, the terminal lobe, ovate, acute, and 3-toothed, the middle division the largest and furnished with 2 small lateral teeth, the lateral lobes obovate, oblique or spreading, sometimes falcate, usually broad and oblique at the coarsely toothed apex, when they unfold bright red covered with loose pale pubescence above and below with silvery white tomentum, green at the end of a few days, at maturity thin and firm, bright green, glabrous and very lustrous above, paler and less lustrous and sometimes furnished with small tufts of rusty pubescence in the axils of the veins below, 3′—6′ long, 2½′—4′ broad, with a yellow midrib and primary veins, late in the autumn turning brilliant scarlet; petioles slender, terete, 1½′—2½′ in length. Flowers: staminate in slender glabrous aments 3′—4′ long; calyx pubescent, bright red before opening, divided into 4 or 5 ovate acute segments shorter than the stamens; pistillate on pubescent peduncles sometimes ½′ long, bright red, their involucral scales ovate, pubescent, shorter than the acute calyx-lobes. Fruit sessile or stalked, solitary or in pairs; nut oval, oblong-ovate or hemispheric, truncate or rounded at base, rounded at apex, ½′—1′ long, ⅓′—⅔′ thick, light reddish brown and occasionally striate, inclosed for one third to one half its length in a deep cup-shaped or turbinate thin cup light reddish brown on the inner surface, covered by closely imbricated oblong-ovate acute thin, or rarely much thickened (var. tuberculata Sarg.) light reddish brown slightly puberulous scales.

A tree, 70°—80° high, with a trunk 2°—3° in diameter, comparatively small branches spreading gradually and forming a rather narrow open head, and slender branchlets coated at first with loose scurfy pubescence, soon pale green and lustrous, light red or orange-red in their first winter and light or dark brown the following year; usually much smaller. Winter-buds ellipsoidal or ovoid, gradually narrowed at apex, ⅛′—¼′ long, dark reddish brown, and pale-pubescent above the middle. Bark of young stems and branches smooth, light brown, becoming on old trunks ½′—1′ thick and divided by shallow fissures into irregular ridges covered by small light brown scales slightly tinged with red. Wood heavy, hard, strong, coarse-grained, light or reddish brown, with thicker darker colored sapwood.

Distribution. Light dry usually sandy soil; valley of the Androscoggin River, Maine, [248] southern New Hampshire and Vermont to southern Ontario, southward to the District of Columbia and along the Appalachian Mountains to eastern Kentucky and Tennessee, and northern Georgia; in central Georgia and northeastern Mississippi (near Corinth, Alcorn County), and westward through New York, Ohio, Indiana, Illinois and southern Wisconsin to central Missouri (Jerome, Phelps County); in eastern Oklahoma (Arkansas River valley near Fisher, Creek County, G. W. Stevens); ascending to altitudes of nearly 5000° on the southern mountains; the prevailing Oak above 2500° to the summits of the Blue Ridge of the Carolinas; very abundant in the coast region from Massachusetts Bay to southern New Jersey; less common in the interior, growing on dry gravelly uplands, and on the prairies skirting the western margins of the eastern forest.

Occasionally planted in the northeastern states and in Europe as an ornamental tree valued chiefly for the brilliant autumn color of the foliage.

× Quercus Robbinsii Trel., believed to be a hybrid of Quercus coccinea and Q. ilicifolia, occurs at North Easton, Bristol County, Massachusetts.

× Quercus Benderi Baenitz, a supposed hybrid of Quercus coccinea and Q. borealis var. maxima, appeared several years ago in Silesia, and a similar tree has been found in the Blue Hills Reservation near Boston.

6. Quercus palustris Muench. Pin Oak. Swamp Spanish Oak.

Leaves obovate, narrowed and cuneate or broad and truncate at base, divided by wide deep sinuses rounded in the bottom into 5—7 lobes, the terminal lobe ovate, acute, 3-toothed toward the apex, or entire, the lateral lobes spreading or oblique, sometimes falcate, especially those of the lowest pair, gradually tapering and acute at the dentate apex, or obovate and broad at apex, when they unfold light bronze-green stained with red on the margins, lustrous and puberulous above, coated below and on the petioles with pale scurfy pubescence, at maturity thin and firm, dark green and very lustrous above, pale below, with large tufts of pale hairs in the axils of the conspicuous primary veins; 4′—6′ long, 2′—4′ wide, with a stout midrib; late in the autumn gradually turning deep scarlet; petioles slender, yellow, ½′—2′ in length. Flowers: staminate in hairy aments 2′—3′ long; calyx puberulous and divided into 4 or 5 oblong rounded segments more or less laciniately cut on the margins, shorter than the stamens; pistillate on short tomentose peduncles, their involucral scales broadly ovate, tomentose, shorter than the acuminate calyx-lobes; stigmas bright red. Fruit sessile or short-stalked, solitary or clustered; nut nearly hemispheric, about ½′ in diameter, light brown, often striate, inclosed only at the base in a thin saucer-shaped [249] cup dark red-brown and lustrous within, and covered by closely appressed ovate light red-brown thin puberulous scales.

A tree, usually 70°—80° high, with a trunk 2°—3° in diameter, often clothed with small tough drooping branches, or when crowded in the forest sometimes 120° high, with a trunk 60°—70° tall and 4°—5° in diameter, slender branches beset with short-ridged spur-like laterals a few inches in length, forming on young trees a broad pyramidal head, becoming on older trees open and irregular, with rigid and more pendulous branches often furnished at first with small drooping branchlets, and slender tough branchlets dark red and covered by short pale silvery tomentum, soon becoming green and glabrous, lustrous dark red-brown or orange color in their first winter, growing darker in their second year and ultimately dark gray-brown. Winter-buds ovoid, gradually narrowed and acute at apex, about ⅛′ long, with imbricated light chestnut-brown scales puberulous toward the thin sometimes ciliate margins. Bark of young trunks and branches smooth, lustrous, light brown frequently tinged with red, becoming on older trunks ¾′—1¼′ thick, light gray-brown, generally smooth and covered by small closely appressed scales. Wood heavy, hard, strong, coarse-grained, light brown, with thin rather darker colored sapwood; sometimes used in construction, and for shingles and clapboards.

Distribution. Borders of swamps and river-bottoms in deep rich moist soil; valley of the Connecticut River in western Massachusetts and Connecticut; on Grand Isle in the Niagara River, New York to southern Ontario and southwestern Michigan, and westward to eastern Iowa (Muscatine County), and southward to southern West Virginia (Hardy and Mercer Counties), southwestern Virginia (Wythe County), central North Carolina (on Bowling’s Creek, near Chapel Hill, Orange County, and on Dutchman’s Creek, Forsyth County); and to southern Kentucky, central Tennessee, southern Arkansas (Fulton, Hempstead County), and northeastern Oklahoma; rare and of small size in New England; exceedingly common on the coast plain south of the Hudson River; very abundant on the bottom-lands of the streams of the lower Ohio River.

Often cultivated as an ornamental tree in the northeastern states and occasionally in the countries of western and central Europe.

7. Quercus georgiana M. A. Curtis.

Leaves convolute in the bud, elliptic or obovate, gradually narrowed and cuneate at base, divided generally about half way to the midrib by wide or narrow oblique sinuses rounded in the bottom into 3—7 lobes, the terminal lobe ovate, acute, or rounded and entire or frequently furnished with 1 or 2 small lateral teeth, the lateral lobes oblique or [250] spreading, mostly triangular, acute and entire, or those of the upper and of the middle pair often broad and repand-lobulate at the oblique ends, sometimes gradually 3-lobed at the broad apex and narrowed and entire below, or equally 3-lobed, with broad or narrow spreading lateral lobes, or occasionally pinnatifid, when they unfold bright green tinged with red, ciliate on the margins and coated on the midrib, veins, and petioles with loose pale pubescence, at maturity thin, bright green and lustrous above, paler below, and glabrous or furnished with tufts of hairs in the axils of the primary veins, usually about 2½′ long and 1½′ wide; turning dull orange and scarlet in the autumn; petioles slender, ½′—¾′ in length. Flowers: staminate in slender glabrous or pubescent aments 2′—3′ long; calyx divided into 4 or 5 broadly ovate rounded segments rather shorter than the stamens; pistillate on short glabrous slender peduncles; their involucral scales rather shorter than the acute calyx-lobes, pubescent or puberulous; stigmas bright red. Fruit short-stalked; nut ellipsoidal or subglobose, ⅓′—½′ long, light red-brown and lustrous, inclosed for one third to nearly one half its length in a thick cup-shaped cup light red-brown and lustrous on the inner surface, and covered by thin ovate bright light red-brown truncate erose scales.

Distribution. Georgia; on Stone Mountain, and Little Stone Mountain, Dekalb County; on a few other granite hills between the Yellow and Oconee rivers in the region south and east of Stone Mountain (Winder, Jackson County, Rockmart, Polk County and at Warm Springs, Meriwether County).

× Quercus Smallii Trel., a possible hybrid of Quercus georgiana and Q. marilandica, occurs on the slopes and summit of Little Stone Mountain, Dekalb County, Georgia.

8. Quercus velutina Lam. Black Oak. Yellow-bark Oak.

Leaves ovate or oblong, rounded, cuneate or truncate at base, mostly 7-lobed and sometimes divided nearly to the middle by wide rounded sinuses into narrow obovate more or less repand-dentate lobes, or into elongated nearly entire mucronate lobes tapering gradually from a broad base, the terminal lobe oblong, elongated, acute, furnished with small lateral teeth, or broad, rounded, and coarsely repand-dentate, or slightly divided into broad dentate lobes or sinuate-dentate, bright crimson when they unfold, and covered above by long loose scattered white hairs and below with thick pale or silvery white tomentum, hoary-pubescent when half grown, and at maturity thick and firm or subcoriaceous, dark green and lustrous above, below yellow-green, brown or dull copper color and more or less pubescent or glabrous with the exception of tufts of rusty hairs in the axils of the principal veins, 3′—12′ long and 2′—10′ wide, but usually 5′—6′ long and 3′—4′ wide, with a stout midrib and primary veins; late in the autumn turning dull red, dark orange color, or brown, and falling gradually during the winter; petioles stout, yellow, glabrous or puberulous, 3′—6′ in length. Flowers: staminate in tomentose or pubescent aments 4′—6′ long; calyx coated with pale hairs, with ovate acute lobes; pistillate on short tomentose peduncles, their involucral scales ovate, shorter than the acute calyx-lobes; stigmas bright red. Fruit sessile or short-stalked, solitary or in pairs; nut ovoid-oblong, obovoid, oval or hemispheric, broad and rounded at base, full and rounded at apex, light red-brown, often striate, frequently coated with soft rufous pubescence, ½′—¾′ long and broad, or rarely 1′ long and broad, inclosed for about half its length or rarely nearly to the apex in the thin deeply cup-shaped or turbinate cup dark red-brown on the inner surface, covered by thin light chestnut-brown acute hoary scales closely appressed at the base of the cup, loosely imbricated above the middle, with free scarious tips forming a fringe-like border to its rim.

A tree, often 70°—80° and occasionally 150° high, with a trunk 3°—4° in diameter, slender branches spreading gradually into a narrow open head, stout branchlets coated at first with pale or fulvous scurfy tomentum, becoming in their first winter glabrous, dull red or reddish brown, growing dark brown in their second year or brown slightly tinged with red. Winter-buds ovoid, strongly angled, gradually narrowed and obtuse at apex, hoary-tomentose, ¼′—½′ long. Bark of young stems and branches smooth, dark brown, deep [251] orange color internally, becoming ¾′—1½′ thick on old trunks, and deeply divided into broad rounded ridges broken on the surface into thick dark brown or nearly black closely appressed plate-like scales. Wood heavy, hard, strong, coarse-grained, bright brown tinged with red, with thin lighter colored sapwood; of little value except as fuel. The bark abounds in tannic acid and is largely used in tanning, as a yellow dye, and in medicine.

Distribution. Dry gravelly uplands and ridges; coast of southern Maine to northern Vermont, southern and western Ontario, the southern peninsula of Michigan, northwestern, eastern and southern Iowa, and southeastern Nebraska, and southward to western Florida, southern Alabama, Mississippi, Louisiana, eastern Kansas, northeastern Oklahoma and eastern Texas to the valley of the Brazos River; one of the commonest Oaks on the gravelly drift of southern New England and the middle states; ascending on the southern Appalachian Mountains to altitudes of about 4000°, and often forming a large part of the forest growth on their foothills; abundant in all parts of the Mississippi basin, and of its largest size in the valley of the lower Ohio River; the common species of the Black Oak group reaching the south-Atlantic and Gulf Coast, and here generally scattered on dry ridges through the maritime Pine belt.

Quercus velutina, which is more variable in the form of its leaves than the other North American Black Oaks, is easily recognized by the bright yellow color of the inner bark, in early spring by the deep red color of the unfolding leaves, becoming pale and silvery in a few days, and by the large tomentose winter-buds. From western Missouri to northwestern Arkansas a form occurs (var. missouriensis Sarg.) with the mature leaves covered above with fascicled hairs, and coated below and on the petioles and summer branchlets with rusty pubescence, and with broader more loosely imbricated hoary-tomentose cup-scales.

9. Quercus Kelloggii Newb. Black Oak.

Leaves oblong or obovate, truncate, cuneate or rounded at the narrow base, 7 or rarely 5-lobed by wide and deep or shallow and oblique sinuses rounded in the bottom, the terminal lobe ovate, 3-toothed at the acute apex, the lateral lobes tapering gradually from the base or broad and obovate, coarsely repand-dentate with acute pointed teeth, or rarely entire, when they unfold dark red or purple and pilose above and coated below and on the petioles with thick silvery white tomentum, at maturity thick and firm, [252] lustrous, dark yellow-green and glabrous or rarely pubescent above, light yellow-green or brownish and glabrous or pubescent, or occasionally hoary-tomentose below, 3′—6′ long, 2′—4′ wide; turning yellow or brown in the autumn before falling; petioles slender, yellow, 1′—2′ in length. Flowers: staminate in hairy aments 4′—5′ long; calyx pubescent, divided into 4 or 5 ovate acute segments shorter than the stamens; anthers bright red; pistillate on short tomentose peduncles, their involucral scales ovate, coated like the acute calyx-lobes with pale tomentum; stigmas dark red. Fruit short-stalked, solitary or clustered; nut oblong, ellipsoidal or obovoid, broad and rounded at base, full and rounded or gradually narrowed and acute at the puberulous apex, 1′—1½′ long, about ¾′ broad, light chestnut-brown, often striate, inclosed for one fourth to two thirds of its length in the deep cup-shaped cup light brown on the inner surface, and covered by thin ovate-lanceolate lustrous light chestnut brown scales, sometimes rounded and thickened on the back toward the base of the cup, their tips elongated, thin and erose on the margins, often forming a narrow fringe-like border to the rim of the cup.

A tree, occasionally 100° high, with a trunk 3°—4° in diameter, stout spreading branches forming an open round-topped head, and branchlets coated at first with thick hoary caducous tomentum, bright red or brown tinged with red, and usually glabrous or pubescent or puberulous during their first winter, becoming dark red-brown in their second year; frequently much smaller and at high elevations a small shrub (f. cibata Jeps.). Winter-buds ovoid, gradually narrowed and acute at apex, about ¼′ long, with closely imbricated pale chestnut-brown scales ciliate on the thin scarious margins and pubescent toward the point of the bud. Bark of young stems and branches smooth, light brown, becoming on old trunks 1′—1½′ thick, dark brown slightly tinged with red or nearly black, divided into broad ridges at the base of old trees and broken above into thick irregular oblong plates covered by minute closely appressed scales. Wood heavy, hard, strong, very brittle, bright red, with thin lighter colored sapwood; occasionally used as fuel.

Distribution. Valleys and mountain slopes; basin of the Mackenzie River in western Oregon, southward over the California coast ranges, and along the western slopes of the Sierra Nevada up to altitudes of 6500° to the Cuyamaca Mountains near the southern boundary of California; extending across the Sierra Nevada to the foothills of Owens valley (Jepson) in eastern California; rare in the immediate neighborhood of the coast; the largest and most abundant Oak-tree of the valleys of southwestern Oregon and of the Sierra Nevada, sometimes forming groves of considerable extent in coniferous forests; of its largest size at altitudes of about 6000° above the sea.

10. Quercus Catesbæi Michx. Turkey Oak.

Leaves oblong or obovate or nearly triangular, gradually narrowed and cuneate at base, deeply divided by wide rounded sinuses into 3 or 5 or rarely 7 lobes, the terminal lobe ovate, elongated, acute and entire or repand-dentate, or obovate and coarsely equally or irregularly 3-toothed at apex, the lateral lobes spreading, usually falcate, entire and acute, tapering from the broad base, and broad, oblique, and repand-lobulate at apex, or 3-toothed at the broad apex and gradually narrowed to the base, coated when they unfold with rufous fascicled hairs, and when fully grown thick and rigid, bright yellow-green and lustrous above, paler, lustrous, and glabrous below, with large tufts of rusty hairs in the axils of the veins, 3′—12′ long, 1′—10′ wide, but usually about 5′ long and wide, with a broad yellow or red-brown midrib; turning bright scarlet before falling in the late autumn or early winter; petioles stout, grooved, ¼′—¾′ in length. Flowers: staminate in slender hairy red-stemmed aments 4′—5′ long; calyx puberulous and divided into 4 or 5 ovate acute lobes; pistillate on short stout tomentose peduncles, their involucral scales bright red, pubescent, hairy at the margins; stigmas dark red. Fruit short-stalked, usually solitary; nut oval, full and rounded at the ends, about 1′ long and ¾′ broad, dull light brown, covered at the apex by a thin coat of snow-white tomentum, inclosed for about one third its length in a thin turbinate cup often gradually narrowed into a stout stalk-like base, light red-brown and lustrous on the inner surface, covered by ovate-oblong rounded scales extending above the rim of the cup and down over the upper third of the inner surface, and hoary-pubescent except their thin bright red margins.

A tree, usually 20°—30°, or occasionally 50°—60° high, with a trunk rarely exceeding 2° in diameter, stout spreading more or less contorted branches forming a broad or narrow open irregular generally round-topped head, and stout branchlets coated at first with fascicled hairs, nearly glabrous and deep red when the leaves are half grown, dark red in their first winter, gradually growing dark brown; generally much smaller and sometimes shrubby. Winter-buds elongated, acute, ½′ long, with light chestnut-brown scales erose on the thin margins, and coated, especially toward the point of the bud, with rusty pubescence. Bark ½′—1′ thick, red internally, dark gray tinged with red on the surface, and at the base of old trunks becoming nearly black, deeply and irregularly furrowed and broken into small appressed scales. Wood heavy, hard, strong, rather close-grained, light brown tinged with red, with thick lighter colored sapwood; largely used for fuel.

Distribution. Dry barren sandy ridges and sandy bluffs and hummocks in the neighborhood of the coast; southeastern Virginia (near Zuni, Isle of Wight County) to the shores [254] of Indian River and Peace Creek, Florida, and westward to eastern Louisiana; comparatively rare toward the western limits of its range, and most abundant and of its largest size on the high bluff-like shores of bays and estuaries in South Carolina and Georgia; the prevailing tree with Quercus cinerea in the flat woods of the interior of the Florida peninsula as far south as the sandy ridges in the neighborhood of Lake Istokpoga, De Soto County.

× Quercus Mellichampii Trel. believed to be a hybrid of Quercus Catesbæi and Q. laurifolia occurs at Bluffton on the coast of South Carolina, in the neighborhood of Orlando, Orange County and near San Mateo, Putnam County, Florida.

× Quercus Ashei Trel. believed to be a hybrid of Quercus Catesbæi with Q. cinerea occurs at Folkston and near Trader’s Hill, Charlton County and St. Mary’s, Camden County, Georgia.

× Quercus blufftonensis Trel., a probable hybrid of Quercus Catesbæi and Q. rubra L., has been found at Bluffton, South Carolina.

× Quercus Walteriana Ashe, believed to be a hybrid of Quercus Catesbæi and Q. nigra, is not rare in the immediate neighborhood of the coast of South Carolina and Georgia, and occurs on sand hills in Sampson County, North Carolina, near Jacksonville, Duval County, Florida, at Mount Vernon, Mobile County and in the neighborhood of Selma, Dallas County, Alabama.

11. Quercus ilicifolia Wang. Bear Oak. Scrub Oak.

Leaves obovate or rarely oblong, gradually or abruptly cuneate at base, divided by wide shallow sinuses into 3—7, usually 5, acute lobes, the terminal lobe ovate, elongated, rounded and 3-toothed or acute and dentate or entire at apex, the lateral lobes spreading, mostly triangular and acute, or those of the upper pair broad, oblique and repand-lobulate or broad at apex, slightly 3-lobed and entire below, or deeply 3-lobed above and sinuate below, or occasionally oblong to oblong-obovate and entire, with undulate margins, when they unfold dull red and puberulous or pubescent on the upper surface and coated on the lower and on the petioles with thick pale tomentum, with conspicuous tufts of silvery white hairs in the axils of the veins, at maturity thick and firm, dark green and lustrous above, covered below with pale or silvery white pubescence, 2′—5′ long, 1½′—3′ wide, [255] with a stout yellow midrib and slender primary veins; turning dull scarlet or yellow in the autumn; petioles slender, glabrous, or pubescent, 1′—1½′ in length. Flowers: staminate in hairy aments 4′—5′ long, and often persistent until midsummer; calyx red or green tinged with red and irregularly divided into 3—5 ovate rounded lobes shorter than the stamens; anthers bright red ultimately yellow; pistillate on stout tomentose peduncles, their involucral scales ovate, about as long as the acute calyx-lobes, red and tomentose; stigmas dark red. Fruit produced in great profusion, sessile or stalked, in pairs or rarely solitary; nut ovoid, broad, flat or rounded at base, gradually narrowed and acute or rounded at apex, about ½′ long and broad, light brown, lustrous, usually faintly striate, inclosed for about one half its length in the cup-shaped or saucer-shaped cup often abruptly enlarged above the stalk-like base, thick, light reddish brown within, and covered by thin ovate closely imbricated red-brown puberulous scales acute or truncate at apex, the minute free tips of the upper scales forming a fringe-like border to the cup.

A tree, occasionally 18°—20° high, with a trunk 5′—6′ in diameter, with slender spreading branches usually forming a round-topped head, and slender branchlets dark green more or less tinged with red and hoary-pubescent at first, during their first winter red-brown or ashy gray and pubescent or puberulous, becoming glabrous and darker in their second year and ultimately dark brown or nearly black; more frequently an intricately branched shrub, with numerous contorted stems 3°—10° tall. Winter-buds ovoid, obtuse, about ⅛′ long, with dark chestnut-brown rather loosely imbricated glabrous or pilose scales. Bark thin, smooth, dark brown, covered by small closely appressed scales.

Distribution. Dry sandy barrens and rocky hillsides; coast of eastern Maine southward through eastern and southern New England to southern and southwestern Pennsylvania and along the Appalachian Mountains, principally on their eastern slopes, to southern Virginia; on Crowder and King Mountains, Gaston County, North Carolina; and westward to the shores of Lake George and the valley of the Hudson River; common in eastern and southern New England, in the Pine barrens of New Jersey, and in eastern Pennsylvania.

× Quercus Brittonii Davis, believed to be a hybrid of Quercus ilicifolia and Q. marilandica, has been found on Staten Island, New York, and at Ocean Grove, Monmouth County, New Jersey.

× Quercus Giffordii Trel., believed to be a hybrid of Quercus ilicifolia and Q. Phellos, has been found at May’s Landing, Atlantic County, New Jersey.

× Quercus Rehderi Trel., believed to be a hybrid of Quercus ilicifolia and Q. velutina, is not rare in eastern Massachusetts and occurs on Martha’s Vineyard (Chilmark).

12. Quercus rubra L. Red Oak. Spanish Oak.

Leaves ovate to obovate, narrowed and rounded or cuneate at base, the terminal lobe long-acuminate, entire or slightly lobed, often falcate, usually longer than the 2 or 4 acuminate entire lateral lobes narrowed from a broad base and often falcate, or oblong-obovate and divided at the broad apex by wide or narrow sinuses broad and rounded in the bottom into 3 rounded or acute entire or dentate lobes, and entire and gradually narrowed below into an acute or rounded base (var. triloba Ashe), the two forms usually occurring on different but sometimes on the same tree, at maturity thin and firm, dark green and lustrous above, coated below with soft close pale or rusty pubescence, 6′—7′ long and 4′—5′ wide, obscurely reticulate-venulose, with a stout tomentose midrib and primary veins; turning brown or dull orange color in the autumn; petioles slender, flattened, 1′—2′ in length. Flowers: staminate in tomentose aments, 3′—5′ long; calyx thin and scarious, pubescent on the outer surface, divided into 4 or 5 ovate rounded segments; pistillate on stout tomentose peduncles, their involucral scales coated with rusty tomentum, as long or rather shorter than the acute calyx-lobes; stigmas dark red. Fruit sessile or short-stalked; nut subglobose to ellipsoidal, full and rounded at apex, truncate and rounded at base, about [256] ½′ long, bright orange-brown, inclosed only at base or sometimes for one third its length in a thin saucer-shaped cup flat on the bottom or gradually narrowed from a stalk-like base, or deep and turbinate, bright red-brown on the inner surface, covered by thin ovate-oblong reddish scales acute or rounded at apex and pale-pubescent except on the margins.

A tree, usually 70°—80° high, with a trunk 2°—3° in diameter, large spreading branches forming a broad round-topped open head, and stout branchlets coated at first, like the young leaves, with thick rusty or orange-colored clammy tomentum, dark red or reddish brown and pubescent or rarely glabrous during their first winter, becoming in their second year dark red-brown or ashy gray. The var. triloba usually 20°—30° rarely 40°—50° high. Winter-buds ovoid or oval, acute, ⅛′—¼′ long, with bright chestnut-brown puberulous or pilose scales ciliate with short pale hairs. Bark ¾′—1′ thick, dark brown or pale, and divided by shallow fissures into broad ridges covered by thin closely appressed scales. Wood hard, strong, not durable, coarse-grained, light red, with thick lighter colored sapwood; sometimes used in construction, and largely as fuel. The bark is rich in tannin, and is used in tanning leather and occasionally in medicine.

Distribution. Southeastern and southern Pennsylvania and southern New Jersey southward to central Florida, through the Gulf states to the valley of the Brazos River, Texas, and through eastern Oklahoma and southwestern Missouri to central Tennessee and Kentucky, southern Indiana and Illinois, southern Ohio (Black Fork Creek, Lawrence County), and Kanawha County, West Virginia; in the north Atlantic states only in the neighborhood of the coast and comparatively rare; very common in the south Atlantic and Gulf states on dry hills between the coast plain and the Appalachian Mountains; less abundant in the southern maritime Pine belt. The var. triloba: rare and local. Pleasant Grove, Lancaster County, Pennsylvania and Jefferson County, Indiana, southward to central and western Florida, southern Alabama and Mississippi, western Arkansas and eastern Texas; on dry uplands near Milledgeville, Baldwin County, Georgia, the prevailing form.

Quercus rubra var. pagodæfolia Ashe. Swamp Spanish Oak. Red Oak.

Leaves elliptic to oblong, acuminate, gradually narrowed and cuneate or full and rounded or rarely truncate at base, deeply divided by wide sinuses rounded in the bottom into 5—11 acuminate usually entire repand-dentate lobes often falcate and spreading at right angles [257] to the midrib or pointed toward the apex of the leaf, when they unfold coated with pale tomentum, thickest on the lower surface, and dark red on the upper surface, at maturity dark green and very lustrous above, pale and tomentose below, 6′—8′ long and 5′—6′ wide, with a stout midrib usually puberulous on the upper side, slender primary veins arched to the points of the lobes, and conspicuous reticulate veinlets; turning bright clear yellow before falling; petioles stout, pubescent or tomentose, 1½′—2′ in length. Flowers and Fruit as in the species.

A tree, sometimes 120° high, with a trunk 4°—5° in diameter, heavy branches forming in the forest a short narrow crown, or in more open situations wide-spreading or ascending and forming a great open head, and slender branchlets hoary tomentose at first, tomentose or pubescent during their first winter, and dark reddish brown and puberulous during their second year. Winter-buds ovoid, acute, often prominently 4-angled, about ¼′ long, with light red-brown puberulous scales sometimes ciliate at the apex. Bark about 1′ thick and roughened by small rather closely appressed plate-like light gray, gray-brown or dark brown scales. Wood light reddish brown, with thin nearly white sapwood; largely manufactured into lumber in the Mississippi valley, and valued almost as highly as white oak.

Distribution. Rich bottom-lands and the alluvial banks of streams; Maryland (Queen Anne County) and coast of Virginia to northern Florida, and through the Gulf states and Arkansas to southern Missouri, western Tennessee and Kentucky, and southern Illinois and Indiana; most abundant and one of the largest and most valuable timber-trees in the river swamps of the Yazoo basin, Mississippi, and of eastern Arkansas. Differing chiefly from the type in the more numerous and more acuminate lobes of the usually more elongated leaves usually paler on the lower surface, and in the generally paler bark of the trunk; passing into Quercus rubra var. leucophylla Ashe with leaves on upper branches nearly as broad as long thickly covered below with brownish pubescence and deeply divided into 5—7 lobes, and on lower branches slightly obovate, less deeply divided, thin, dark green, sometimes pubescent becoming glabrous above and often covered below with pale or brown pubescence.

A tree sometimes 120° high; in low rich soil; coast region of southeastern Virginia, southward to western Florida and through the Gulf states to the valley of the Neches River, Texas, and northward to northern Arkansas; in southern Illinois (near Mt. Carmel, Wabash County) and southwestern Indiana (near Hovey Lake, Posey County); abundant in low woods about River Junction, Gadsden County, Florida, and in central Mississippi.

× Quercus Willdenoviana Zabel is believed in Europe to be a hybrid of Quercus rubra and Quercus velutina.

13. Quercus marilandica Muench. Black Jack. Jack Oak.

Leaves broadly obovate, rounded or cordate at the narrow base, usually 3 or rarely 5-lobed at the broad and often abruptly dilated apex, with short or long, broad or narrow, rounded or acute, entire or dentate lobes, or entire or dentate at apex, sometimes oblong-obovate, undulate-lobed at the broad apex and entire below, or equally 3-lobed with elongated spreading lateral lobes broad and lobulate at apex, when they unfold coated with a clammy tomentum of fascicled hairs and bright pink on the upper surface, at maturity thick and firm or subcoriaceous, dark yellow-green and very lustrous above, yellow, orange color, or brown and scurfy-pubescent below, usually 6′—7′ long and broad, with a thick broad orange-colored midrib; turning brown or yellow in the autumn; petioles stout, yellow, glabrous or pubescent, ½′—¾′ in length. Flowers: staminate in hoary aments 2′—4′ long; calyx thin and scarious, tinged with red above the middle, pale-pubescent on the outer surface, divided into 4 or 5 broad ovate rounded lobes; anthers apiculate, dark red; pistillate on short rusty-tomentose peduncles coated like their involucral scales with thick rusty tomentum; stigmas dark red. Fruit, solitary or in pairs, usually pedunculate; nut oblong, full and rounded at the ends, rather broader below than above the middle, about ¾′ long, light yellow-brown and often striate, the shell lined with dense fulvous tomentum, inclosed for one third to nearly two thirds of its length in a thick turbinate light brown cup puberulous on the inner surface, and covered by large reddish brown loosely imbricated scales often ciliate and coated with loose pale or rusty tomentum, the upper scales smaller, erect, inserted on the top of the cup in several rows, and forming a thick rim round its inner surface, or occasionally reflexed and covering the upper half of the inner surface of the cup.

A tree, 20°—30°, or occasionally 40°—50° high, with a trunk rarely more than 1′ in diameter, short stout spreading often contorted branches forming a narrow compact round-topped or sometimes an open irregular head, and stout branchlets coated at first with thick pale tomentum, light brown and scurfy-pubescent during their first summer, becoming reddish brown and glabrous or puberulous in the winter, and ultimately brown or ashy gray. Winter-buds ovoid or oval, prominently angled, light red-brown, coated with rusty brown hairs, about ¼′ long. Bark 1′—1½′ thick, and deeply divided into nearly square plates 1′—3′ long and covered by small closely appressed dark brown or nearly black scales. Wood heavy, hard, strong, dark rich brown, with thick lighter colored sapwood; largely used as fuel and in the manufacture of charcoal.

Distribution. Dry sandy or clay barrens; Long Island and Staten Island, New York, eastern and southern Pennsylvania, and southern New Jersey to the shores of Matanzas Inlet and Tampa Bay, Florida, and westward through the Gulf states to western Texas [259] (Callahan County) and to western Oklahoma (Dewey and Kiowa Counties), Arkansas, eastern Kansas, southeastern Nebraska and through Missouri to northeastern Illinois, southwestern and southern Indiana, and northeastern Kentucky (South Portsmouth, Greenup County, R. E. Horsey); rare in the north, very abundant southward; west of the Mississippi River often forming on sterile soils a great part of the forest growth; of its largest size in southern Arkansas and eastern Texas.

× Quercus Rudkinii Britt., with characters intermediate between those of Quercus marilandica and Q. Phellos, and probably a hybrid of these species, has been found near Tottenville, Staten Island, New York, at Keyport, Monmouth County, New Jersey, and at the Falls of the Yadkin River, Stanley County, North Carolina.

× Quercus sterilis Trel., believed to be a hybrid of Quercus marilandica and Q. nigra has been found in Bladen County, North Carolina.

× Quercus Hastingsii Sarg., believed to be a hybrid of Quercus marilandica and Q. texana, occurs near Boerne, Kendall County, and at Brownwood, Brown County, Texas.

× Quercus Bushii Sarg., believed to be a hybrid of Quercus marilandica and Q. velutina, although not common, occurs in eastern Oklahoma (Sapulpa, Creek County), Mississippi (Oxford, Lafayette County), Alabama (Dothan, Houston County, near Berlin, Dallas County, and Daphne, Baldwin County), Florida (Sumner, Levey County), and in Georgia (Climax, Decatur County).

14. Quercus arkansana Sarg.

Leaves broadly obovate, slightly 3-lobed or dentate at the wide apex, cuneate at base, on sterile branches often oblong-ovate, acute or rounded at apex, rounded at base, the lobes ending in long slender mucros, when they unfold tinged with red, thickly covered with pale fascicled hairs persistent until summer, the midrib and veins more thickly clothed with long straight hairs, and at maturity glabrous, with the exception of small axillary tufts of pubescence on the lower surface, light yellow-green above, paler below, 2′—2¾′ long and broad, with a slender light yellow midrib, thin primary veins and prominent veinlets; on sterile branches often 4½′—5½′ long and 2½′—2¾′ wide; petioles slender, coated at first with clusters of pale hairs, becoming glabrous or puberulous, ⅗′—⅘′ in length. Flowers: staminate in aments covered with clusters of long pale hairs, 2′—2½′ long; calyx [260] usually 4 rarely 3-lobed, thinly covered with long white hairs; stamens usually 4; anthers ovoid-oblong, apiculate, dark red; pistillate on stout peduncles, hoary-tomentose like the scales of the involucre; stigmas dark red. Fruit solitary or in pairs, on short glabrous peduncles; nut broad-ovoid, rounded at apex, sparingly pubescent especially below the middle with fascicled hairs, light brown, obscurely striate, ¼′—⅓′ long, ½′—⅝′ thick, inclosed only at base in the flat saucer-shaped cup, pubescent on the inner surface, covered with closely appressed scales obtuse at their narrow apex, red on the margins, pale pubescent, those of the upper rank smaller, erect, inserted on the top of the cup and forming a thin rim round its inner surface.

A tree when crowded in the forest often 60°—70° high, with a tall trunk, stout ascending branches forming a long narrow head, and slender branchlets thickly coated early in the season with pale fascicled hairs, pubescent or nearly glabrous in their first autumn and darker and glabrous in their second year, when not crowded by other trees rarely 40° high with a short trunk occasionally 1° in diameter. Winter-buds ovoid, acute, with thin light chestnut-brown slightly pubescent or nearly glabrous scales. Bark thick, nearly black, divided by deep fissures into long narrow ridges covered with thick closely appressed scales.

Distribution. Low woods and on rolling sand hills four miles north of Fulton, Hempstead County, Arkansas; rare and local.

15. Quercus nigra L. Water Oak.

Leaves oblong-obovate, gradually narrowed and cuneate at base and enlarged often abruptly at the broad rounded entire or occasionally 3-lobed apex, on vigorous young branchlets sometimes pinnatifid with acute, acuminate or rounded lobes or broadly oblong-obovate and rounded at apex with entire or undulate margins, on upper branches occasionally linear-lanceolate, on occasional trees narrowed below to an elongated cuneate base and gradually widened above into a more or less deeply 3-lobed apex, the lobes rounded or acute (var. tridentifera Sarg.), or often acute at the ends, and on upper branchlets sometimes linear-lanceolate to linear-obovate, acute or rounded at apex, divided above the middle by deep wide rounded sinuses into elongated lanceolate acute entire lobes, or pinnatifid above the middle, when they unfold thin, light green more or less tinged with red and covered by fine caducous pubescence, with conspicuous tufts of pale hairs in the axils of the veins below, at maturity thin, dull bluish green, paler below than above, glabrous or with axillary tufts of rusty hairs, usually about 2½′ long and 1½′ wide, or on fertile branches sometimes 6′ long and 2½′ wide; turning yellow and falling gradually during the winter; petioles stout, flattened, ⅛′—½′ in length; leaves of seedling plants linear-lanceolate with entire or undulate margins, or occasionally lobed with 1 or 2 pointed lobes, often deeply 3-lobed at a wide apex, and occasionally furnished below the middle with a single acuminate lobe, all the forms often occurring on a plant less than three feet high. Flowers: staminate in red hairy-stemmed aments 2′—3′ long; calyx thin and scarious, covered on the outer surface with short hairs, divided into 4 or 5 ovate rounded segments; pistillate on short tomentose peduncles, their involucral scales a little shorter than the acute calyx-lobes and coated with rusty hairs; stigmas deep red. Fruit usually solitary, sessile or short-stalked; nut ovoid, broad and flat at base, full and rounded at the pubescent apex, light yellow-brown, often striate, ⅓′—⅔′ long and nearly as thick, usually inclosed only at the base in a thin saucer-shaped cup, or occasionally for one third its length in a cup-shaped cup, coated on the inner surface with pale silky tomentum and covered by ovate acute closely appressed light red-brown scales clothed with pale pubescence except on their darker colored margins.

A tree, occasionally 80° high, with a trunk 2°—3½° in diameter, numerous slender branches spreading gradually from the stem and forming a symmetrical round-topped head, and slender glabrous branchlets light or dull red during their first winter, becoming grayish brown in their second season. Winter-buds ovoid, acute, strongly angled, covered by loosely imbricated dark red-brown puberulous scales slightly ciliate on the thin margins. [261] Bark ½′—¾′ thick, with a smooth light brown surface slightly tinged with red and covered by smooth closely appressed scales. Wood heavy, hard, strong, close-grained, light brown, with thick lighter colored sapwood; little valued except as fuel.

Distribution. High sandy borders of swamps and streams and the rich bottom-lands of rivers, or northward sometimes in dry woods; southern Delaware, southward to the shores of the Indian River and Tampa Bay, Florida, ranging inland in the south Atlantic states through the Piedmont region, and westward through the Gulf states to the valley of the Colorado River, Texas, and through eastern Oklahoma and Arkansas to southeastern Missouri and to central Tennessee and Kentucky. The var. tridentifera Sarg. rare and local; southwest Virginia to Alabama (near Selma, Dallas County), central and western Mississippi, eastern Louisiana; valley of Navidad River, Lavaca County, Texas. A form (f. microcarya Sarg.—Quercus microcarya Small) occurs in the dry soil on slopes of Little Stone Mountain, Dekalb County, Georgia.

The Water Oak is commonly planted as a shade-tree in the streets and squares of the cities and towns of the southern states.

16. Quercus rhombica Sarg.

Leaves rhombic, rarely oblong-obovate to lanceolate, acute or rounded and apiculate at apex, cuneate at base, the margins entire or slightly undulate, those on vigorous shoots occasionally furnished on each side near the middle with a short lobe, when they unfold deeply tinged with red, covered with short pale caducous pubescence and furnished below with usually persistent tufts of axillary hairs, at maturity thin, dark green and lustrous above, pale below, 3½′—4′ long, 1½′—2′ wide, with a stout conspicuous yellow midrib and slender forked primary veins; turning yellow and falling gradually in early winter, rarely at the ends of branches, obovate and rounded, slightly 3-lobed or undulate at the broad apex (var. obovatifolia Sarg.); petioles yellow, ⅕′—½′ in length. Flowers not seen. Fruit sessile or short-stalked; nut ovoid, rounded at apex, thickly covered with pale pubescence, ⅖′—½′ long, ⅗′ thick; inclosed only at the base in a saucer-shaped cup, rounded on the bottom, silky pubescent on the inner surface, and covered with slightly pubescent reddish brown loosely appressed scales rounded at apex, with free tips, those of the upper rank thin and ciliate on the margins.

A tree often 120°—150° high, with a tall trunk 3°—4½° in diameter, stout, wide-spreading smooth branches forming a broad open head, and slender glabrous branchlets red-brown during their first season and dark gray the following year. Bark pale gray, slightly furrowed and covered with closely appressed scales, ½′—¾′ thick.

Distribution. Borders of swamps and low wet woods of the coast region; southeastern Virginia (Dismal Swamp) to northern Florida, and through the Gulf states to the valley of the Neches River (Beaumont, Jefferson County), eastern Texas; in Louisiana northward to the valley of the Red River; most abundant in south central Alabama and in Louisiana.

× Quercus beaumontiana Sarg., believed to be a hybrid of Quercus rhombica and Q. rubra has been found growing by a street in Beaumont, Jefferson County, Texas.

× Quercus Cocksii Sarg., probably a hybrid of Quercus rhombica and Q. velutina, has been found at Pineville, Rapides Parish, Louisiana.

17. Quercus Phellos L. Willow Oak.

Leaves ovate-lanceolate or rarely obovate-lanceolate, often somewhat falcate, gradually narrowed and acute at the ends, and entire with slightly undulate margins, when they fold light yellow-green and lustrous on the upper surface, coated on the lower with pale caducous pubescence, at maturity glabrous, light green and rather lustrous above, dull and paler or rarely hoary-pubescent below, conspicuously reticulate-venulose, 2½′—5′ long, ¼′—1′ wide, with a slender yellow midrib and obscure primary veins forked and united [263] about halfway between the midrib and margins; turning pale yellow in the autumn; petioles stout, about ⅛′ in length. Flowers: staminate in slender-stemmed aments 2′—3′ long; calyx yellow, hirsute, with 4 or 5 acute segments; pistillate on slender glabrous peduncles, their involucral scales brown covered by pale hairs, about as long as the acute calyx-lobes; stigmas bright red. Fruit short-stalked or nearly sessile, solitary or in pairs; nut hemispheric, light, yellow-brown, coated with pale pubescence, inclosed only at the very base in the thin pale reddish brown saucer-shaped cup silky-pubescent on the inner surface, and covered by thin ovate hoary-pubescent closely appressed scales rounded at apex.

A tree, often 70°—90° high, with a trunk 2° or rarely 4° in diameter, small branches spreading into a comparatively narrow open or conical round-topped head, and slender glabrous reddish brown branchlets roughened by dark lenticels, becoming in their second year dark brown tinged with red or grayish brown; usually much smaller. Winter-buds ovoid, acute, about ⅛′ long, with dark chestnut-brown scales pale and scarious on the margins. Bark ½′—¾′ thick, light red-brown slightly tinged with red, generally smooth but on old trees broken by shallow narrow fissures into irregular plates covered by small closely appressed scales. Wood heavy, strong, not hard, rather coarse-grained, light brown tinged with red, with thin lighter colored sapwood; occasionally used in construction, for clapboards and the fellies of wheels.

Distribution. Low wet borders of swamps and streams and rich sandy uplands; Staten Island, New York, southern New Jersey and southeastern Pennsylvania and southward to northeastern Florida, through the Gulf states to the valley of the Navasota River, Brazos County, Texas, and through Arkansas, eastern Oklahoma and southeastern Missouri to central Tennessee and northwestern Kentucky (Ballard County), and in southwestern Illinois (Massac and Pope Counties); in the Atlantic states usually confined to the maritime plain; less common in the middle districts, rarely extending to the Appalachian foothills.

Occasionally planted as a shade-tree in the streets of southern towns, and rarely in western Europe; hardy in eastern Massachusetts.

Quercus heterophylla Michx. f.

This has usually been considered a hybrid between Quercus Phellos and Quercus velutina or Quercus borealis var. maxima; first known in the eighteenth century from an individual growing in a field belonging to John Bartram on the Schuylkill River, Philadelphia. What appears to be the same form has since been discovered in a number of stations [264] from New Jersey to Texas, and it is possible that Quercus heterophylla may, as many botanists have believed, best be considered a species.

× Quercus subfalcata Trel., believed to be a hybrid of Quercus Phellos and Q. rubra has been found at Wickliffe, Ballard County, Illinois, at Campbell, Lawrence County, Mississippi, Fulton, Hempstead County, Arkansas, and Houston, Harris County, Texas; its var. microcarpa Sarg., probably of the same parentage, originated in a Dutch nursery.

× Quercus ludoviciana Sarg., believed to be a hybrid of Quercus Phellos and Q. rubra var. pagodæfolia grows in low wet woods ten miles west of Opelousas, St. Landry Parish, Louisiana.

18. Quercus laurifolia Michx. Laurel Oak. Water Oak.

Leaves elliptic or rarely slightly broadest above the middle, acuminate at the ends, apiculate at apex, occasionally lanceolate or oblong-obovate and rounded at apex (var. hybrida Michx.) sometimes 3-lobed at apex, the terminal lobe acuminate, much larger than the others (var. tridentata Sarg.), frequently unequally lobed on vigorous branches of young trees, with small nearly triangular lobes, when they unfold in spring yellow-green, or later in the season often pink or bright red, and slightly puberulous, at maturity thin, green, and very lustrous above, light green and less lustrous below, usually 3′—4′ long and ¾′ wide, with a conspicuous yellow midrib; falling abruptly in early spring leaving the branches bare during only a few weeks; petioles stout, yellow, rarely more than ¼′ in length. Flowers: staminate in red-stemmed hairy aments 2′—3′ long; calyx pubescent on the outer surface, divided into 4 ovate rounded lobes; pistillate on stout glabrous peduncles, their involucral scales brown and hairy, about as long as the acute calyx-lobes; stigmas dark red. Fruit sessile or subsessile, generally solitary; nut ovoid to hemispheric, broad and slightly rounded at base, full and rounded at the puberulous apex, dark brown, about ½′ long, inclosed for about one fourth its length in a thin saucer-shaped cup red-brown and silky-pubescent on the inner surface, and covered by thin ovate light red-brown scales rounded at apex and pale-pubescent except on their darker colored margins.

A tree, occasionally 100° high, with a tall trunk 3°—4° in diameter, and comparatively slender branches spreading gradually into a broad dense round-topped shapely head, and slender glabrous branchlets dark red when they first appear, dark red-brown during their first winter, becoming reddish brown or dark gray in their second season. Winter-buds broadly ovoid or oval, abruptly narrowed and acute at apex, ¹/₁₆′—⅛′ long with numerous [265] thin closely imbricated bright red-brown scales ciliate on the margins. Bark of young trees ½′—1′ thick, dark brown more or less tinged with red, roughened by small closely appressed scales, becoming at the base of old trees 1′—2′ thick, nearly black, and divided by deep fissures into broad flat ridges. Wood heavy, very strong and hard, coarse-grained, liable to check badly in drying, dark brown tinged with red, with thick lighter colored sapwood; probably used only as fuel.

Distribution. Sandy banks of streams and swamps and rich hummocks in the neighborhood of the coast; North Carolina (near Newbern) southward to the shores of Bay Biscayne and the valley of the Caloosahatchie River, Florida, and in the interior of the peninsula to the neighborhood of Lake Istokpaga, De Soto County, and westward to eastern Louisiana, ranging inland to Darlington, Darlington County, South Carolina, to the neighborhood of Augusta, Richmond County, Mayfield, Hancock County, Albany, Dougherty County, Cuthbert, Randolph County, and Bainbridge, Decatur County, Georgia, Georgiana, Butler County, and Berlin, Dallas County, Alabama, Rockport, Copiah County, Mississippi, and to the neighborhood of Bogalusa, Washington Parish, Louisiana (R. S. Cocks); nowhere abundant, but most common and of its largest size in eastern Florida.

19. Quercus cinerea Michx. Blue Jack. Upland Willow Oak.

Leaves oblong-lanceolate to oblong-obovate, gradually narrowed and cuneate or sometimes rounded at base, acute or rounded and apiculate at apex, entire with slightly thickened undulate margins, or at the ends of vigorous sterile branches occasionally 3-lobed at the apex and variously lobed on the margins (β dentato-lobata A. De Candolle), when they unfold bright pink and pubescent on the upper surface, coated on the lower with thick silvery white tomentum, at maturity firm in texture, blue-green, lustrous, conspicuously reticulate venulose above, pale-tomentose below, 2′—5′ long, ½′—1½′ wide, with a stout yellow midrib and remote obscure primary veins forked and united within the margins; turning red and falling gradually late in the autumn or in early winter; petioles stout, ¼′—½′ in length. Flowers: staminate in hoary-tomentose aments 2′—3′ long; calyx pubescent, bright red and furnished at apex with a thick tuft of silvery white hairs before opening, divided into 4 or 5 ovate acute lobes, becoming yellow as it opens; stamens 4 or 5; anthers apiculate, dark red in the bud, becoming yellow; pistillate on short stout tomentose peduncles, their involucral scales about as long as the acute calyx-lobes and coated with [266] pale tomentum; stigmas dark red. Fruit produced in great profusion, sessile or raised on a short stalk rarely ¼′ long; nut ovoid, full and rounded at the ends or subglobose, about ½′ long, often striate, hoary-pubescent at apex, inclosed only at the base or for one half its length in a thin saucer-shaped or cup-shaped cup bright red-brown and coated with lustrous pale pubescence on the inner surface, and covered by thin closely imbricated ovate-oblong scales hoary-tomentose except on the dark red-brown margins.

A tree on dry hills, usually 15°—20° high, with a trunk 5′—6′ in diameter, stout branches forming a narrow irregular-head, and thick rigid branchlets coated at first with a dense fulvous or hoary tomentum of fascicled hairs, soon becoming glabrous or puberulous, dark brown sometimes tinged with red during their first winter and darker in their second year; or in low moist soil often 60°—75° high, with a trunk 18′—20′ in diameter, and a broad round-topped shapely head of drooping branches. Winter-buds ovoid, acute, with numerous rather loosely imbricated bright chestnut-brown scales ciliate on the margins, often ¼′ long on vigorous branches, frequently obtuse and occasionally much smaller. Bark ¾′—1½′ thick, and divided into thick nearly square plates 1′—2′ long, and covered by small dark brown or nearly black scales slightly tinged with red. Wood hard, strong, close-grained, light brown tinged with red, with thick darker colored sapwood; probably only used as fuel.

Distribution. Sandy barrens and dry upland ridges, and in the rich moist soil of the pine-covered flats of the Florida peninsula; North Carolina southward to the shores of the Indian River and Peace Creek, Florida, and along the Gulf coast to the valley of the Brazos River, Texas; in the Atlantic and middle Gulf states mostly confined to a maritime belt 40°—60° wide, extending across the Florida peninsula as far south as the sand hills in the neighborhood of Lake Istokpoga, De Soto County, and west of the Mississippi River, ranging inland to the neighborhood of Dallas, Dallas County, Texas and to southeastern Oklahoma (near Antlers, Pushmataha County).

× Quercus dubia Ashe, believed to be a hybrid of Quercus cinerea and Q. laurifolia occurs at Abbottsburg, Bladen County, North Carolina, on the coast of South Carolina, in southern Georgia and northern and central Florida, and at Mississippi City, Lincoln County, Mississippi.

× Quercus subintegra Trel., a supposed hybrid of Quercus cinerea and Q. rubra occurs at Lumber City, Telfair County, Georgia, Lake City, Columbia County, Florida, and at Berlin, Dallas County, Alabama.

× Quercus sublaurifolia Trel., a supposed hybrid of Quercus cinerea and Q. laurifolia occurs at Folkston, Charlton County, Georgia, and at Biloxi, Harrison County, Mississippi.

× Quercus carolinensis Trel., believed to be a hybrid of Quercus cinerea and Q. marilandica occurs at Newbern, Craven County, North Carolina, Lumber City, Telfair County and Climax, Decatur County, Georgia, and near Fletcher, Hardin County, Texas.

× Quercus caduca Trel., believed to be a hybrid of Quercus cinerea and Q. nigra, occurs at Folkston, Charlton County and Lumber City, Telfair County, Georgia, Jacksonville, Duval County, and Gainsville, Alachua County, Florida, Mississippi City, Harrison County, Mississippi, and at Milano, Milano County and Bryan, Brazos County, Texas.

× Quercus oviedoensis Sarg., believed to be a hybrid of Quercus cinerea and Q. myrtifolia, has been found near Oviedo, Orange County, Florida.

20. Quercus imbricaria Michx. Shingle Oak. Laurel Oak.

Leaves oblong-lanceolate to oblong-obovate, apiculate and acute or rounded at apex, gradually narrowed and cuneate or rounded at base, entire with slightly thickened, revolute often undulate margins, or sometimes more or less 3-lobed, or on sterile branches occasionally repand-lobulate, when they unfold bright red, soon becoming yellow-green, covered with scurfy rusty pubescence on the upper surface and hoary-tomentose on the lower, at maturity thin, glabrous, dark green, and very lustrous above, pale green or light brown and pubescent below, 4′—6′ long, ¾′—2′ wide, with a stout yellow midrib, numerous slender yellow veins arcuate and united at some distance from the margins, and reticulate [267] veinlets; late in the autumn turning dark red on the upper surface; petioles stout, pubescent, rarely more than ½′ in length. Flowers: staminate in hoary-tomentose aments, 2′—3′ long; calyx light yellow, pubescent, and divided into 4 acute segments; pistillate on slender tomentose peduncles, their involucral scales covered with pale pubescence and about as long as the acute calyx-lobes; stigmas greenish yellow. Fruit solitary or in pairs, on stout peduncles often nearly ½′ in length; nut nearly as broad as long, full and rounded at the ends, dark chestnut-brown, often obscurely striate, ½′—⅔′ long, inclosed for one third to one half its length in a thin cup-shaped or turbinate cup bright red-brown and lustrous on the inner surface, and covered by thin ovate light red-brown scales rounded or acute at the apex and pubescent except on their darker colored margins.

A tree, usually 50°—60° high, with a trunk rarely exceeding 3° in diameter, or rarely 100° high, with a long naked stem 3°—4° in diameter, slender tough horizontal or somewhat pendulous branches forming a narrow round-topped picturesque head, and slender branchlets dark green, lustrous, and often suffused with red when they first appear, soon glabrous, light reddish brown or light brown during their first winter and dark brown in their second year. Winter-buds ovoid, acute, about ⅛′ long, obscurely angled, and covered by closely imbricated light chestnut-brown lustrous scales erose and often ciliate on the margins. Bark on young stems and branches thin, light brown, smooth, and lustrous, becoming on old trunks ¾′—1½′ thick, and slightly divided by irregular shallow fissures into broad ridges covered by close slightly appressed light brown scales somewhat tinged with red. Wood heavy, hard, rather coarse-grained, light brown tinged with red, with thin lighter colored sapwood; occasionally used in construction, and for clapboards and shingles.

Distribution. Rich hillsides and the fertile bottom-lands of streams; Lehigh County (Allentown to Dorney’s Park), Bedford, Huntington, Franklin and Union Counties, Pennsylvania, westward through Ohio to southern Michigan, southern Wisconsin and southeastern and southern Iowa (Muscatine to Taylor County), and southward to the District of Columbia, along the Appalachian Mountains and their foothills, up to altitudes of 2200°, to the valley of the Little Tennessee River, North Carolina, and to northern Georgia (Wilkes County), and middle Tennessee; through Missouri to northeastern Kansas and southeastern Nebraska, and in northern and southern Arkansas (Fulton, Hempstead County); comparatively rare in the east; one of the most abundant Oaks of the lower Ohio basin; probably growing to its largest size in southern Indiana and Illinois.

Occasionally planted as an ornamental tree in the northern states, and hardy as far north as Massachusetts.

× Quercus Leana Nutt., scattered usually in solitary individuals from the District of Columbia and western North Carolina to southern Michigan, central and northern Illinois and southeastern Missouri, is believed to be a hybrid between this species and Quercus velutina.

× Quercus tridentata Engelm., described as a hybrid of Quercus imbricaria and Q. marilandica first found at Allenton, Saint Louis County, Missouri, occurs also near Olney, Richland County, Illinois.

× Quercus exacta Trel., believed to be a hybrid of Quercus imbricaria and Q. palustris, occurs near Olney, Richland County, Illinois, and at Crown Point, Lake County, Indiana.

21. Quercus hypoleuca Engelm.

Leaves lanceolate or oblong-lanceolate to elliptic, occasionally somewhat falcate, acute and often apiculate at apex, cuneate or rounded or cordate at the narrow base, entire or repandly serrate above the middle with occasionally small minute rigid spinose teeth, or on vigorous shoots serrate-lobed with oblique acute lobes, when they unfold light red, covered with close pale pubescence above and coated below with thick hoary tomentum, at maturity thick and firm, dark yellow-green and lustrous on the upper surface, covered on the lower with thick silvery white or fulvous tomentum, 2′—4′ long, ½′—1′ wide, with thickened revolute margins; turning yellow or brown and falling gradually during the spring after the appearance of the new leaves; petioles stout, flattened, pubescent or tomentose, ⅛′—¼′ in length. Flowers: staminate in slender aments 4′—5′ long; calyx slightly tinged with red, covered with pale hairs and divided into 4 or 5 broadly ovate rounded lobes; anthers acute, apiculate, bright red becoming yellow; pistillate mostly solitary, sessile or short-stalked, their involucral scales thin, scarious, and soft-pubescent; stigmas dark red. Fruit sessile or borne on a stout peduncle up to ½′ in length, usually solitary; nut ovoid, acute or rounded at the narrow hoary-pubescent apex, dark green and often striate when ripe, becoming light chestnut-brown in drying, ½′—⅔′ long, the shell lined with white tomentum, inclosed for about one third its length in a turbinate thick cup pubescent on the inner surface, and covered by thin broadly ovate light chestnut-brown scales rounded at apex and clothed, especially toward the base of the cup, with soft silvery pubescence.

A tree, usually 20°—30° or sometimes 60° high, with a tall trunk 10′—15′ in diameter, slender branches spreading into a narrow round-topped inversely conic head, and stout rigid branchlets coated at first with thick hoary tomentum disappearing during the first winter, becoming light red-brown often covered with a glaucous bloom and ultimately nearly black; frequently a shrub. Winter-buds ovoid, obtuse, about ⅛′ long, with thin [269] light chestnut-brown scales. Bark ¾′—1′ thick, nearly black, deeply divided into broad ridges broken on the surface into thick plate-like scales. Wood heavy, very strong, hard, close-grained, dark brown, with thick lighter colored sapwood.

Distribution. Scattered but nowhere abundant through Pine-forests on the slopes of cañons and on high ridges usually at altitudes between 6000°—7000° above the sea on the mountains of western Texas, and of southern New Mexico and Arizona; in northern Chihuahua and Sonora.

22. Quercus agrifolia Née. Live Oak. Encina.

Leaves oval, orbicular or oblong, rounded or acute and apiculate at apex, rounded or cordate at base, entire or sinuate-dentate with slender rigid spinose teeth, when they unfold tinged with red and coated with caducous hoary tomentum, at maturity subcoriaceous, convex, dark or pale green, dull and obscurely reticulate above, paler, rather lustrous, glabrous or pubescent below, with tufts of rusty hairs in the axils of the principal veins, or sometimes covered above with fascicled hairs and coated below with thick hoary pubescence, ¾′—4′ long and ½′—3′ wide, with thickened strongly revolute margins; falling gradually during the winter and early spring; petioles stout or slender, pubescent or glabrous, ½′—1′ in length. Flowers: staminate in slender hairy aments 3′—4′ long; calyx bright purple-red in the bud, sometimes furnished with a tuft of long pale hairs at the apex, glabrous or glabrate, divided nearly to the base into 5—7 ovate acute segments reddish above the middle; pistillate sessile or short-stalked, their involucral scales bright red and covered with thick hoary tomentum, or glabrous or puberulous; stigmas bright red. Fruit sessile or nearly so, solitary or in few-fruited clusters; nut elongated, ovate, abruptly narrowed at base, gradually narrowed to the acute puberulous apex, light chestnut-brown, ¾′—1½′ long, ¼′—¾′ thick, the shell lined with a thick coat of pale tomentum, inclosed for one third its length or only at the base in a thin turbinate light brown cup coated on the inner surface with soft pale silky pubescence, and covered by thin papery scales rounded at the narrow apex, and slightly puberulous, especially toward the base of the cup.

A tree, occasionally 80°—90° high, with a short trunk 3°—4° or rarely 6°—7° in diameter, dividing a few feet above the base into numerous great limbs often resting on the ground and forming a low round-topped head frequently 150° across, and slender dark gray or brown branchlets tinged with red, coated at first with hoary tomentum persistent until the second or third year; or with a trunk, rising to the height of 30° or 40°, and crowned by a narrow head of small branches; often much smaller; frequently shrubby in habit, [270] with slender stems only a few feet high. Winter-buds globose and usually about ¹/₁₆′ thick, or ovoid-oblong, acute, and sometimes on vigorous shoots nearly ¼′ in length, with thin broadly ovate closely imbricated light chestnut-brown glabrous or pubescent scales. Bark of young stems and branches thin, close, light brown or pale bluish gray, becoming on old trunks 2′—3′ thick, dark brown slightly tinged with red, and divided into broad rounded ridges separating on the surface into small closely appressed scales. Wood heavy, hard, close-grained, very brittle, light brown or reddish brown, with thick darker colored sapwood; valued and largely used for fuel.

Distribution. Usually in open groves of great extent from Sonoma County, California, southward over the coast ranges and islands to the San Pedro Mártir Mountains, Lower California; less common at the north; very abundant and of its largest size in the valleys south of San Francisco Bay and their commonest and characteristic tree; frequently covering with semiprostrate and contorted stems the sand dunes on the coast in the central part of the state; in southwestern California the largest and most generally distributed Oak-tree between the mountains and the sea, often covering low hills and ascending to altitudes of 4500° in the cañons of the San Jacinto Mountains.

Occasionally cultivated as an ornamental tree in temperate western, and in southern Europe.

23. Quercus Wislizenii A. DC. Live Oak.

Leaves narrowly lanceolate to broadly elliptic, generally oblong-lanceolate, acute or rounded and generally apiculate at apex, rounded or truncate or gradually narrowed and cuneate at base, entire, serrulate or serrate or sinuate-dentate with spreading rigid spinescent teeth, when they unfold thin, dark red, ciliate, and covered with pale scattered fascicled hairs, at maturity thick and coriaceous, glabrous and lustrous, dark green on the upper and paler and yellow-green on the lower surface, usually 1′—1½′ long and about ⅔′ wide, with obscure primary veins and conspicuous reticulate veinlets, gradually deciduous during their second summer and autumn; petioles coated at first with hoary tomentum, usually pubescent or puberulous at maturity, ⅛′ to nearly 1′ in length. Flowers: staminate in hairy aments 3′—4′ long; calyx tinged with red in the bud, divided into broadly ovate ciliate glabrous light yellow lobes shorter than the 3—6 stamens; pistillate sessile or short-stalked, their involucral scales and peduncle hoary-tomentose. Fruit sessile, short-stalked or occasionally spicate; nut slender, oblong, abruptly narrowed at base, pointed and pilose at the apex, ¾′—1½′ long, about ⅓′ thick, light chestnut-brown, often striate, the shell lined with a scanty coat of pale tomentum, more or less inclosed in the thin turbinate sometimes [271] tubular cup ½′—1′ deep, or rarely cup-shaped and shallow, light green and puberulous within, and covered by oblong lanceolate light brown closely imbricated thin scales, sometimes toward the base of the cup thickened and rounded on the back, usually pubescent or puberulous, especially above the middle, and frequently ciliate on the margins.

A tree, usually 70°—80° high, with a short trunk 4°—6° in diameter, stout spreading branches forming a round-topped head, and slender rigid branchlets coated at first with hoary tomentum or covered with scattered fascicled hairs, puberulous or glabrous and rather light brown during their first season, gradually growing darker in their second year; usually much smaller and sometimes reduced to an intricately branched shrub, with numerous stems only a few feet tall. Winter-buds ovoid or oval, acute, ⅛′—¼′ long, with closely imbricated light chestnut-brown ciliate scales. Bark on young trees and large branches thin, generally smooth and light-colored, becoming on old trunks 2′—3′ thick, and divided into broad rounded often connected ridges separating on the surface into small thick closely appressed dark brown scales slightly tinged with red. Wood heavy, very hard, strong, close-grained, light brown tinged with red, with thick lighter colored sapwood; sometimes used for fuel.

Distribution. Lower slopes of Mt. Shasta southward through the coast region of California to the Santa Lucia Mountains, and to Santa Rosa and Santa Cruz Islands, and along the slopes of the Sierra Nevada to Kern County, up to altitudes of 2000° at the north and of 4500° at the south; as a shrub 4°—6° high with small thick leaves (var. frutescens Engelm.) on the desert slopes of the San Bernardino, San Jacinto and Cuyamaca mountains, at altitudes of 5000°—7000° above the sea, and on San Pedro Mártir in Lower California; nowhere common as a tree, but most abundant and of its largest size in the valleys of the coast region of central California at some distance from the sea, and on the foothills of the Sierra Nevada; very common as a shrub in the cañons of the desert slopes of the mountains of southern California; near the coast and on the islands small and mostly shrubby.

× Quercus morehus Kell., a supposed hybrid between Quercus Wislizenii and Q. Kelloggii occurs in Lake County, California.

24. Quercus myrtifolia Willd.

Leaves oval to oblong-obovate, acute and apiculate or broad and rounded at apex, gradually narrowed and cuneate or broad and rounded or cordate at base, entire, with much thickened revolute sometimes undulate margins, or on vigorous shoots sinuate-dentate and lobed above the middle, when they unfold, thin, dark red, coated below and on the [272] petioles with clammy rusty tomentum and densely pubescent above, at maturity thick and coriaceous, lustrous, dark green, glabrous and conspicuously reticulate-venulose above, paler, yellow-green, or light orange-brown, glabrous or pubescent below, with tufts of rusty hairs in the axils of the veins, ½′—2′ long and ¼′—1′ wide; falling gradually during their second year; petioles stout, pubescent, yellow, rarely more than ⅛′ in length. Flowers: staminate in hoary pubescent aments 1′—1½′ long; calyx coated on the outer surface with rusty hairs and divided into 5 ovate-acute, segments shorter than the 2 or 3 stamens; pistillate sessile or nearly sessile, solitary or in pairs, their involucral scales tomentose and tinged with red. Fruit solitary or in pairs, sessile or short-stalked; nut subglobose or ovoid, acute, ¼′—½′ long, dark brown, lustrous and often striate, puberulous at apex, the shell lined with a thick coat of rusty tomentum, inclosed for one fourth to one third its length in a saucer-shaped or turbinate cup light brown and puberulous within, and covered by closely imbricated broad-ovate light brown pubescent scales ciliate on the margins and rounded at their broad apex.

A round-topped tree, rarely 40° high, with a trunk 4′—5′ or rarely up to 15′ in diameter, short or rarely long spreading branches and slender branchlets coated at first with a thick pale fulvous tomentum of articulate hairs usually persistent during the summer, light brown more or less tinged with red or dark gray, and pubescent or puberulous during their first winter, becoming darker and glabrous in their second season; more often an intricately branched shrub, with slender rigid stems 3°—4° or rarely 15°—20° high and 1′—3′ in diameter. Winter-buds ovoid or oval, gradually narrowed to the acute apex, with closely imbricated dark chestnut-brown slightly puberulous scales. Bark thin and smooth, becoming near the ground dark and slightly furrowed.

Distribution. Dry sandy ridges on the coast and islands of South Carolina to Bay Biscayne, Florida, crossing the central peninsula and from the valley of the Caloosahatchee River, westward along the coast of Florida, Alabama, and Mississippi; most abundant on the islands off the coast of east Florida, and of Alabama and Mississippi; often covering large areas with low impenetrable thickets; perhaps of its largest size in Orange County, on Jupiter Island, and on the coast west of the Apalachicola River, Florida.

25. Quercus chrysolepis Liebm. Live Oak. Maul Oak.

Leaves oblong-ovate to elliptic, acute or cuspidate at apex, cordate, rounded or cuneate at base, mostly entire on old trees, often dentate or sinuate-dentate on young trees with 1 or 2 or many spinescent teeth, the two forms often appearing together on vigorous shoots, clothed when they unfold with a thick tomentum of fulvous hairs soon deciduous from the [273] upper and more gradually from the lower surface, at maturity thick and coriaceous, bright yellow-green and glabrous above, more or less fulvous-tomentose below during their first year, ultimately becoming glabrate and bluish white, 1′—4′ long, ½′—2′ wide, with thickened revolute margins; deciduous during their third and fourth years; petioles slender, yellow, rarely ½′ in length. Flowers: staminate in slender tomentose aments 2′—4′ long; calyx light yellow, pubescent, divided usually into 5—7 broadly ovate acute ciliate lobes often tinged with red above the middle; pistillate sessile or subsessile or rarely in short few-flowered spikes, their broadly ovate involucral scales coated with fulvous tomentum; stigmas bright red. Fruit usually solitary, sessile or short-stalked; nut ellipsoidal or ovoid, acute or rounded at the full or narrow slightly puberulous apex, light chestnut-brown, ½′—2′ long and about as thick, the shell lined with a thin coat of loose tomentum, with abortive ovules scattered irregularly over the side of the seed, inclosed only at the base in a thin hemispheric or in a thick turbinate broad-rimmed cup pale green or dark reddish brown within, and covered by small triangular closely appressed scales with a short free tip, clothed with hoary pubescence, or often hidden in a dense coat of fulvous tomentum.

A tree, usually not more than 40°—50° high, with a short trunk 3°—5° in diameter, dividing into great horizontal limbs sometimes forming a head 150° across, and slender rigid or flexible branchlets coated at first with thick fulvous tomentum, becoming during their first winter dark brown somewhat tinged with red, tomentose, pubescent, or glabrous, and ultimately light brown or ashy gray; occasionally in sheltered cañons producing trunks 8°—9° in diameter; on exposed mountain sides forming dense thickets 15°—20° high. Winter-buds broadly ovoid or oval, acute, about ⅛′ long, with closely imbricated light chestnut-brown usually puberulous scales. Bark ¾′—1½′ thick, light or dark gray-brown tinged with red, and covered by small closely appressed scales. Wood heavy, very strong, hard, tough, close-grained, light brown, with thick darker colored sapwood; used in the manufacture of agricultural implements and wagons.

Distribution. Southern Oregon, along the California coast ranges and the western slopes of the Sierra Nevada to the San Bernardino and San Jacinto mountains; of its largest size in the cañons of the coast ranges of central California and on the foothills of the Sierra Nevada; ascending to altitudes of 8000°—9000° above the sea; near the southern boundary of California, on the mountains of northern Lower California and Sonora and in Arizona (Santa Rita and Huachuca Mountains, on Beaver Creek and in Copper Cañon near Camp Verde, and in Sycamore Cañon south of Flagstaff), usually shrubby, with rigid branches, rigid coriaceous oblong or semiorbicular spinose-dentate leaves, subsessile or pedunculate fruit, with ovoid acute nuts 1′—1½′ long, their shells lined with thick or thin pale tomentum, and purple cotyledons (var. Palmeri Engelm.—Quercus Wilcoxii Rydb.)

26. Quercus tomentella Engelm.

Leaves oblong-lanceolate, acute, sometimes cuspidate or occasionally rounded at apex, broad and rounded or gradually narrowed and abruptly cuneate at base, remotely crenate-dentate with small remote spreading callous tipped teeth, or entire, when they unfold light green tinged with red, covered above with scattered pale fascicled hairs and below and on the petioles with thick hoary tomentum, at maturity thick and coriaceous, dark green, glabrous and lustrous on the upper surface, pale and covered with fascicled hairs on the lower surface, 2′—4′ long, 1′—2′ wide, with thickened strongly revolute margins, and a pubescent midrib; gradually deciduous during their third season; petioles stout, pubescent, about ½′ in length. Flowers: staminate in pubescent aments 2½′—14′ long, calyx light yellow, pubescent, divided into 5—7 ovate acute lobes; pistillate subsessile or in few-flowered spikes on short or elongated pubescent peduncles, their involucral scales like the calyx coated with fascicled hairs; stigmas red. Fruit subsessile or short-stalked; nut ovoid, broad at base, full and rounded at apex, about 1½′ long and ¾′ thick, inclosed only at the base in a cup-shaped shallow cup thickened below, light brown and pubescent on the inner surface, and covered by thin ovate acute scales, their free chestnut-brown tips more or less hidden in a thick coat of hoary tomentum.

A tree, 30°—40°, or occasionally 60° high, with a trunk 1°—2° in diameter, spreading branches forming a shapely round-topped head, and slender branchlets coated at first with hoary tomentum, becoming light brown tinged with red or orange color. Winter-buds ovoid, acute or obtuse, nearly ¼′ long, with many loosely imbricated light chestnut-brown scales more or less clothed with pale pubescence. Bark thin, reddish brown, broken into large closely appressed scales. Wood heavy, hard, close-grained, compact, pale yellow-brown, with lighter colored sapwood.

Distribution. Deep narrow cañons and high wind-swept slopes of Santa Rosa, Santa Cruz, and Santa Catalina islands, California; on Guadalupe Island off the coast of Lower California.

27. Quercus Emoryi Torr. Black Oak.

Leaves oblong-lanceolate, acute and mucronate at apex, cordate or rounded at the slightly narrowed base, entire or remotely repand-serrate with 1—5 pairs of acute rigid oblique teeth, when they unfold thin, light green more or less tinged with red and covered with silvery white tomentum, at maturity thick, rigid, coriaceous, dark green, very lustrous and glabrous or coated above with minute fascicled hairs, pale and glabrous or puberulous below, usually with 2 large tufts of white hairs at the base of the slender midrib, obscurely reticulate-venulose, 1′—2½′ long, ½′—1′ wide; falling gradually in April with the appearance of the new leaves; petioles stout, pubescent, about ¼′ in length. Flowers: staminate in hoary tomentose aments; calyx light yellow, hairy on the outer surface, divided into 5—7 ovate acute lobes; pistillate sessile or short-stalked, their involucral scales covered with hoary tomentum. Fruit ripening irregularly from June to September, sessile or short-stalked; nut oblong, oval, or ovate, narrowed at base, rounded at the narrow pilose apex, ½′—¾′ long, about ⅓′ thick, dull light green when fully grown, dark chestnut-brown or nearly black at maturity, with a thin shell lined with thick white tomentum, inclosed for from one third to one half its length in the deeply cup-shaped or nearly hemispheric cup light green and pubescent within, and covered by closely imbricated broadly ovate acute thin and scarious light brown scales clothed with short soft pale pubescence.

A tree, usually 30°—40° high, with a short trunk 2°—3° in diameter, stout rigid rather drooping branches forming a round-topped symmetrical head, and slender rigid branchlets covered at first with close hoary tomentum, bright red, pubescent or tomentose in their first winter, ultimately glabrous and dark red-brown or black; sometimes 60°—70° high, with a trunk 4°—5° in diameter, with a head occasionally 100° across; or at high altitudes [275] or on exposed mountain slopes a low shrub. Winter-buds ellipsoidal, acute, about ¼′ long, pale pubescent toward the apex, with thin closely imbricated light chestnut-brown ciliate scales. Bark 1′—2′ thick, dark brown or nearly black, deeply divided into large oblong thick plates separating into small thin closely appressed scales. Wood heavy, strong, brittle, close-grained, dark brown or almost black, with thick bright brown sapwood tinged with red. The sweet acorns are an important article of food for Mexicans and Indians, and are sold in the towns of southern Arizona and northern Mexico.

Distribution. Mountain ranges of western Texas, southern New Mexico, Arizona south of the Colorado plateau, and of northern Mexico; in Texas common in the cañons and on the southern slopes of the Limpio and Chisos mountains; the most abundant Oak of southern New Mexico and Arizona, forming a large part of the forests covering the mountain slopes and extending from the upper limits of the mesa nearly to the highest ridges; attaining its largest size and beauty in the moist soil of sheltered cañons.

28. Quercus dumosa Nutt. Scrub Oak.

Leaves oblong, rounded and acute at apex, broad and abruptly cuneate or rounded at base, usually about ¾′ long and ½′ wide, spinescent with a few minute teeth, or undulate and entire or coarsely spinescent, with an obscure midrib and primary veins, conspicuous reticulate veinlets, and stout petioles rarely ⅛′ long; or sometimes oblong to oblong-obovate and divided by deep sinuses into 5—9 oblong acute rounded or emarginate bristle-tipped lobes, the terminal lobe 3-lobed, rounded or acute, 2′—4′ long and 1′—1½′ wide, with primary veins running to the points of the lobes, obscure reticulate veinlets, and petioles sometimes 1′ long, thin when they unfold and clothed with scattered fascicled hairs, or rarely tomentose above and coated below and on the petioles with hoary tomentum, at maturity thick and firm, dark green and glabrous on the upper surface, paler and more or less pubescent on the lower surface; mostly deciduous during the winter. Flowers: staminate in pubescent aments; calyx divided into 4—7 ovate lanceolate hairy segments; pistillate sessile or stalked, in long many-flowered tomentose spikes, their involucral scales and calyx hoary-tomentose; stigmas red. Fruit sessile or short-stalked; nut ovoid, broad at base, broad and rounded or acute at apex, ½′—1′ long, ⅓′—⅔′ thick, inclosed for one half to two thirds its length in a deep cup-shaped or hemispheric cup light brown and pubescent within, covered by ovate pointed scales coated with pale or rufous tomentum, usually much thickened, united and tuberculate, those above with free acute tips forming a fringe to the rim of the cup, or frequently with basal scales but little thickened and furnished with long free tips; in var. Alvordiana Jeps., with a nut 1½′—1⅝′ long, ¼′—½′ thick, gradually narrowed and acute at apex, inclosed only at base in a shallow cup-shaped cup.

A tree, rarely 20° high, with a trunk 12′—18′ in diameter, small branches forming a round-topped head, and slender branchlets coated at first with hoary tomentum, becoming in their first winter ashy gray or light or dark reddish brown and usually pubescent or tomentose; more often an intricately branched rigid shrub, with stout stems covered by pale gray bark and usually 6°—8° high, often forming dense thickets. Winter-buds ellipsoidal, generally acute, ¹/₁₆′—⅛′ long, with thin pale red often pilose and ciliate scales. Bark of the trunk bright brown and scaly.

Distribution. California; western slopes of the central Sierra Nevada; common on the coast ranges south of San Francisco Bay and on the islands off the coast of the southern part of the state, ranging inland to the borders of the Mohave Desert and to the cañons of the desert slopes of the San Bernardino and San Jacinto mountains, and southward into Lower California; arborescent only in sheltered cañons of the islands; the var. Alvordiana, in the San Emidio Cañon of the coast ranges of Kern County and on the San Carlos Range, Fresno County; north of San Francisco Bay replaced by the variety bullata Engelm. ranging to Mendocino County and to Napa valley.

× Quercus MacDonaldii Greene, a shrub or small tree with characters intermediate between those of Quercus dumosa and Q. Engelmannii, is usually considered a hybrid of these species. It occurs on Santa Cruz and Santa Catalina Islands, and in Santa Barbara, and Los Angeles Counties, California.

29. Quercus virginiana Mill. Live Oak.

Leaves oblong, elliptic or obovate, rounded or acute at apex, gradually narrowed and cuneate or rarely rounded or cordate at base, usually entire with slightly revolute margins, or rarely spinose-dentate above the middle, thin, dark green and lustrous on the upper surface, pale and pubescent on the lower surface, 2′—5′ long, ½′—2½′ wide, and inconspicuously reticulate-venulose, with a narrow yellow midrib, and few slender obscure primary veins forked and united at some distance from the margins; gradually turning yellow or brown at the end of the winter and falling with the appearance of the new leaves in the spring; petioles stout, rarely more than ¼′ in length. Flowers: staminate in hairy aments 2′—3′ long; calyx light yellow, hairy, divided into 5—7 ovate rounded segments; anthers hirsute; pistillate in spikes on slender pubescent peduncles 1′—3′ long, their involucral scales and ovate calyx-lobes coated with hoary pubescence; stigmas bright red. Fruit usually in 3—5 fruited spikes or rarely in pairs or single on stout light brown puberulous peduncles 1′—5′ long; nut ellipsoidal or slightly obovoid, narrowed at base, rounded or acute at apex, dark chestnut-brown and lustrous, about 1′ long and ⅓′ thick, inclosed for about one fourth its length in a turbinate light reddish brown cup puberulous within, [277] its scales thin, ovate, acute, slightly keeled on the back, covered by dense lustrous hoary tomentum and ending in small closely appressed reddish tips; seed sweet, with light yellow connate cotyledons.

A tree, 40°—50° high, with a trunk 3°—4° in diameter above its swollen buttressed base, usually dividing a few feet from the ground into 3 or 4 horizontal wide-spreading limbs forming a low dense round-topped head sometimes 130° across, and slender rigid branchlets coated at first with hoary tomentum, becoming ashy gray or light brown and pubescent or puberulous during their first winter and darker and glabrous the following season; occasionally 60°—70° tall, with a trunk 6°—7° in diameter; often shrubby and occasionally not more than a foot high. Winter-buds globose or slightly obovoid, about ⅙′ long, with thin light chestnut-brown scales white and scarious on the margins. Bark of the trunk and large branches ½′—1′ thick, dark brown tinged with red, slightly furrowed, separating on the surface into small closely appressed scales. Wood very heavy, hard, strong, tough, close-grained, light brown or yellow, with thin nearly white sapwood; formerly largely and still occasionally used in shipbuilding.

Distribution. Shores of Mobjack Bay, Virginia, southward along the coast and islands to southern Florida, and along the shores of the Gulf of Mexico to northeastern Mexico, spreading inland through Texas to the valley of the Red River and to the mountains in the extreme western part of the state; on the mountains of Cuba, southern Mexico, and Central America; most abundant and of its largest size on the Atlantic and east Gulf coasts on rich hummocks and ridges a few feet above the level of the sea; abundant in Texas in the coast region, near the banks of streams, and westward toward the valley of the Rio Grande often forming the principal part of the shrubby growth on low moist soil; in sandy barren soil in the immediate vicinity of the seacoast or on the shores of salt water estuaries and bays often a shrub, sometimes bearing fruit on stems not more than a foot high (var. maritima, Sarg., and var. dentata Sarg.).

Occasionally planted as a shade and ornamental tree in the southern United States.

Variable in habit and in the size and thickness of the leaves the different forms of Quercus virginiana show little variation in their fruit. The most important of these varieties is

Quercus virginiana var. geminata Sarg.

Leaves oblong-obovate to elliptic, rounded or acute at apex, cuneate or narrowed and rounded at base, occasionally slightly and irregularly dentate above the middle on vigorous shoots, conspicuously reticulate-venulose, hoary tomentose below, 1½′—3′ long, ⅓′—1′ [278] wide, with thickened strongly revolute margins; persistent until after the leaves of the typical Q. virginiana in the same locality have all fallen; occasionally in Florida with oblong-elliptic to slightly obovate leaves 4½′—5′ long and 1′—2′ wide (f. grandifolia Sarg.). Flowers and Fruit as in the species.

A tree often 75° high with a trunk 3° in diameter, with the habit, branchlets, winter-buds and bark of the typical form; often much smaller and occasionally a shrub.

Distribution. Sandy soil; coast region of North Carolina south of the Cape Fear River, South Carolina and Georgia, and southward in Florida to Jupiter Island on the east coast and the valley of the Caloosahatchee River on the west coast; abundant and often the common Live Oak in the central part of the peninsula, at least as far south as Orange County, and westward through western Florida, southeastern and southern Alabama to the Gulf coast and islands of Mississippi.

Other varieties of Quercus virginiana are var. macrophylla Sarg., differing from the type in its much larger ovate or slightly obovate leaves rounded or acute at base, entire or occasionally repand-dentate, pale tomentose below, 3½′—4′ long and 1¼′—2½′ wide. Large trees forming groves; sandy bottoms of the Atascosa River and in flat woods above them, Pleasanton, Atascosa County, Texas: var. virescens Sarg., differing from the type in the green glabrous or rarely puberulous lower surface of the leaves and in the glabrous branchlets. A large tree in sandy soil; Gainesville, Alachua County, Sanford, Seminole County, Sumner, Levey County, Simpson’s Hummock, and near Long Key in the Everglades, Dade County, Florida: var. eximea Sarg., differing from the type in its narrow elliptic to narrow oblong-obovate leaves and pale bark; a tree rarely 20° high, with a trunk 8′—12′ in diameter; rarely a shrub; dry sandy open woods, near Springfield, Livingston Parish and near Hammond, Tangipahoa Parish, eastern Louisiana. The following small shrubby small-leaved forms are recognized: var. fusiformis Sarg., with oblong-ovate leaves acute at apex, rounded or cuneate at base, entire or occasionally dentate, and pale pubescent below, and small fruit; dry limestone ridges and flat-topped hills of the Edwards Plateau (Kerr and Comal Counties), western Texas: var. dentata Chapm., distinct in the oblong-obovate repand-dentate lower leaves with large triangular teeth, acute at the broad apex, often 4′ long and 1¼′ wide at the base of the stems, and much larger than the oblong-lanceolate entire upper leaves; common in sterile pine-barrens near the coast of Florida: var. maritima Sarg., with oblong-obovate or rarely lanceolate leaves, acute and apiculate or rounded at apex, cuneate at base, and entire or slightly and irregularly toothed above the middle; fruit solitary or in pairs, or rarely in elongated spikes (Quercus succulenta [279] Small); sandy barrens near the coast, South Carolina to Miami, Dade County, Florida: var. pygmaea

The Project Gutenberg eBook of Manual of the Trees of North America (Exclusive of Mexico) 2nd ed.

MANUAL OF THE TREES OF NORTH AMERICA (EXCLUSIVE OF MEXICO)

PREFACE TO THE SECOND EDITION

PREFACE

TABLE OF CONTENTS

SYNOPSIS OF THE FAMILIES OF PLANTS DESCRIBED IN THIS BOOK

ANALYTICAL KEY TO THE GENERA OF PLANTS INCLUDED IN THIS BOOK, BASED CHIEFLY ON THE CHARACTER OF THE LEAVES

TREES OF NORTH AMERICA (Exclusive of Mexico)

Class 1. GYMNOSPERMÆ.

I. PINACEÆ.

CONSPECTUS OF THE NORTH AMERICAN GENERA.

1. PINUS Duham. Pine.

SOFT PINES.

1. Pinus Strobus L. White Pine.

2. Pinus monticola D. Don. White Pine.

3. Pinus Lambertiana Dougl. Sugar Pine.

4. Pinus flexilis James. Rocky Mountain White Pine.

5. Pinus albicaulis Engelm. White Pine.

6. Pinus Balfouriana Balf. Foxtail Pine.

7. Pinus aristata Engelm. Foxtail Pine. Hickory Pine.

8. Pinus cembroides Zucc. Nut Pine. Piñon.

Pinus cembroides var. Parryana Voss. Nut Pine. Piñon.

Pinus cembroides var. edulis Voss. Nut Pine. Piñon.

Pinus cembroides var. monophylla Voss. Nut Pine. Piñon.

PITCH PINES.

9. Pinus leiophylla Schlecht. and Cham. Yellow Pine.

10. Pinus ponderosa Laws. Yellow Pine. Bull Pine.

Pinus ponderosa var. Jeffreyi Vasey.

Pinus ponderosa var. arizonica Shaw. Yellow Pine.

11. Pinus palustris Mill. Long-leaved Pine. Southern Pine.

12. Pinus caribæa Morelet. Slash Pine. Swamp Pine.

13. Pinus tæda L. Loblolly Pine. Old Field Pine.

14. Pinus rigida Mill. Pitch Pine.

Pinus rigida var. serotina Loud. Pond Pine. Marsh Pine.

15. Pinus radiata D. Don. Monterey Pine.

16. Pinus attenuata Lemm. Knob-cone Pine.

17. Pinus Sabiniana Dougl. Digger Pine. Bull Pine.

18. Pinus Coulteri D. Don. Pitch Pine.

19. Pinus resinosa Ait. Red Pine. Norway Pine.

20. Pinus contorta Loud. Scrub Pine.

Pinus contorta var. latifolia S. Wats. Lodge-pole Pine.

21. Pinus Banksiana Lamb. Gray Pine. Jack Pine.

22. Pinus glabra Walt. Spruce Pine. Cedar Pine.

23. Pinus echinata Mill. Yellow Pine. Short-leaved Pine.

24. Pinus virginiana Mill. Jersey Pine. Scrub Pine.

25. Pinus clausa Sarg. Sand Pine. Spruce Pine.

26. Pinus muricata D. Don. Prickle-cone Pine.

27. Pinus pungens Lamb. Table Mountain Pine. Hickory Pine.

28. Pinus Torreyana Carr. Torrey Pine.

2. LARIX Adans. Larch.

CONSPECTUS OF THE NORTH AMERICAN SPECIES.

1. Larix laricina K. Koch. Tamarack. Larch.

2. Larix occidentalis Nutt. Tamarack.

3. Larix Lyallii Parl. Tamarack.

3. PICEA Dietr. Spruce.

CONSPECTUS OF THE NORTH AMERICAN SPECIES.

1. Picea mariana B. S. P. Black Spruce.

2. Picea rubra Link. Red Spruce.

3. Picea glauca Voss. White Spruce.

4. Picea Engelmannii Engelm. White Spruce. Engelmann Spruce.

5. Picea pungens Englm. Blue Spruce. Colorado Spruce.

6. Picea Breweriana S. Wats. Weeping Spruce.

7. Picea sitchensis Carr. Tideland Spruce. Sitka Spruce.

4. TSUGA Carr. Hemlock.

CONSPECTUS OF THE NORTH AMERICAN SPECIES.

1. Tsuga canadensis Carr. Hemlock.

2. Tsuga caroliniana Engelm. Hemlock.

3. Tsuga heterophylla Sarg. Hemlock.

4. Tsuga Mertensiana Sarg. Mountain Hemlock. Black Hemlock.

5. PSEUDOTSUGA Carr.

CONSPECTUS OF THE NORTH AMERICAN SPECIES.

1. Pseudotsuga taxifolia Britt. Douglas Spruce. Red Fir.

2. Pseudotsuga macrocarpa Mayr. Hemlock.

6. ABIES Link. Fir.

CONSPECTUS OF THE NORTH AMERICAN SPECIES.

1. Abies Fraseri Poir. Balsam Fir. She Balsam.

2. Abies balsamea Mill. Balsam Fir.

3. Abies lasiocarpa Nutt. Balsam Fir.

4. Abies grandis Lindl. White Fir.

5. Abies concolor Lindl. & Gord. White Fir.

MANUAL OF THE TREES OF NORTH AMERICA
(EXCLUSIVE OF MEXICO)

SYNOPSIS
OF THE FAMILIES OF PLANTS DESCRIBED IN THIS BOOK

ANALYTICAL KEY
TO THE GENERA OF PLANTS INCLUDED IN THIS BOOK, BASED CHIEFLY ON THE CHARACTER OF THE LEAVES

TREES OF NORTH AMERICA
(Exclusive of Mexico)